Republic, locality B4131: Ypresian, Washington

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Angiospermae - Nymphaeales - Nymphaeaceae
Nuphar cf. carlquisti
DeVore et al. 2015
SR 12-002-007A&B (rhizome), UWBM 57225B (rhizome), SR 01-01-10A&B (rhizome), SR 00-05-22A&B (rhizome), UWBM 95518 (Stigmatic disk), SR 13-004-009 (tepal), SR 13-004-011A&B (tepal)
Nuphar carlquistii n. sp. DeVore et al. 2015
DeVore et al. 2015
SR 02-17-03 A&B
Insecta - Odonata - Euphaeidae
Republica weatbrooki n. gen., n. sp. Archibald and Cannings 2021
Archibald and Cannings 2021 1 specimen
SR 06-59-08 (1 measurement)
Insecta - Odonata
Cephalozygoptera indet. Archibald et al. 2021
Archibald et al. 2021 1 specimen
SR 14-003-004
Insecta - Odonata - Dysagrionidae
Dysagrion pruettae n. sp. Archibald and Cannings 2021
Archibald et al. 2021 1 specimen
SR 13-005-012A,B (1 measurement)
Stenodiafanus westersidei n. gen., n. sp. Archibald and Cannings 2021
Archibald et al. 2021 1 specimen
SR 06-01-42 (1 measurement)
Dysagrionites informal sp. A
Archibald et al. 2021 1 specimen
SR 96-12-03
Dysagrionites delinei n. sp. Archibald and Cannings 2021
Archibald et al. 2021 2 specimens
SR 06-01-046AB; 94-05-26
Okanopteryx sp. Archibald and Cannings 2021
Archibald et al. 2021 2 specimens
SRUI99-72-96AB; SR 13-005-015AB
Okanopteryx jeppesenorum n. sp. Archibald and Cannings 2021
Archibald et al. 2021 7 specimens
SR 13-005-013; 11-43-09; 18-002-001; 13-003-001; 02-32-23; 07-22-05AB; 19-005-002AB (1 measurement)
Archibald et al. 2021 2 specimens
SR 11-51-07AB, 06-08-18
Okanagrion hobani Archibald and Cannings 2021
Archibald et al. 2021 9 specimens
SR 11-21-33A,B; 02-24-14A,B; 94-05-22A,B; 00-04-05; 10-72-03; 06-62-06; 13-005-017; 99-71-07
Okanagrion worleyae Archibald and Cannings 2021
Archibald et al. 2021 1 specimen
SR 11-21-09
Okanagrion liquetoalatum n. sp. Archibald and Cannings 2021
Archibald et al. 2021 1 specimen
SR 06-69-17A,B (1 measurement)
Insecta - Odonata - Whetwhetaksidae
Whetwhetaksa millerae n. gen., n. sp. Archibald and Cannings 2021
Archibald et al. 2021 5 specimens
SR 06-01-38; SR 10-36-10; SRUI99-98-34; SR 06-01-36; SR 10-14-19 (3 measurements)
Insecta - Odonata
Anisoptera informal indet. A
Archibald and Cannings 2019 1 specimen
SR 07-16-14
Insecta - Odonata - Aeshnidae
Aeshnidae informal indet. A
Archibald and Cannings 2019 1 specimen
SRUI 09-97-61
Idemlinea versatilis n. gen., n. sp. Archibald and Cannings 2019
Archibald and Cannings 2019 1 specimen
SR 07-05-14A,B
Ypshna brownleei n. sp. Archibald and Cannings 2019
Archibald and Cannings 2019 1 specimen
SR 01-06-13 (1 measurement)
Antiquiala snyderae n. gen., n. sp. Archibald and Cannings 2019
Archibald and Cannings 2019 1 specimen
SR 08-10-08A,B
Insecta - Neuroptera
Neuroptera indet. Linnaeus 1758
Archibald and Makarkin 2006 1 individual
SR95-25-31ab (1 measurement)
Insecta - Neuroptera - Osmylidae
Osmylidia glastrai n. sp. Makarkin et al. 2021
Makarkin et al. 2021 1 specimen
SR-02-22-15 (1 measurement)
Insecta - Neuroptera - Ithonidae
Allorapisma chuorum n. gen., n. sp. Makarkin and Archibald 2009
Makarkin and Archibald 2009 1 individual
SR08-14-01; collected by Chu family (1 measurement)
Polystoechotidae indet. Handlirsch 1906
Archibald and Makarkin 2006 1 individual
SR97-14-8a
Polystoechotites barksdalae n. sp. Archibald and Makarkin 2006
Archibald and Makarkin 2006 2 individuals
SR97-3-9ab, SR01-01-14 (2 measurements)
Palaeopsychops timmi n. sp. Archibald and Makarkin 2006
Archibald and Makarkin 2006 1 individual
SR02-25-01 (1 measurement)
Palaeopsychops marringerae n. sp. Archibald and Makarkin 2006
Archibald and Makarkin 2006 1 individual
SR97-08-05 (1 measurement)
Insecta - Neuroptera - Chrysopidae
Adamsochrysa wilsoni n. sp. Makarkin and Archibald 2013
Makarkin and Archibald 2013 1 specimen
SR06-23-01 (1 measurement)
Insecta - Raphidioptera - Raphidiidae
Raphidiidae informal indet. A
Archibald and Makarkin 2021 1 specimen
UWBM 74306
Insecta - Mecoptera - Cimbrophlebiidae
Cimbrophlebia brooksi n. sp. Archibald 2009
2 specimens
SR06-20-05, SR99-04-05 (2 measurements)
Cimbrophlebia westae n. sp. Archibald 2009
1 specimen
SRUI 099600 (1 measurement)
Insecta - Mecoptera - Dinopanorpidae
Dinokanaga cf. dowsonae Archibald 2005
Archibald 2005 2 specimens
SR01-01-03, part only, collected by Zach Adamsen, September 1997; SR1-1-4a, b, collected by Bill Richards, 1997,
Dinokanaga andersoni n. sp. Archibald 2005
Archibald 2005 1 specimen
SR01–06–01 Collected by Eric Anderson, August 2001 (1 measurement)
Insecta - Mecoptera - Eorpidae
Eorpa elverumi n. sp. Archibald et al. 2013
Archibald et al. 2013 1 specimen
SRUI 08-07-07 (1 measurement)
Eorpa ? ypsipeda Archibald et al. 2013
Archibald et al. 2013 8 specimens
SR 01-06-02, 08-35-04, 08-35-07, 09-11-01, 05-03-19, 95-10-03, 10-02-04, 12-004-002 (3 measurements)
Insecta - Hymenoptera - Formicidae
Propalosoma gutierrezae n. gen., n. sp. Dlussky and Rasnitsyn 1999
Dlussky and Rasnitsyn 1999 1 specimen
SR93-8-4 (2 measurements)
Klondikia whiteae n. gen., n. sp. Dlussky and Rasnitsyn 2002
Dlussky and Rasnitsyn 2002 1 specimen
SR-94-5-7 (coll Pat White 1997) (1 measurement)
Insecta
Eoprephasma hichensi n. gen., n. sp. Archibald and Bradler 2015
Archibald and Bradler 2015 1 specimen
SR 12-004-007 (1 measurement)
Insecta - Orthoptera - Palaeorehniidae
Republicopteron douseae n. gen., n. sp. Archibald et al. 2022
Archibald et al. 2022 2 specimens
SR 00-04-06, SR 21-005-001
Insecta - Diptera - Pipunculidae
Metanephrocerus belgardeae n. sp. Archibald et al. 2014
Archibald et al. 2014 1 specimen
SR 08-06-02 (1 measurement)
Reptilia - Theropoda
Aves indet. Linnaeus 1758
Mayr et al. 2019
see common names

Geography
Country:United States State/province:Washington
Coordinates: 48.7° North, 118.7° West (view map)
Paleocoordinates:54.4° North, 101.9° West
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period: Paleogene Epoch: Eocene
Stage: Ypresian 10 m.y. bin: Cenozoic 2
Key time interval: Ypresian
Age range of interval: 56 - 47.8 m.y. ago
Age estimate: 49.43 ± 0.54 Ma (Ar/Ar)
Stratigraphy
Formation:Klondike Mountain Member:Tom Thumb
Stratigraphic resolution:group of beds
Lithology and environment
Primary lithology:planar lamination,brown lithified mudstone
Environment:lacustrine - large Tectonic setting:rift
Geology comments: Basin developed in Republic graben
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Reason for describing collection:taxonomic analysis
Collection method comments: Material housed in the Burke Museum of Natural History and Culture, Seattle (UWBM), partly belonging to the Stonerose Interpretive Center, Republic, Washington (SR).
Metadata
Also known as:Boot Hill exposure
Database number:103206
Authorizer:M. Clapham, M. Uhen, K. Volkman Enterer:M. Clapham, K. Volkman, M. Uhen, J. Karr
Modifier:M. Clapham Research group:paleoentomology
Created:2011-01-23 17:43:42 Last modified:2013-11-25 13:02:06
Access level:the public Released:2011-01-23 17:43:42
Creative Commons license:CC BY
Reference information

Primary reference:
34867. S. B. Archibald. 2009. New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA. Zootaxa 2249:51-62 [M. Clapham/M. Clapham]

Secondary references:

46655 S. B. Archibald. 2005. New Dinopanorpidae (Insecta: Mecoptera) from the Eocene Okanagan Highlands (British Columbia, Canada and Washington State, USA). Canadian Journal of Earth Sciences 42:119-136 [M. Clapham/J. Karr]
56829 S. B. Archibald and S. Bradler. 2015. Stem-group stick insects (Phasmatodea) in the early Eocene at McAbee, British Columbia, Canada, and Republic, Washington, United States of America. The Canadian Entomologist 147:744-753 [M. Clapham/M. Clapham]
70811 S. B. Archibald and R. A. Cannings. 2019. Fossil dragonflies (Odonata: Anisoptera)from the early Eocene Okanagan Highlands, western North America. The Canadian Entomologist 151:783-816 [M. Clapham/M. Clapham]
78952 S. B. Archibald and R. A. Cannings. 2021. A new genus and species of Euphaeidae (Odonata, Zygoptera) from the early Eocene Okanagan Highlands locality at Republic, Washington, U.S.A. Zootaxa 4966:392-400 [M. Clapham/M. Clapham]
75770 S. B. Archibald, R. A. Cannings, R. J. Erickson, S. M. Bybee, and R. W. Mathewes. 2021. The Cephalozygoptera, a new, extinct suborder of Odonata with new taxa from the early Eocene Okanagan Highlands, western North America. Zootaxa 4934:1-133 [M. Clapham/M. Clapham]
87276 S. B. Archibald, J. J. Gu, and R. W. Mathewes. 2022. The Palaeorehniidae (Orthoptera, Ensifera, "Zeuneropterinae"), and new taxa from the early Eocene Okanagan Highlands, western North America. Zootaxa 5100:559-572 [M. Clapham/M. Clapham]
51914 S. B. Archibald, C. Kehlmaier, and R. W. Mathewes. 2014. Early Eocene big headed flies (Diptera: Pipunculidae) from the Okanagan Highlands, western North America. The Canadian Entomologist 146:429-443 [M. Clapham/M. Clapham]
36713 S. B. Archibald and V. N. Makarkin. 2006. Tertiary Giant Lacewings (Neuroptera: Polystoechotidae): revision and description of new taxa from Western North America and Denmark. Journal of Systematic Palaeontology 4(2):119-155 [M. Clapham/J. Karr/J. Karr]
78958 S. B. Archibald and V. N. Makarkin. 2021. Early Eocene snakeflies (Raphidioptera) of western North America from the Okanagan Highlands and Green River Formation. Zootaxa 4951:41-79 [M. Clapham/M. Clapham]
47557 S. B. Archibald, R. W. Mathewes, and D. R. Greenwood. 2013. The Eocene apex of panorpoid scorpionfly family diversity. Journal of Paleontology 87:677-695 [M. Clapham/M. Clapham]
83102 M. DeVore, W. Taylor, and K. Pigg. 2015. Nuphar carlquistii sp. nov. (Nymphaeaceae): A Water Lily from the Latest Early Eocene, Republic, Washington. International Journal of Plant Sciences 176(4):365-377 [K. Volkman/K. Volkman]
40968 G. M. Dlussky and A. P. Rasnitsyn. 1999. Two new species of aculeate hymenopterans (Vespida=Hymenoptera) from the Middle Eocene of the United States. Paleontological Journal 33:546-549 [M. Clapham/M. Clapham]
44418 G. M. Dlussky and A. P. Rasnitsyn. 2002. Ants (Hymenoptera: Formicidae) of Formation Green River and some other middle Eocene deposits of North America. Russian Entomological Journal 11:411-436 [M. Clapham/M. Clapham]
36851 V. N. Makarkin and S. B. Archibald. 2009. A new genus and first Cenozoic fossil record of moth lacewings (Neuroptera: Ithonidae) from the Early Eocene of North America. Zootaxa 2063:55-63 [M. Clapham/J. Karr]
44347 V. N. Makarkin and S. B. Archibald. 2013. A diverse new assemblage of green lacewings (Insecta, Neuroptera, Chrysopidae) from the early Eocene Okanagan Highlands, western North America. Journal of Paleontology 87:123-146 [M. Clapham/M. Clapham]
78951 V. N. Makarkin, S. B. Archibald, and R. W. Mathewes. 2021. New Protosmylinae (Neuroptera: Osmylidae) from the early Eocene of western North America, with taxonomic remarks. Zootaxa 4980:142-156 [M. Clapham/M. Clapham]
61691 G. Mayr, S. B. Archibald, G. W. Kaiser and R. W. Mathewes. 2019. Early Eocene (Ypresian) birds from the Okanagan Highlands, British Columbia (Canada) and Washington State (USA). Canadian Journal of Earth Sciences 56:803-813 [M. Uhen/B. Shipps/M. Uhen]