PBDB Collection

Peñacerrada outcrop I: Late/Upper Albian, Spain

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Arachnida - Araneae - Oonopidae

	Orchestina informal sp. 2 Smith 1882	Saupe et al. 2012	1 specimen
	MCNA 12593
Arachnida - Araneae - Lagonomegopidae

	? Lagonomegops cor n. sp. Pérez-de la Fuente et al. 2013	Pérez-de la Fuente et al. 2013	1 specimen
	MCNA 13295

	Spinomegops arcanus n. gen., n. sp. Pérez-de la Fuente et al. 2013	Pérez-de la Fuente et al. 2013	1 specimen
	MCNA 13307
Arachnida - Acariformes - Archaeorchestidae

	Archaeorchestes minguezae n. gen., n. sp. Arillo and Subías 2000	Arillo and Subías 2000	1 specimen
	recombined as Strieremaeus minguezae
	MCNA 8866
Malacostraca - Isopoda

	Oniscidea indet. Latreille 1802	Sánchez-García et al. 2021	3 specimens
	MCNA 9458, 9924.2, 14907
Malacostraca - Isopoda - Trichoniscidae

	Autrigoniscus resinicola n. gen., n. sp. Sánchez-García et al. 2021	Sánchez-García et al. 2021	4 specimens
	MCNA 12522, 12678, 12617, 13823.1
Malacostraca - Isopoda - Ligiidae

	Ligiidae informal indet. morphotype I Brandt 1833	Sánchez-García et al. 2021	1 specimen
	MCNA 9513

	Ligiidae informal indet. morphotype II Brandt 1833	Sánchez-García et al. 2021	1 specimen
	MCNA 14274

	Eoligiiscus tarraconensis n. gen., n. sp. Sánchez-García et al. 2021	Sánchez-García et al. 2021	1 specimen
	MCNA 9751
Malacostraca - Isopoda - Detonidae

	Heraclitus helenae n. gen., n. sp. Sánchez-García et al. 2021	Sánchez-García et al. 2021	1 specimen
	MCNA 12546
Malacostraca - Tanaidacea

	Proleptochelia tenuissima n. gen., n. sp. Vonk and Schram 2007	Vonk and Schram 2007	1 specimen
	MCNA 9846A
Malacostraca - Tanaidacea - Alavatanaidae

	Electrotanais monolithus n. gen., n. sp. Sánchez-García et al. 2015	Sánchez-García et al. 2015	1 specimen
	MCNA 12530

	Eurotanais terminator n. gen., n. sp. Sánchez-García et al. 2015	Sánchez-García et al. 2015	1 specimen
	MCNA 13285

	Alavatanais carabe n. gen., n. sp. Vonk and Schram 2007	Vonk and Schram 2007	8 specimens
	MCNA 9537, 9088; 13888,14301a,b, 13890, 10226, 14478 in Sánchez-García et al., 2015

	Alavatanais cf. carabe Vonk and Schram 2007	Sánchez-García et al. 2015	2 specimens
	MCNA 12104, 12105

	Alavatanais aff. carabe Vonk and Schram 2007	Sánchez-García et al. 2015	2 specimens
	MCNA 9201a,b (paratypes of Proleptochelia euskadiensis in Vonk & Schram, 2007)

	Proleptochelia euskadiensis n. sp. Vonk and Schram 2007	Vonk and Schram 2007	3 specimens
	synonym of Alavatanais carabe
	MCNA 9846B, 8637, 9449

	Alavatanais margulisae n. sp. Sánchez-García et al. 2015	Sánchez-García et al. 2015	5 specimens
	MCNA 9583a,b, 9924, 12703, 12749
Insecta - Diptera - Limoniidae

	Gonomyia (Iberiana) penalveri n. sp. Kania‑Kłosok et al. 2022	Kania‑Kłosok et al. 2022	1 specimen
	MCNA 8818 (2 measurements)

	Helius hispanicus n. sp. Kania‑Kłosok et al. 2021	Kania‑Kłosok et al. 2021	1 specimen
	MCNA 9946

	Helius (Helius) spiralis n. sp. Kania et al. 2018	Kania et al. 2018	1 specimen
	MCNA 15078 (1 measurement)

	Helius (Helius) alavensis n. sp. Kania et al. 2016	Kania et al. 2016	1 specimen
	MCNA 9112

	Trichoneura (Cretalinea) xavieri n. sp. Kania-Kłosok et al. 2021	Kania-Kłosok et al. 2021	1 specimen
	MCNA 9735 (1 measurement)
Insecta - Diptera - Ceratopogonidae

	Archiaustroconops alavensis n. sp. Szadziewski and Arillo 1998	Szadziewski and Arillo 1998	5 specimens
	MCNA 8836.1, 8836.2, 8839, 8840, 8842 (2 measurements)

	Leptoconops zherikhini n. sp. Szadziewski and Arillo 2003		4 specimens
	recombined as Leptoconops (Leptoconops) zherikhini
	MCNA 9176, 8897, 8976.2, 9571 (only says Alava amber) (2 measurements)

	Protoculicoides skalskii n. sp. Szadziewski and Arillo 1998	Szadziewski and Arillo 1998	1 specimen
	recombined as Gerontodacus skalskii
	MCNA 8645 (2 measurements)
Insecta - Diptera - Cecidomyiidae

	Eltxo cretaceus n. gen., n. sp. Arillo and Nel 2000	Arillo and Nel 2000	1 specimen
	MCNA 8824 (2 measurements)

	Cretohaplusia ortunoi n. gen., n. sp. Arillo and Nel 2000	Arillo and Nel 2000	1 specimen
	MCNA 9076 (2 measurements)
Insecta - Diptera - Keroplatidae

	Hegalari minor n. sp. Blagoderov and Arillo 2002	Blagoderov and Arillo 2002	1 specimen
	MCNA-9292 (2 measurements)

	Hegalari antzinako n. gen., n. sp. Blagoderov and Arillo 2002	Blagoderov and Arillo 2002	1 specimen
	MCNA-8820 (2 measurements)
Insecta - Diptera - Mycetophilidae

	Allocotocera xavieri n. sp. Blagoderov and Arillo 2002	Blagoderov and Arillo 2002	2 specimens
	MCNA-9941.1, MCNA-9403 (2 measurements)

	Alavamanota hispanica n. gen., n. sp. Blagoderov and Arillo 2002	Blagoderov and Arillo 2002	1 specimen
	MCNA-8819 (2 measurements)
Insecta - Diptera - Phoridae

	Euliphora grimaldii n. gen., n. sp. Arillo and Mostovski 1999	Arillo and Mostovski 1999	1 specimen
	MCNA 8648 (2 measurements)
Insecta - Diptera - Atelestidae

	Alavesia subiasi n. gen., n. sp. Waters and Arillo 1999	Waters and Arillo 1999	3 specimens
	MCNA-8837, 8838, 8841 (3 measurements)
Insecta - Diptera - Chimeromyiidae

	Chimeromyina concilia n. gen., n. sp. Arillo and Grimaldi 2009	Grimaldi et al. 2009	1 specimen
	MCNA 8882 (1 measurement)

	Chimeromyia alava n. sp. Arillo and Grimaldi 2009	Grimaldi et al. 2009	5 specimens
	MCNA 9238, 9318, 8883, 8743.2, 8886 (6 measurements)
Insecta - Diptera - Stratiomyidae

	Lysistrata emerita n. gen., n. sp. Grimaldi and Arillo 2011	Grimaldi et al. 2011	1 specimen
	MCNA 12698 (1 measurement)
Insecta - Diptera - Xylomyidae

	? Xylomyidae indet. Verrall 1909	Grimaldi et al. 2011	1 specimen
	MCNA 8833 (1 measurement)
Insecta - Diptera - Tethepomyiidae

	Tethepomyia buruhandi n. sp. Grimaldi and Arillo 2008	Grimaldi and Arillo 2008	1 specimen
	MCNA 8821 (1 measurement)

	Tethepomima holomma n. gen., n. sp. Grimaldi and Arillo 2008	Grimaldi and Arillo 2008	1 specimen
	MCNA 9915 (2 measurements)
Insecta - Diptera - Psychodidae

	Espanoderus barbarae n. gen., n. sp. Skibińska et al. 2017	Skibińska et al. 2017	3 specimens
	MCNA 13034, 9911, 12574 (2 measurements)
Insecta - Thysanoptera - Melanthripidae

	Gymnopollisthrips minor n. gen., n. sp. Peñalver et al. 2012	Peñalver et al. 2012	2 specimens
	MCNA 10731, 9472

	Gymnopollisthrips maior n. sp. Peñalver et al. 2012	Peñalver et al. 2012	1 specimen
	MCNA 9283
Insecta - Hemiptera - Hispanocaderidae

	Hispanocader lisae n. gen., n. sp. Golub et al. 2012	Golub et al. 2012	1 specimen
	MCNA 10656 (1 measurement)
Insecta - Hemiptera - Hydrometridae

	Alavametra popovi n. gen., n. sp. Sánchez-García and Nel 2016	Sánchez-García et al. 2016	2 specimens
	MCNA 12686, 12685 (2 measurements)
Insecta - Hemiptera - Mesoveliidae

	Iberovelia quisquilia n. gen., n. sp. Sánchez-García and Nel 2017	Sánchez-García et al. 2017	1 specimen
	MCNA 12804

	Glaesivelia pulcherrima n. gen., n. sp. Sánchez-García and Solórzano Kraemer 2017	Sánchez-García et al. 2017	2 specimens
	MCNA 12805, 12806
Insecta - Hemiptera - Tajmyraphididae

	Alavesiaphis margaritae n. gen., n. sp. Peñalver and Wegierek 2008	Peñalver and Wegierek 2008	1 specimen
	MCNA 10021
Insecta - Psocodea - Archaeatropidae

	Archaeatropos alavensis n. gen., n. sp. Baz and Ortuño 2000	Baz and Ortuño 2000	6 specimens
	MCNA 8834, 8646, 8868, 8640, 8802, 8889 (2 measurements)

	Archaeatropos alavensis Baz and Ortuño 2000	Álvarez-Parra et al. 2022	1 specimen
	MCNA-14912.1
Insecta - Psocodea - Empheriidae

	Preempheria antiqua n. gen., n. sp. Baz and Ortuño 2001	Baz and Ortuño 2001	2 specimens
	MCNA 8888, 8872 (2 measurements)

	Empheropsocus arilloi n. gen., n. sp. Baz and Ortuño 2001	Baz and Ortuño 2001	3 specimens
	MCNA 8677, 8814, 9731 (3 measurements)

	Empheropsocus margineglabrus n. sp. Baz and Ortuño 2001	Baz and Ortuño 2001	1 specimen
	MCNA 9755 (3 measurements)
Insecta - Psocodea - Electrentomidae

	Manicapsocidus enigmaticus n. gen., n. sp. Baz and Ortuño 2001	Baz and Ortuño 2001	3 specimens
	MCNA 8867, 8798-1, 8874 (2 measurements)
Insecta - Orthoptera - Elcanidae

	Hispanelcana lopezvallei n. sp. Peñalver and Grimaldi 2010	Peñalver and Grimaldi 2010	1 specimen
	MCNA 9898

	Hispanelcana alavensis n. sp. Peñalver and Grimaldi 2010	Peñalver and Grimaldi 2010	1 specimen
	MCNA 9569

	Hispanelcana arilloi n. gen., n. sp. Peñalver and Grimaldi 2010	Peñalver and Grimaldi 2010	4 specimens
	MCNA 9588, 9150, 9746, 9287
Insecta - Dermaptera

	Autrigonoforceps iberica n. gen., n. sp. Engel and Peris 2015	Engel et al. 2015	1 specimen
	MCNA 13964 (1 measurement)
Insecta - Blattodea

	Morazatermes krishnai n. gen., n. sp. Engel and Delclòs 2010	Engel and Delclòs 2010	9 specimens
	MCNA 12621, 10034, 8939, 9140, 9573, 10753, 12063.2, 12662, 12665 (2 measurements)

	Cantabritermes simplex n. gen., n. sp. Engel and Delclòs 2010	Engel and Delclòs 2010	1 specimen
	MCNA 12803 (1 measurement)

	Ithytermes montoyai n. gen., n. sp. Sánchez-García et al. 2020	Sánchez-García et al. 2020	2 specimens
	MCNA 14936.1, 2 (2 measurements)
Insecta - Hymenoptera - Anaxyelidae

	Eosyntexis parva n. sp. Ortega-Blanco et al. 2008	Ortega-Blanco et al. 2008	1 specimen
	MCNA 8756
Insecta - Hymenoptera - Diapriidae

	Diapriidae indet. (Haliday 1833)	Engel et al. 2013	1 specimen
	corrected as Diapriinae
	MCNA 9543
Insecta - Hymenoptera - Spathiopterygidae

	Spathiopteryx alavarommopsis n. gen., n. sp. Engel and Ortega-Blanco 2013	Engel et al. 2013	1 specimen
	MCNA 12603 (1 measurement)
Insecta - Hymenoptera

	Iberopria perialla n. gen., n. sp. Engel et al. 2013	Engel et al. 2013	1 specimen
	MCNA 9896 (2 measurements)
Insecta - Hymenoptera - Proterosceliopsidae

	Proterosceliopsis masneri n. gen., n. sp. Ortega-Blanco et al. 2014	Ortega-Blanco et al. 2014	5 specimens
	MCNA 13631, 12753, 12579, 12738, 9837 (2 measurements)
Insecta - Hymenoptera - Scelionidae

	Scelionidae indet. Thomson 1858	Ortega-Blanco et al. 2014	27 specimens
	original and current combination Scelioninae
	MCNA 8764, 8876, 8956, 8997, 9090, 9215, 9379, 9512, 9589(x2),9606, 9819, 9836, 10011, 10046, 10055, 12527, 12533, 12573, 12612(x2), 12626, 12792.1, 12792.2, 12793, 13276, 13288
Insecta - Hymenoptera

	Amissascelio temporarius n. gen., n. sp. Ortega-Blanco et al. 2014	Ortega-Blanco et al. 2014	8 specimens
	MCNA 13643, 9584, 9978, 10015, 10744, 10776, 11389, 12708

	Bruescelio platycephalus n. gen., n. sp. Ortega-Blanco et al. 2014	Ortega-Blanco et al. 2014	3 specimens
	MCNA 9536, 13284, 9499 (2 measurements)

	Perimoscelio tyrbastes n. gen., n. sp. Ortega-Blanco et al. 2014	Ortega-Blanco et al. 2014	5 specimens
	MCNA 9844, 8987, 9900, 10002, 9921

	Perimoscelio confector n. sp. Ortega-Blanco et al. 2014	Ortega-Blanco et al. 2014	11 specimens
	MCNA 9806, 8755.1, 8774, 9227, 9438, 10737, 10754, 12584, 12586, 9773, 9812

	Juxtascelio interitus n. gen., n. sp. Ortega-Blanco et al. 2014	Ortega-Blanco et al. 2014	11 specimens
	MCNA 12600, 8779, 8794, 9157, 9249, 9785, 9893, 11391, 12557, 9412, 9877

	Tithonoscelio resinalis n. gen., n. sp. Ortega-Blanco et al. 2014	Ortega-Blanco et al. 2014	1 specimen
	MCNA 8633.3 (2 measurements)

	Alavascelio delvallei n. gen., n. sp. Ortega-Blanco et al. 2014	Ortega-Blanco et al. 2014	14 specimens
	MCNA 9912.1, 9912.2, 9782, 12564(x8), 12726, 9892, 12666

	Electroteleiopsis hebdomas n. gen., n. sp. Ortega-Blanco et al. 2014	Ortega-Blanco et al. 2014	4 specimens
	MCNA 8909.1, 8793, 9878, 13834.1
Insecta - Hymenoptera - Serphitidae

	Aposerphites angustus n. sp. Ortega-Blanco et al. 2011	Ortega-Blanco et al. 2011	1 specimen
	MCNA 8651

	Serphites lamiak n. sp. Ortega-Blanco et al. 2011	Ortega-Blanco et al. 2011	1 specimen
	MCNA 8753
Insecta - Hymenoptera - Gallorommatidae

	Galloromma alavaensis n. sp. Ortega-Blanco et al. 2011	Ortega-Blanco et al. 2011	3 specimens
	MCNA 12630, 13281,13268 (2 measurements)
Insecta - Hymenoptera - Alavarommatidae

	Alavaromma orchamum n. gen., n. sp. Ortega-Blanco et al. 2011	Ortega-Blanco et al. 2011	1 specimen
	MCNA 9127 (2 measurements)
Insecta - Hymenoptera - Mymarommatidae

	Archaeromma hispanicum n. sp. Ortega-Blanco et al. 2011	Ortega-Blanco et al. 2011	1 specimen
	MCNA 9482 (2 measurements)
Insecta - Hymenoptera - Braconidae

	Protorhyssalopsis perrichoti n. gen., n. sp. Ortega-Blanco et al. 2011	Ortega-Blanco et al. 2011	1 specimen
	MCNA 9796 (2 measurements)

	Archephedrus stolamissus n. gen., n. sp. Ortega-Blanco et al. 2009	Ortega-Blanco et al. 2009	1 specimen
	MCNA 8785
Insecta - Hymenoptera - Maimetshidae

	Iberomaimetsha rasnitsyni n. gen., n. sp. Ortega-Blanco et al. 2011	Perrichot et al. 2011	2 specimens
	MCNA 9928, 8765

	Iberomaimetsha nihtmara n. sp. Ortega-Blanco et al. 2011	Perrichot et al. 2011	6 specimens
	MCNA 13049, 8758, 8790, 9373, 9918, 10732
Insecta - Hymenoptera - Megalyridae

	Megalava truncata n. gen., n. sp. Perrichot 2009	Perrichot 2009	1 specimen
	MCNA 9416

	Valaa delclosi n. gen., n. sp. Perrichot 2009	Perrichot 2009	1 specimen
	MCNA 12578a
Insecta - Hymenoptera - Evaniidae

	Iberoevania roblesi n. gen., n. sp. Peñalver et al. 2010	Peñalver et al. 2010	1 specimen
	MCNA 8759

	Cretevania alonsoi n. sp. Peñalver et al. 2010	Peñalver et al. 2010	3 specimens
	MCNA 9601, 9757, 9879
Insecta - Hymenoptera - Radiophronidae

	Radiophron ibericus n. gen., n. sp. Ortega-Blanco et al. 2010	Ortega-Blanco et al. 2010	7 specimens
	MCNA 8754, MCNA 8789, MCNA 8989, MCNA 9561, MCNA 9576, MCNA 13030.1, MCNA 13030.2

	Microcostaphron parvus n. gen., n. sp. Ortega-Blanco et al. 2010	Ortega-Blanco et al. 2010	1 specimen
	MCNA 8769
Insecta - Hymenoptera - Embolemidae

	Embolemus periallus n. sp. Ortega-Blanco et al. 2011	Ortega-Blanco et al. 2011	1 specimen
	recombined as Ampulicomorpha perialla
	MCNA 12709 (2 measurements)
Insecta - Hymenoptera - Bethylidae

	Lancepyris alavaensis n. sp. Ortega-Blanco and Engel 2013	Ortega-Blanco and Engel 2013	1 specimen
	recombined as Zophepyris alavaensis
	MCNA 9759

	Cretepyris martini n. gen., n. sp. Ortega-Blanco and Engel 2013	Ortega-Blanco and Engel 2013	3 specimens
	MCNA 8982, 9613, 12548

	Cretepyris ? martini Ortega-Blanco and Engel 2013	Ortega-Blanco and Engel 2013	1 specimen
	MCNA 9971

	Liztor pilosus n. gen., n. sp. Ortega-Blanco and Engel 2013	Ortega-Blanco and Engel 2013	3 specimens
	MCNA 9200, 9392, 8974
Insecta - Hymenoptera - Scolebythidae

	Scolebythidae informal indet. 1 Evans 1963	Engel et al. 2013	1 specimen
	MCNA 8757

	Scolebythidae informal indet. 2 Evans 1963	Engel et al. 2013	1 specimen
	MCNA 9895

	Ectenobythus iberiensis n. gen., n. sp. Engel et al. 2013	Engel et al. 2013	2 specimens
	MCNA 9815, 9483 (2 measurements)
Insecta - Hymenoptera - Stigmaphronidae

	Burmaphron sorginak n. sp. Ortega-Blanco et al. 2011	Ortega-Blanco et al. 2011	11 specimens
	MCNA 12618, 8982, 9405, 10726, 11226.1, 11226.2, 12589, 12781.1, 12781.2, 12795, 13841

	Burmaphron iratxoak n. sp. Ortega-Blanco et al. 2011	Ortega-Blanco et al. 2011	4 specimens
	MCNA 9346.2, 8806.1, 9180, 9526

	Burmaphron jentilak n. sp. Ortega-Blanco et al. 2011	Ortega-Blanco et al. 2011	9 specimens
	MCNA 8981, 8775.2, 9081, 9146, 9306, 10769, 12562, ?8780, ?12588

	Libanophron sugaar n. sp. Ortega-Blanco et al. 2011	Ortega-Blanco et al. 2011	16 specimens
	MCNA 8983, 8784 (13 individuals), 12728, 9189

	Hippocoon basajauni n. sp. Ortega-Blanco et al. 2011	Ortega-Blanco et al. 2011	2 specimens
	MCNA 9765, 9495

	Tagsmiphron olentzero n. sp. Ortega-Blanco et al. 2011	Ortega-Blanco et al. 2011	4 specimens
	MCNA 12733, 8806.2, 9578, 9976

	Elasmophron mari n. sp. Ortega-Blanco et al. 2011	Ortega-Blanco et al. 2011	1 specimen
	MCNA 9502
Insecta - Raphidioptera - Baissopteridae

	? Baissoptera cretaceoelectra n. sp. Pérez-de la Fuente et al. 2012	Pérez-de la Fuente et al. 2012	1 specimen
	MCNA 12068.4
Insecta - Raphidioptera - Mesoraphidiidae

	Mesoraphidiidae informal indet. 1 Martynov 1925	Pérez-de la Fuente et al. 2012	1 specimen
	MCNA 9218

	Mesoraphidiidae informal indet. 3 Martynov 1925	Pérez-de la Fuente et al. 2012	1 specimen
	MCNA 9316

	Alavaraphidia imperterrita n. gen., n. sp. Pérez-de la Fuente et al. 2012	Pérez-de la Fuente et al. 2012	1 specimen
	MCNA 13608

	? Styporaphidia hispanica n. sp. Pérez-de la Fuente et al. 2012	Pérez-de la Fuente et al. 2012	1 specimen
	MCNA 9343
Insecta - Neuroptera - Berothidae

	Berothidae indet. Handlirsch 1906	Pérez-de la Fuente et al. 2020	1 specimen
	MCNA 9294

	Berothidae informal indet. 1 Handlirsch 1906	Pérez-de la Fuente et al. 2020	1 specimen
	MCNA 10033

	Berothidae informal indet. 2 Handlirsch 1906	Pérez-de la Fuente et al. 2020	1 specimen
	MCNA 9902

	Berothidae informal indet. 3 Handlirsch 1906	Pérez-de la Fuente et al. 2020	1 specimen
	MCNA 13379.1

	Berothidae informal indet. 4 Handlirsch 1906	Pérez-de la Fuente et al. 2020	1 specimen
	MCNA 12590
Insecta - Coleoptera - Leiodidae

	Cretaciella sorianoae n. gen., n. sp. Perreau 2019	Perreau 2019	1 specimen
	nomen nudum belonging to Oritocatopini
	MCNA 9423 (1 measurement)
Insecta - Coleoptera - Staphylinidae

	Cretasonoma corinformibus n. gen., n. sp. Peris et al. 2014	Peris et al. 2014	7 specimens
	MCNA 8654, 9990, 12663, 13254, 14251, 14247, 14265

	Penarhytus tenebris n. gen., n. sp. Peris et al. 2014	Peris et al. 2014	1 specimen
	MCNA 12683

	Paederinae indet. (Fleming 1821)	Peris et al. 2014	1 specimen
	MCNA-8912

	Prosolierius parvus n. sp. Peris et al. 2014	Peris et al. 2014	3 specimens
	MCNA 14190, 12752, 13611
Insecta - Coleoptera - Trogossitidae

	Trogossitidae indet. (Latreille 1802)	Peris et al. 2014	2 specimens
	original and current combination Trogossitarii
	MCNA 9259, 10763
Insecta - Coleoptera - Oedemeridae

	Darwinylus marcosi n. gen., n. sp. Peris 2017	Peris 2017	1 specimen
	MCNA 11229 (2 measurements)
Insecta - Coleoptera - Mordellidae

	Mediumiuga sinespinis n. gen., n. sp. Peris and Ruzzier 2013	Peris and Ruzzier 2013	1 specimen
	MCNA 8839
Insecta - Coleoptera - Monotomidae

	Monotomidae indet. (Laporte 1840)	Peris and Delclòs 2015	1 specimen
	MCNA 9158

	Cretakarenni hispanicus n. sp. Peris and Delclòs 2015	Peris and Delclòs 2015	1 specimen
	MCNA 8655 (1 measurement)

	Rhizophtoma longus n. sp. Peris and Delclòs 2015	Peris and Delclòs 2015	3 specimens
	MCNA 9184, 9371, 9552 (1 measurement)
Collembola - Entomobryomorpha

	Entomobryomorpha indet. Börner 1913	Sánchez-García and Engel 2016	1 specimen
	MCNA 9560
Collembola - Entomobryomorpha - Isotomidae

	Protoisotoma autrigoniensis n. sp. Sánchez-García and Engel 2016	Sánchez-García and Engel 2016	2 specimens
	MCNA 12788, ?12787

	Proisotoma communis Sánchez-García and Engel 2016	Sánchez-García and Engel 2016	68 specimens
	MCNA 10040.28,30-35,37-49, 10744.2, 11231.2-46, 12609, 12674.1

	? Cryptopygus sp. Willem 1901	Simón-Benito et al. 2002	11 specimens
	MCNA 9273.1, 9273.2, 9273.3, 8969.1-2, 9273, 9324, 9612
	= Proisotoma communis n. sp. Sánchez-García and Engel 2016	Sánchez-García and Engel 2016

	Proisotoma (Ballistura) sp. Borner 1901	Simón-Benito et al. 2002	4 specimens
	MCNA 9148, 9464.1-2, 10061
	= Proisotoma communis Sánchez-García and Engel 2016	Sánchez-García and Engel 2016

	? Anurophorus sp. Nicolet 1842	Simón-Benito et al. 2002	3 specimens
	MCNA 10070, 10071.1-2
	= Proisotoma communis Sánchez-García and Engel 2016	Sánchez-García and Engel 2016

	? Onychiurus sp. Gervais 1841	Simón-Benito et al. 2002	1 specimen
	MCNA 9162
	= Proisotoma communis Sánchez-García and Engel 2016	Sánchez-García and Engel 2016

	? Micranurida sp. Börner 1901	Simón-Benito et al. 2002	2 specimens
	MCNA 9612.1-7
	= Proisotoma communis Sánchez-García and Engel 2016	Sánchez-García and Engel 2016

	Burmisotoma spinulifera n. sp. Sánchez-García and Engel 2016	Sánchez-García and Engel 2016	1 specimen
	MCNA 12583
Collembola - Symphypleona

	? Sminthurus informal sp. 1 (Latreille 1802)	Simón-Benito et al. 2002	1 specimen
	MCNA 10016
	= Symphypleona indet. Borner 1901	Sánchez-García and Engel 2016
Collembola - Symphypleona - Sminthurididae

	Pseudosminthurides stoechus n. gen., n. sp. Sánchez-García and Engel 2016	Sánchez-García and Engel 2016	1 specimen
	MCNA 12788.1
Collembola - Symphypleona - Sminthuridae

	Archeallacma dolichopoda n. gen., n. sp. Sánchez-García and Engel 2016	Sánchez-García and Engel 2016	3 specimens
	MCNA 13850.4, 13850.5, 14284.2

	? Sminthurus informal sp. 2 (Latreille 1802)	Simón-Benito et al. 2002	1 specimen
	MCNA 10048
	= Katiannasminthurus xenopygus n. gen., n. sp. Sánchez-García and Engel 2016	Sánchez-García and Engel 2016

	Arrhopalites sp. Börner 1906	Simón-Benito et al. 2002	1 specimen
	MCNA 9311
	= Sphyrotheciscus senectus n. gen., n. sp. Sánchez-García and Engel 2016	Sánchez-García and Engel 2016
Collembola - Symphypleona - Katiannidae

	? Fasciosminthurus sp. Gisin 1960	Simón-Benito et al. 2002	1 specimen
	MCNA 10047
	= Cretokatianna bucculenta n. gen., n. sp. Sánchez-García and Engel 2016	Sánchez-García and Engel 2016
see common names

Geography

Country:	Spain	State/province:	Álava
Coordinates:	42.6° North, 2.7° West (view map)
Paleocoordinates:	30.4° North, 1.4° East
Basis of coordinate:	based on nearby landmark
Geographic resolution:	outcrop

Time

Period:	Cretaceous	Epoch:	Early/Lower Cretaceous
Stage:	Albian	10 m.y. bin:	Cretaceous 4
Key time interval:	Late/Upper Albian
Age range of interval:	106.20000 - 100.50000 m.y. ago

Stratigraphy

Formation:

Escucha

Lithology and environment

Primary lithology:	amber
Environment:	terrestrial indet.

Taphonomy

Modes of preservation:	body,soft parts,amber
Size of fossils:	mesofossils,microfossils

Collection methods and comments

Reason for describing collection:	taxonomic analysis
Collection method comments: Housed in Museo de Ciencias Naturales de Álava

Metadata

Also known as:	Alava amber
Database number:	108769
Authorizer:	M. Clapham	Enterer:	M. Clapham, A. Noshirvan, J. Karr
Modifier:	M. Clapham	Research group:	paleoentomology
Created:	2011-05-04 05:10:26	Last modified:	2018-02-23 11:33:35
Access level:	the public	Released:	2011-05-04 05:10:26
Creative Commons license:	CC BY

Reference information

Primary reference:

39277.

R. Szadziewski and A. Arillo. 2003. The oldest fossil record of the extant subgenus Leptoconops (Leptoconops) (Diptera: Ceratopogonidae). Acta Zoologica Cracoviensia 46(Suppl):271-275 [M. Clapham/M. Clapham/M. Clapham]

Secondary references:

81784		S. Álvarez-Parra, E. Peñalver, A. Nel and X. Delclòs. 2022. New barklice (Psocodea, Trogiomorpha) from Lower Cretaceous Spanish amber. Papers in Palaeontology 8(e1436) [M. Clapham/M. Clapham]
51691		A. Arillo and M. B. Mostovski. 1999. A new genus of Prioriphorinae (Diptera Phoridae) from the Lower Cretaceous amber of Alava (Spain). Studia Dipterologica 6:251-255 [M. Clapham/M. Clapham]
50845		A. Arillo and A. Nel. 2000. Two new fossil cecidomyiids flies from the Lower Cretaceous amber of Alava (Spain) (Diptera, Cecidomyiidae). Bulletin de la Société Entomologique de France 105:285-288 [M. Clapham/M. Clapham]
53474		A. Arillo and L. S. Subías. 2000. A new fossil oribatid mite, Archaeorchestes minguezae n. gen. n. sp. from the Spanish Lower Cretaceous amber. Description of a new family Archaeorchestidae (Acariformes, Oribatida, Zetorchestoidea). Mitteilungen aus dem Geologisch-Paläontologischen Institut der Universität Hamburg 84:231-236 [M. Clapham/M. Clapham]
40220		A. Baz and V. M. Ortuño. 2000. Archaeatropidae, a new family of Psocoptera from the Cretaceous Amber of Alava, northern Spain. Annals of the Entomological Society of America 93:367-373 [M. Clapham/M. Clapham]
39636		A. Baz and V. M. Ortuño. 2001. New genera and species of empheriids (Psocoptera: Empheriidae) from the Cretaceous amber of Alava, northern Spain. Cretaceous Research 22:575-584 [M. Clapham/M. Clapham]
40406		A. Baz and V. Ortuño. 2001. A new electrotomoid psocid (Psocoptera) from the Cretaceous amber of Alava (northern Spain). Deutsche Entomologische Zeitschrift 48:27-32 [M. Clapham/M. Clapham/M. Clapham]
40877		V. A. Blagoderov and A. Arillo. 2002. New Sciaroidea (Insecta: Diptera) in Lower Cretaceous amber from Spain. Studia Dipterologica 9:31-40 [M. Clapham/A. Noshirvan/M. Clapham]
39358		M. S. Engel and X. Delclòs. 2010. Primitive termites in Cretaceous amber from Spain and Canada (Isoptera). Journal of the Kansas Entomological Society 83:111-128 [M. Clapham/M. Clapham]
48549		M. S. Engel, J. Ortega-Blanco, and R. C. McKellar. 2013. New scolebythid wasps in Cretaceous amber from Spain and Canada, with implications for the phylogeny of the family (Hymenoptera: Scolebythidae). Cretaceous Research 46:31-42 [M. Clapham/M. Clapham]
47622		M. S. Engel, J. Ortega-Blanco, C. Soriano, D. A. Grimaldi, and X. Delclòs. 2013. A new lineage of enigmatic diaprioid wasps in Cretaceous amber (Hymenoptera: Diaprioidea). American Museum Novitates 3771:1-23 [M. Clapham/M. Clapham]
52343		M. S. Engel, D. Peris, S. Chatzimanolis and X. Delclòs. 2015. An earwig (Insecta: Dermaptera) in Early Cretaceous amber from Spain. Insect Systematics & Evolution 46:291-300 [M. Clapham/M. Clapham/M. Clapham]
42071		V. B. Golub, Y. A. Popov, and A. Arillo. 2012. Hispanocaderidae n. fam. (Hemiptera: Heteroptera: Tingoidea), one of the oldest lace bugs from the Lower Cretaceous Álava amber (Spain). Zootaxa 3270:41-50 [M. Clapham/M. Clapham]
45397		D. A. Grimaldi and A. Arillo. 2008. The Tethepomyiidae, a new family of enigmatic Cretaceous Diptera. Alavesia 2:259-265 [M. Clapham/M. Clapham]
39348		D. A. Grimaldi, A. Arillo, J. M. Cumming and M. Hauser. 2011. Brachyceran Diptera (Insecta) in Cretaceous ambers, Part IV, significant new Orthorrhaphous Taxa. ZooKeys 148:293-332 [M. Clapham/M. Clapham]
48656		D. A. Grimaldi, J. M. Cumming, and A. Arillo. 2009. Chimeromyiidae, a new family of Eremoneuran Diptera from the Cretaceous. Zootaxa 2078:34-54 [M. Clapham/M. Clapham]
58481		I. Kania, W. Krzemiński, and A. Arillo. 2016. First representative of the genus Helius Lepeletier and Serville, 1828 (Diptera, Limoniidae) from the Lower Cretaceous Álava amber (Spain). Cretaceous Research 63:33-38 [M. Clapham/M. Clapham/M. Clapham]
64891		I. Kania, W. Krzemiński, and A. Arillo. 2018. A new peculiar species of the genus Helius Lepeletier & Serville, 1828 (Diptera, Limoniidae) from Cretaceous Alava amber (Spain). Earth and Environmental Science Transactions of the Royal Society of Edinburgh 107:231-237 [M. Clapham/M. Clapham/M. Clapham]
76897		I. Kania-Kłosok, W. Krzemiński, K. Kopeć and A. Arillo. 2021. The oldest evolutionary lineage of Trichoneura Loew, 1850 (Diptera, Limoniidae) and the first evidence of this genus in Cretaceous Spanish amber. Insects 12(411) [M. Clapham/M. Clapham]
83346		I. Kania‑Kłosok, A. Arillo, M. Tuchowski, Q. Q. Zhang, and W. Krzemiński. 2022. Early stage of evolution of Gonomyia (Diptera: Limoniidae), new significant discoveries in the Cretaceous Iberian and Kachin amber. Scientific Reports 12(21118) [M. Clapham/M. Clapham]
77190		I. Kania‑Kłosok, W. Krzemiński, and A. Arillo. 2021. Two new long‑rostrum cranefly species from the Cretaceous Iberian amber (Diptera, Limoniidae, Helius). Scientific Reports 11(12851) [M. Clapham/M. Clapham]
40136		J. Ortega-Blanco, D. J. Bennett, X. Delclòs and M. S. Engel. 2009. A Primitive Aphidiine Wasp in Albian Amber from Spain and a Northern Hemisphere Origin for the Subfamily (Hymenoptera: Braconidae: Aphidiinae). Journal of the Kansas Entomological Society 82(4):273-282 [M. Clapham/A. Noshirvan/M. Clapham]
35978		J. Ortega-Blanco, X. Declòs, and M. S. Engel. 2011. A Protorhyssaline Wasp in Early Cretaceous Amber from Spain (Hymenoptera: Braconidae). Journal of the Kansas Entomological Society 84(1):51-57 [M. Clapham/M. Clapham]
35977		J. Ortega-Blanco, X. Declòs, and M. S. Engel. 2011. The Wasp Family Embolemidae in Early Cretaceous Amber from Spain (Hymenoptera: Chrysidoidea). Journal of the Kansas Entomological Society 84(1):36-42 [M. Clapham/M. Clapham]
39574		J. Ortega-Blanco, X. Delclòs, and M. S. Engel. 2011. Diverse stigmaphronid wasps in Early Cretaceous amber from Spain (Hymenoptera: Ceraphronoidea: Stigmaphronidae). Cretaceous Research 32:762-773 [M. Clapham/M. Clapham/M. Clapham]
39579		J. Ortega-Blanco, X. Delclòs, E. Peñalver and M. S. Engel. 2011. Serphitid wasps in Early Cretaceous amber from Spain (Hymenoptera: Serphitidae). Cretaceous Research 32:143-154 [M. Clapham/M. Clapham]
48251		J. Ortega-Blanco and M. S. Engel. 2013. Bethylidae from Early Cretaceous Spanish amber (Hymenoptera: Chrysidoidea). Journal of the Kansas Entomological Society 86:264-276 [M. Clapham/M. Clapham]
51738		J. Ortega-Blanco, R. C. McKellar, and M. S. Engel. 2014. Diverse scelionid wasps from Early Cretaceous Álava amber, Spain (Hymenoptera: Platygastroidea). Bulletin of Geosciences 89:553-571 [M. Clapham/M. Clapham]
36176		J. Ortega-Blanco, E. Peñalver, X. Declòs and M. S. Engel. 2011. False fairy wasps in Early Cretaceous amber from Spain (Hymenoptera: Mymarommatoidea). Palaeontology 54(3):511-523 [M. Clapham/M. Clapham]
48386		J. Ortega-Blanco, A. P. Rasnitsyn, and X. Delclòs. 2008. First record of anaxyelid woodwasps (Hymenoptera: Anaxyelidae) in Lower Cretaceous Spanish amber. Zootaxa 1937:39-50 [M. Clapham/M. Clapham]
40901		J. Ortega-Blanco, A. P. Rasnitsyn, and X. Delclos. 2010. A New Family of Ceraphronoid Wasps form Early Cretaceous Álava Amber. Acta Palaeontologica Polonica 55(2):265-276 [M. Clapham/A. Noshirvan]
39644		E. Peñalver and D. A. Grimaldi. 2010. Latest occurrences of the Mesozoic family Elcanidae (Insecta: Orthoptera), in Cretaceous amber from Myanmar and Spain. Annales de la Société Entomologique de France 46:88-99 [M. Clapham/M. Clapham]
41709		E. Peñalver, C. C. Labandeira, E. Barrón, X. Delclòs, P. Nel, A. Nel, P. Tafforeau and C. Soriano. 2012. Thrips pollination of Mesozoic gymnosperms. Proceedings of the National Academy of Sciences 109:8623-8628 [M. Clapham/M. Clapham/M. Clapham]
44262		E. Peñalver, J. Ortega-Blanco, and X. Delclòs. 2010. Mesozoic Evaniidae (Insecta: Hymenoptera) in Spanish amber: reanalysis of the phylogeny of the Evanioidea. Acta Geologica Sinica 84:809-827 [M. Clapham/M. Clapham]
39677		E. Peñalver and P. Wegierek. 2008. A new genus and species of the family Tajmyraphididae (Hemiptera: Sternorrhyncha) in Early Cretaceous amber from Peñacerrada I (Spain). Alavesia 2:187-192 [M. Clapham/J. Karr]
69955		R. Pérez-de la Fuente, M. S. Engel, X. Delclòs and E. Peñalver. 2020. Straight-jawed lacewing larvae (Neuroptera) from Lower Cretaceous Spanish amber, with an account on the known amber diversity of neuropterid immatures. Cretaceous Research 106(104200) [M. Clapham/M. Clapham/M. Clapham]
42190		R. Pérez-de la Fuente, E. Peñalver, X. Delclòs and M. S. Engel. 2012. Snakefly diversity in Early Cretaceous amber from Spain (Neuropterida, Raphidioptera). ZooKeys 204:1-40 [M. Clapham/M. Clapham]
75086		R. Pérez-de la Fuente, E. Peñalver, and M. S. Engel. 2020. Beaded lacewings (Neuroptera: Berothidae) in amber from the Lower Cretaceous of Spain. Cretaceous Research 119(104705) [M. Clapham/M. Clapham]
47526		R. Pérez-de la Fuente, E. E. Saupe, and P. A. Selden. 2013. New lagonomegopid spiders (Araneae: †Lagonomegopidae) from Early Cretaceous Spanish amber. Journal of Systematic Palaeontology 11:531-553 [M. Clapham/M. Clapham]
62265		D. Peris. 2017. Early Cretaceous origin of pollen-feeding beetles (Insecta: Coleoptera: Oedemeridae). Cladistics 33:268-278 [M. Clapham/M. Clapham]
49685		D. Peris, S. Chatzimanolis, and X. Delclòs. 2014. Diversity of rove beetles (Coleoptera: Staphylinidae) in Early Cretaceous Spanish amber. Cretaceous Research 48:85-95 [M. Clapham/M. Clapham]
55468		D. Peris and X. Delclòs. 2015. Fossil Monotomidae (Coleoptera: Polyphaga) from Laurasian Cretaceous amber. Organisms Diversity & Evolution 15:333-342 [M. Clapham/M. Clapham/M. Clapham]
52620		D. Peris, J. Kolibáč, and X. Delclòs. 2014. Cretamerus vulloi gen. et sp. nov., the oldest bark-gnawing beetle (Coleoptera: Trogossitidae) from Cretaceous amber. Journal of Systematic Palaeontology 12:879-891 [M. Clapham/M. Clapham/M. Uhen]
47963		D. Peris and E. Ruzzier. 2013. A new tribe, new genus, and new species of Mordellidae (Coleoptera: Tenebrionoidea) from the Early Cretaceous amber of Spain. Cretaceous Research 45:1-6 [M. Clapham/M. Clapham]
71338		M. Perreau. 2019. Cretaciella sorianoae gen. et sp. nov. (Coleoptera, Leiodidae, Cholevinae, Oritocatopini), anophthalmic species from Albian amber of the Escucha Formation (Alava, Spain). Geosciences 9(521) [M. Clapham/M. Clapham/M. Clapham]
42084		V. Perrichot. 2009. Long-tailed wasps (Hymenoptera: Megalyridae) from Cretaceous and Paleogene European amber. Paleontological Contributions 1:1-35 [M. Clapham/M. Clapham]
40362		V. Perrichot, J. Ortega-Blanco, R. C. McKellar, X. Delclòs, D. Azar, A. Nel, P. Tafforeau and M. S. Engel. 2011. New and revised maimetshid wasps from Cretaceous ambers (Hymenoptera, Maimetshidae). ZooKeys 130:421-453 [M. Clapham/M. Clapham]
56224		A. Sánchez-García, A. Arillo, and A. Nel. 2016. The first water measurers from the Lower Cretaceous amber of Spain (Heteroptera, Hydrometridae, Heterocleptinae). Cretaceous Research 57:111-121 [M. Clapham/M. Clapham]
62339		A. Sánchez-García and M. S. Engel. 2016. Long-term stasis in a diverse fauna of Early Cretaceous springtails (Collembola: Symphypleona). Journal of Systematic Palaeontology 15:513-537 [M. Clapham/M. Clapham]
59725		A. Sánchez-García and M. S. Engel. 2016. Springtails from the Early Cretaceous amber of Spain (Collembola: Entomobryomorpha), with an annotated checklist of fossil Collembola. American Museum Novitates 3862:1-47 [M. Clapham/M. Clapham]
63234		A. Sánchez-García, A. Nel, A. Arillo and M. M. Solórzano Kraemer. 2017. The semi-aquatic pondweed bugs of a Cretaceous swamp. PeerJ 5(e3760):1-32 [M. Clapham/M. Clapham/M. Uhen]
60621		A. Sánchez-García, E. Peñalver, R. Pérez-de Fuente and X. Delclòs. 2015. A rich and diverse tanaidomorphan (Crustacea: Tanaidacea) assemblage associated with Early Cretaceous resin-producing forests in North Iberia: palaeobiological implications. Journal of Systematic Palaeontology 13:645-676 [M. Clapham/M. Clapham]
64822		A. Sánchez-García, E. Peñalver, X. Delclòs and M. S. Engel. 2018. Mating and aggregative behaviors among basal hexapods in the Early Cretaceous. PLoS One 13(2):e0191669 [M. Clapham/M. Clapham]
71779		A. Sánchez-García, E. Peñalver, X. Delclòs and M. S. Engel. 2020. Early Cretaceous termites in amber from northern Spain (Isoptera). Cretaceous Research 110(104385) [M. Clapham/M. Clapham/M. Clapham]
85389		A. Sánchez-García, E. Peñalver, X. Delclòs and M. S. Engel. 2021. Terrestrial isopods from Spanish amber (Crustacea: Oniscidea): insights into the Cretaceous soil biota. American Museum Novitates 3974:1-32 [M. Clapham/M. Clapham]
47396		E. E. Saupe, R. Pérez-de la Fuente, P. A. Selden, X. Delclòs, P. Tafforeau and C. Soriano. 2012. New Orchestina Simon, 1882 (Araneae: Oonopidae) from Cretaceous ambers of Spain and France: first spiders described using phase-contrast X-ray synchrotron microtomography. Palaeontology 55:127-143 [M. Clapham/M. Clapham]
59727		J. C. Simón-Benito, V. M. Ortuño, and D. Espantaleón. 2002. Colémbolos (Collembola, Insecta) del ámbar Cretácico de Álava (cuenca vasco-cantábrica, norte de España). Estudios del Museo de Ciencias Naturales de Álava 17:83-92 [M. Clapham/M. Clapham]
69288		K. Skibińska, W. Krzemiński, and A. Arillo. 2017. The first Tanyderidae (Diptera) from Lower Cretaceous Álava amber (Spain). Historical Biology 31:872-878 [M. Clapham/M. Clapham]
51585		R. Szadziewski and A. Arillo. 1998. Biting midges (Diptera : Ceratopogonidae) from the lower Cretaceous Amber from Alava, Spain. Polskie Pismo Entomologiczne 67:291-298 [M. Clapham/M. Clapham]
60619		R. Vonk and F. R. Schram. 2007. Three new tanaid species (Crustacea, Peracarida, Tanaidacea) from the Lower Cretaceous. Journal of Paleontology 81:1502-1509 [M. Clapham/M. Clapham]
51692		S. B. Waters and A. Arillo. 1999. A new genus of Hybotidae (Diptera, Empidoidea) from Lower Cretaceous amber of Alava (Spain). Studia Dipterologica 6:59-66 [M. Clapham/M. Clapham]