PBDB Collection

Crato (SMNS collection): Late Aptian, Brazil

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Myriapoda - Scutigeromorpha - Scutigeridae

	Fulmenocursor tenax n. gen., n. sp. Wilson 2001	Wilson 2001	1 specimen
	SMNS 64275
Myriapoda - Scolopendromorpha - Scolopendridae

	Cratoraricrus oberlii n. gen., n. sp. Wilson 2003	Wilson 2003	1 specimen
	SMNS 64431
Diplura

	Ferrojapyx vivax n. gen., n. sp. Wilson and Martill 2001	Wilson and Martill 2001	3 specimens
	SMNS 64276; two additional in Staniczek & Bechly, 2007
Insecta - Hymenoptera - Sapygidae

	Cretofedtschenkia santanensis n. gen., n. sp. Osten 2007	Martill et al. 2007	1 specimen
	SMNS 66594
Insecta - Hymenoptera - Angarosphecidae

	Cretobestiola sp. Pulawski and Rasnitsyn 2000	Martill et al. 2007	1 specimen
	SMNS 66297
Insecta - Hymenoptera - Ampulicidae

	Ampulicidae indet. Shuckard 1840	Martill et al. 2007	1 specimen
	SMNS 66291

	Cariridris bipetiolata Brandão and Martins-Neto 1990	Martill et al. 2007	1 specimen
	SMNS 66565
Insecta - Hymenoptera - Vespidae

	Vespidae indet. (Latreille 1802)	Martill et al. 2007	1 specimen
	SMNS 66295
Insecta - Hymenoptera

	Parviformosus wohlrabeae n. gen., n. sp. Barling et al. 2013	Barling et al. 2013	1 specimen
	SMNS 70902
Insecta - Hymenoptera - Sepulcidae

	Prosyntexis gouleti Sharkey 1990	Jattiot et al. 2011	2 specimens
	SMNS 66301, 66698a/b (4 measurements)
Insecta - Diptera - Tipulidae

	Leptotarsus grimaldii Ribeiro and Lukashevich 2014	Krzemiński et al. 2017	1 specimen
	SMNS 66523

	Leptotarsus martinsnetoi Ribeiro and Lukashevich 2014	Krzemiński et al. 2017	1 specimen
	SMNS 326
Insecta - Diptera - Zhangsolvidae

	Cratomyoides cretacicus n. gen., n. sp. Wilkommen 2007	Martill et al. 2007	1 specimen
	SMNS 66179 (2 measurements)

	Cratomyiidae indet. Mazzarolo and Amorim 2000	Martill et al. 2007	1 specimen
	SMNS 66180
Insecta - Diptera - Tabanidae

	Tabanidae indet. (Latreille 1802)	Martill et al. 2007	1 specimen
	SMNS 66181
Insecta - Diptera - Rhagionidae

	? Rhagionidae indet. (Latreille 1802)	Martill et al. 2007	1 specimen
	SMNS 66185
Insecta - Diptera

	Asiloidea indet. Latreille 1802	Martill et al. 2007	1 specimen
	SMNS 66182 (Asilidae or Therevidae?)
Insecta - Diptera - Asilidae

	Asilidae indet. (Latreille 1802)	Martill et al. 2007	1 specimen
	SMNS 66183
Insecta - Diptera - Mydidae

	Cretomydas santanensis n. gen., n. sp. Wilkommen 2007	Martill et al. 2007	1 specimen
	SMNS 66178 (2 measurements)
Insecta - Neuroptera - Kalligrammatidae

	Makarkinia kerneri n. sp. Bechly and Makarkin 2016	Bechly and Makarkin 2016	1 specimen
	SMNS 70287 (1 measurement)
Insecta - Neuroptera - Rafaelianidae

	Rafaelia minima Nel et al. 2005	Nel et al. 2005	1 specimen
Insecta - Neuroptera - Palaeoleontidae

	Baisopardus cryptohymen n. sp. Heads et al. 2005	Heads et al. 2005	1 specimen
	SMNS 65470 (2 measurements)

	Parapalaeoleon magnus n. gen., n. sp. Menon and Makarkin 2008	Menon and Makarkin 2008	1 specimen
	SMNS 66000/268 (1 measurement)
Insecta - Neuroptera - Araripeneuridae

	Cratoneura dividens Martins-Neto 1994	Makarkin et al. 2018	1 specimen
	SMNS 66000/265

	Araripeneura sp. Martins-Neto and Vulcano 1989	Makarkin et al. 2018	1 specimen
	SMNS 66000/262

	Araripeneura regia Martins-Neto and Vulcano 1989	Makarkin et al. 2018	1 specimen
	SMNS 66000/259
Insecta - Neuroptera - Myrmeleontidae

	Pseudonymphes ? brunherottae Martins-Neto 1994	Makarkin et al. 2018	1 specimen
	SMNS 66000/256
Insecta - Neuroptera - Babinskaiidae

	Neliana maculata (Martins-Neto and Vulcano 1989)	Makarkin et al. 2017	1 specimen
	SMNS 66000/257
Insecta - Neuroptera - Ithonidae

	Principiala n. gen. incerta Makarkin and Menon 2007		1 specimen
	SMNS 66000/255 (3 measurements)
Insecta - Neuroptera - Mesochrysopidae

	Karenina leilana n. sp. Makarkin and Menon 2005	Makarkin and Menon 2005	1 specimen
	SMNS 65506

	Karenina longicollis n. sp. Makarkin and Menon 2005	Makarkin and Menon 2005	1 specimen
	SMNS 65505

	Triangulochrysopa formosa n. sp. Menon and Makarkin 2008	Menon and Makarkin 2008	1 specimen
	SMNS 66000/271 (3 measurements)
Insecta - Neuroptera - Osmylidae

	Nuddsia longiantennata n. gen., n. sp. Menon and Makarkin 2008	Menon and Makarkin 2008	1 specimen
	SMNS 66000/263 (2 measurements)
Insecta - Megaloptera - Corydalidae

	Cratocorydalopsis brasiliensis n. gen., n. sp. Jepson and Heads 2016	Jepson and Heads 2016	1 specimen
	SMNS 66000-247 (Megaloptera, fig. 11.69b in Martins-Neto et al., 2007) (2 measurements)

	Lithocorydalus fuscata n. gen., n. sp. Jepson and Heads 2016	Jepson and Heads 2016	1 specimen
	SMNS 66000-248 (Megaloptera, fig. 11.69c in Martins-Neto et al., 2007) (2 measurements)
Insecta - Coleoptera - Hydrophilidae

	Hydrophilidae indet. Latreille 1802	Wolf-Schwenninger and Schawaller 2007	1 specimen
	SMNS 66446
Insecta - Coleoptera - Scarabaeidae

	Scarabaeidae indet. (Latreille 1802)	Wolf-Schwenninger and Schawaller 2007	1 specimen
	SMNS 66458
Insecta - Coleoptera - Staphylinidae

	Staphylinidae indet. (Latreille 1802)	Wolf-Schwenninger and Schawaller 2007	1 specimen
	SMNS 66452 (2 measurements)
	= Apticax volans n. gen., n. sp. Schomann and Solodovnikov 2012	Schomann and Solodovnikov 2012

	Apticax solidus n. sp. Schomann and Solodovnikov 2012	Schomann and Solodovnikov 2012	1 specimen
	SMNS 66453 (2 measurements)
Insecta - Coleoptera - Mastigocoleidae

	Mastigocoleus rhinoceros n. sp. Tihelka and Cai 2022	Tihelka et al. 2022	1 specimen
	SMNS 66552 (2 measurements)
Insecta - Coleoptera - Elateridae

	Elateridae indet. Leach 1815	Wolf-Schwenninger and Schawaller 2007	1 specimen
	SMNS 66465
Insecta - Coleoptera - Buprestidae

	Buprestidae indet. (Leach 1815)	Wolf-Schwenninger and Schawaller 2007	1 specimen
	SMNS 66461
Insecta - Coleoptera - Peltidae

	Peltinae indet. Kirby 1837	Wolf-Schwenninger and Schawaller 2007	1 specimen
	SMNS 66467
Insecta - Coleoptera - Cucujidae

	Cucujidae indet. (Latreille 1802)	Wolf-Schwenninger and Schawaller 2007	2 specimens
	SMNS 66468, 66469
Insecta - Coleoptera - Chrysomelidae

	Eumolpinae indet. (Hope 1840)	Wolf-Schwenninger and Schawaller 2007	1 specimen
	SMNS 66471
Insecta - Coleoptera - Nemonychidae

	? Oxycoryninae indet. (Schönherr 1840)	Wolf-Schwenninger and Schawaller 2007	1 specimen
	SMNS 66553 Cratonemonyx martinsnetoi
	= Cratonemonyx martinsnetoi Legalov 2014	Gratshev and Legalov 2014
Insecta - Coleoptera - Attelabidae

	Rhynchitinae indet. (Gistel 1848)	Wolf-Schwenninger and Schawaller 2007	1 specimen
	SMNS 66449
Insecta - Coleoptera - Nitidulidae

	Nitidulidae indet. (Latreille 1802)	Wolf-Schwenninger and Schawaller 2007	1 specimen
	SMNS 66470
Insecta - Coleoptera - Lymexylidae

	Cratoatractocerus grimaldii n. gen., n. sp. Wolf-Schwenninger 2011	Wolf-Schwenninger 2011	1 specimen
	SMNS-66534 (1 measurement)
Insecta - Coleoptera - Tenebrionidae

	? Tenebrionidae indet. (Latreille 1802)	Wolf-Schwenninger and Schawaller 2007	1 specimen
	SMNS 66472
Insecta - Coleoptera - Ommatidae

	Ommatidae indet. Sharp and Muir 1912	Wolf-Schwenninger and Schawaller 2007	1 specimen
	SMNS 66456
Insecta - Coleoptera - Cupedidae

	Cupedidae indet. Laporte 1836	Wolf-Schwenninger and Schawaller 2007	1 specimen
	SMNS 66552
Insecta - Alienoptera - Alienopteridae

	Vcelesvab cratocretokrat n. gen., n. sp. Vršanský et al. 2018	Vršanský et al. 2018	1 specimen
	SNMS 66645
Insecta - Alienoptera - Umenocoleidae

	Ponopterix axelrodi Vršanský and Grimaldi 1999	Martill et al. 2007	6 specimens
	SMNS 66326, 66329, 66334-36, 66338

	Ponopterix maxima n. sp. Bechly 2007	Martill et al. 2007	13 specimens
	SMNS 66332, 66323, 66324, 66325, 66327, 66328, 66330, 66331, 66332, 66333, 66337, 66562, 67574

	Cratovitisma oldreadi n. gen., n. sp. Bechly 2007	Martill et al. 2007	1 specimen
	SMNS 66000-127
Insecta - Mantodea

	Cretophotina santanensis n. sp. Lee 2014	Lee 2014	1 specimen
	SMNS 67583 (1 measurement)
Insecta - Mantodea - Santanmantidae

	Santanmantis axelrodi Grimaldi 2003	Grimaldi 2003	5 specimens
	SMNS 112-115, 172 (112 is now SMNS 66677; Lee 2014) (10 measurements)
Insecta - Blattodea

	Mariconitermes talicei Fontes and Vulcano 1998	Martill et al. 2007	2 specimens
	SMNS 66193, 66194 (2 measurements)

	Meiatermes hariolus Grimaldi 2008	Grimaldi et al. 2008	1 specimen
	SMNS 66190 (part of Nordestinatermes araripena in Bechly, 2007) (1 measurement)

	Nordestinatermes araripena (Krishna 1990)	Martill et al. 2007	2 specimens
	SMNS 66191,66192 (2 measurements)

	Cretatermes pereirai Fontes and Vulcano 1998	Martill et al. 2007	1 specimen
	SMNS 66517
Insecta - Blattodea - Kalotermitidae

	Cratokalotermes santanensis n. gen., n. sp. Bechly 2007	Martill et al. 2007	1 specimen
	SMNS 66195 (3 measurements)
Insecta - Blattodea

	Cretarhinotermes novaolindense n. gen., n. sp. Bechly 2007	Martill et al. 2007	2 specimens
	SMNS 66196, 66197 (2 measurements)
Insecta - Blattodea - Cratomastotermitidae

	Cratomastotermes wolfschwenningeri n. gen., n. sp. Bechly 2007	Martill et al. 2007	4 specimens
	SMNS 66186-66188, ?66189 (4 measurements)
Insecta - Blattodea - Mesoblattinidae

	Blattidae informal indet. B	Martill et al. 2007	4 specimens
	SMNS 66321, 66308, 66309
	= Perlucipecta santanensis n. sp. Lee 2016	Lee 2016
	Also SMNS 66311
Insecta - Blattodea - Ectobiidae

	Blattellidae informal indet. A	Martill et al. 2007	6 specimens
	SMNS 66313, 66317, 66318
	= Piniblattella magna n. sp. Lee 2016	Lee 2016
	Also SMNS 66000-116, 66310, 66320

	Mesoblattina limai Pinto and Purper 1986	Martill et al. 2007	5 specimens
	SMNS 66312, 66314, 66315, 66319, 66322
Insecta - Dermaptera - Anisolabididae

	Kotejalabis haeuseri n. sp. Haas 2007	Martill et al. 2007	1 specimen
	SMNS 66557

	Kotejalabis goethitica Engel and Chatzimanolis 2005	Martill et al. 2007	1 specimen
	SMNS 66515

	Cratoborellia gorbi n. gen., n. sp. Haas 2007	Martill et al. 2007	1 specimen
	SMNS 66556
Insecta - Dermaptera

	Caririlabia berghoffi n. sp. Haas 2007	Martill et al. 2007	1 specimen
	SMNS 66555
Insecta - Chresmodidae

	Chresmoda neotropica n. sp. Engel and Heads 2008	Delclòs et al. 2008	1 specimen
	SMNS 66000-13 (2 measurements)
Insecta - Orthoptera - Proscopiidae

	Eoproscopia martilli n. gen., n. sp. Heads 2008	Heads 2008	1 specimen
	SMNS 66000-135 (2 measurements)
Insecta - Blattaria - Blattulidae

	Ocelloblattula santanensis n. sp. Lee 2016	Lee 2016	1 specimen
	SMNS 66558

	Elisama brevis (Mendes 2000)	Lee 2016	10 specimens
	SMNS 67563, 67564, 67565, 67566, 67567, 67568, 67569, 67570, 67571, 67572

	Elisama americana Vršanský 2002	Martill et al. 2007	2 specimens
	SMNS 66000-125, 66316

	Elisama hindwingnii n. sp. Lee 2016	Lee 2016	1 specimen
	SMNS 67573
Insecta - Hemiptera - Hydrometridae

	Hydrometridae indet. (Billberg 1820)	Perez Goodwyn 2002	1 specimen
	SMNS 64654 (old numbers 218 and 147)
	= Christometra paradoxa Pêgas et al. 2017	Pêgas et al. 2017

	Incertametra santanensis n. gen., n. sp. Perez Goodwyn 2002	Perez Goodwyn 2002	2 specimens
	SMNS 64652, 64653 (3 measurements)
Insecta - Hemiptera - Gelastocoridae

	Cratonerthra estevezae n. sp. López Ruf and Goodwyn 2005	López Ruf et al. 2005	1 specimen
	SMNS 65416 (1 measurement)
Insecta - Hemiptera

	Cratonepa enigmatica n. gen., n. sp. Jattiot et al. 2012	Jattiot et al. 2012	1 specimen
	SMNS 66380 (2 measurements)
Insecta - Hemiptera - Belostomatidae

	Neponymphes godoii Zamboni 2001	Popov and Bechly 2007	1 specimen
	SMNS 66593
Insecta - Hemiptera - Palaeontinidae

	Baeocossus fortunatus n. sp. Menon and Heads 2005	Menon and Heads 2005	2 specimens
	SMNS 65546, 65547 (3 measurements)

	Colossocossus bechlyi n. sp. Menon and Heads 2005	Menon and Heads 2005	1 specimen
	SMNS 65548 (1 measurement)

	Colossocossus rugosa n. sp. Menon et al. 2005	Menon et al. 2005	1 specimen
	SMNS 65503 (1 measurement)

	Colossocossus loveridgei n. gen., n. sp. Menon et al. 2005	Menon et al. 2005	1 specimen
	SMNS 65502 (1 measurement)

	Parawonnacottella penneyi n. sp. Menon et al. 2005	Menon et al. 2005	1 specimen
	SMNS 65504 (1 measurement)
Insecta - Hemiptera - Tettigarctidae

	Tettagalma striata n. gen., n. sp. Menon 2005	Menon 2005	1 specimen
	SMNS 65507 (2 measurements)
Insecta - Odonata - Thaumatoneuridae

	Euarchistigma peterknobli n. sp. Bechly 2010	Bechly 2010	1 specimen
	SMNS 66823 (4 measurements)
Insecta - Odonata - Protoneuridae

	Eoprotoneura hyperstigma Carle and Wighton 1990	Martill et al. 2007	1 specimen
	SMNS 66386
Insecta - Odonata - Hemiphlebiidae

	Parahemiphlebia mickoleiti Bechly 1998	Bechly 1998	1 specimen
	SMNS 63072
Insecta - Odonata

	? Cretarchistigma essweini n. sp. Bechly 1998	Bechly 1998	1 specimen
	SMNS 63071 (7 measurements)

	Santanagrion longipes n. gen., n. sp. Bechly 2010	Bechly 2010	1 specimen
	SMNS 66803b (2 measurements)
Insecta - Odonata - Aeschnidiidae

	Wightonia araripina Carle and Wighton 1990	Martill et al. 2007	2 specimens
	SMNS 66610,66611

	Santanoptera gabbotti Martill and Nel 1996	Martill et al. 2007	1 specimen
	SMNS 66609
Insecta - Odonata - Megaphlebiidae

	Megaphlebia rayandressi n. gen., n. sp. Bechly 2010	Bechly 2010	1 specimen
	SMNS 66617 (old R6) (1 measurement)
Insecta - Odonata - Proterogomphidae

	Paracordulagomphus aberrans Bechly 2010	Bechly 2010	1 specimen
	SMNS 66614

	Cordulagomphus (Procordulagomphus) xavieri (Nel and Escuillé 1994)	Martill et al. 2007	1 specimen
	SMNS 66391

	Cordulagomphus winkelhoferi n. sp. Bechly 2007	Martill et al. 2007	1 specimen
	SMNS 66607 (old M58) (2 measurements)

	Cordulagomphus hanneloreae n. sp. Bechly 2007	Martill et al. 2007	1 specimen
	SMNS 66591 (old O21) (2 measurements)

	Cordulagomphus fenestratus Carle and Wighton 1990	Martill et al. 2007	1 specimen
	SMNS C13

	Cratogomphus erraticus n. gen., n. sp. Bechly 2010	Bechly 2010	1 specimen
	SMNS 66606a/b (old L43/L17) (4 measurements)

	? Araripegomphus informal n. sp. Nel and Paicheler 1994	Bechly 1998	1 specimen
	SMNS 63070
	= Cratogomphus cf. erraticus Bechly 2010	Bechly 2010
	Probably also belongs to this new taxon
Insecta - Odonata - Cretapetaluridae

	Cratopetalura petruleviciusi Nel and Bechly 2009	Bechly 2010	1 specimen
	SMNS 66567 (1 measurement)
Insecta - Odonata - Cratopetaliidae

	Cratopetalia whiteheadi n. gen., n. sp. Bechly 2010	Bechly 2010	1 specimen
	SMNS 66389 (2 measurements)
Insecta - Odonata - Araripephlebiidae

	Araripephlebia mirabilis Bechly 1998	Martill et al. 2007	1 specimen
	SMNS 66618
Insecta - Odonata - Gomphidae

	Cratolindenia knuepfae n. gen., n. sp. Bechly 2000	Bechly 2000	1 specimen
	SMNS 64414 (2 measurements)
Insecta - Odonata - Araripegomphidae

	Araripegomphus andreneli n. sp. Bechly 1998	Bechly 1998	1 specimen
	SMNS 63651 (2 measurements)

	Araripegomphus hanseggeri n. sp. Bechly 2000	Bechly 2000	2 specimens
	SMNS 64415, 6 (4 measurements)
Insecta - Odonata - Mesuropetalidae

	Paraeschnopsis brasiliensis n. gen., n. sp. Bechly 2010	Bechly 2010	1 specimen
	SMNS 63648 (old E100404) (1 measurement)
Insecta - Odonata - Gomphaeschnidae

	Paramorbaeschna araripensis n. gen., n. sp. Bechly et al. 2001	Bechly et al. 2001	2 specimens
	SMNS 63068, 64218 (3 measurements)

	Gomphaeschnaoides magnus Bechly et al. 2001	Martill et al. 2007	1 specimen
	SMNS 64344

	Gomphaeschnaoides obliquus (Wighton 1987)	Martill et al. 2007	1 specimen
	SMNS 63069

	Gomphaeschna obliqua Wighton 1987	Bechly et al. 2001	1 specimen
	SMNS 63069 (2 measurements)
Insecta - Odonata - Liupanshaniidae

	Paramesuropetala gigantea Bechly et al. 2001	Bechly 2010	1 specimen
	SMNS 66613

	Araripeliupanshania annesuseae Bechly et al. 2001	Bechly et al. 2001	3 specimens
	SMNS 64343, 64345, 66616 (4 measurements)
Insecta - Odonata - Hageniidae

	Cratohagenius erichweberi n. gen., n. sp. Bechly 2010	Bechly 2010	4 specimens
	SMNS 66615a/b (old H71a/M71b), 66612 (old R5), 66592 (old H29), 66390 (old E100401) (11 measurements)
Insecta

	Cretereisma sp. Willmann 2007	Martill et al. 2007	10 specimens
	SMNS 66547-9, 66599-605, 66673-5

	Cretereisma schwickertorum n. sp. Willmann 2007	Martill et al. 2007	1 specimen
	SMNS 66598 (2 measurements)

	Cretereisma antiqua n. gen., n. sp. Willmann 2007	Martill et al. 2007	1 specimen
	SMNS 66546 (old 28) (2 measurements)
Insecta - Ephemeroptera

	Ephemeroptera indet. Hyatt and Arms 1890	Martill et al. 2007	2 specimens
	SMNS 66622, 66625
Insecta - Ephemeroptera - Baetiscidae

	Protobaetisca bechlyi n. gen., n. sp. Staniczek 2007	Martill et al. 2007	1 specimen
	SMNS 66620
Insecta - Ephemeroptera - Oligoneuriidae

	Incogemina nubila Storari et al. 2020	Storari et al. 2020	1 specimen
	SMNS 66623 (initially described as putative adult of Colocrus magnum)

	? Colocrus magnum n. sp. Staniczek 2007	Martill et al. 2007	1 specimen
	SMNS 66624

	Colocrus indivicum McCafferty 1990	Martill et al. 2007	2 specimens
	SMNS 66538, 66619
Insecta - Ephemeroptera - Leptophlebiidae

	? Leptophlebiidae informal sp. 1	Martill et al. 2007	1 specimen
	SMNS 66639 (species 1 of McCafferty, 1990)
Insecta - Ephemeroptera - Euthyplociidae

	Pristiplocia rupestris McCafferty 1990	Martill et al. 2007	3 specimens
	SMNS 66541 (adult), 66539, 66621 (adults)
Insecta - Ephemeroptera - Hexagenitidae

	Protoligoneuria limai Demoulin 1955	Martill et al. 2007	4 specimens
	SMNS 66536, 66537, 66635, ?66635 (putative adult)
Insecta - Ephemeroptera

	Ephemeroidea indet. Latreille 1810	Martill et al. 2007	2 specimens
	SMNS 66630, 66634
Insecta - Coxoplectoptera - Mickoleitiidae

	Mickoleitia sp. Staniczek et al. 2011	Staniczek et al. 2011	12 specimens
	putative larvae

	Mickoleitia longimanus n. gen., n. sp. Staniczek et al. 2011	Staniczek et al. 2011	1 specimen
	(3 measurements)
Arachnida - Scorpiones - Chactidae

	Araripescorpius ligabuei Campos 1986	Menon 2007	1 specimen
	SMNS 65536
Arachnida - Scorpiones - Protoischnuridae

	Protoischnurus axelrodorum de Carvalho and Lourenço 2001	Menon 2007	2 specimens
	SMNS 65534, 65535
Arachnida - Amblypygi - Phrynidae

	Britopygus weygoldti n. gen., n. sp. Dunlop and Martill 2002	Dunlop and Barov 2005	2 specimens
	SMNS 64332; 66123 (in Dunlop and Barov, 2005)
Arachnida - Thelyphonida - Thelyphonidae

	Mesoproctus rowlandi Dunlop 1998	Dunlop and Martill 2002	1 specimen
	SMNS 64331
Arachnida - Solifugae - Ceromidae

	Cratosolpuga wunderlichi Selden 1996	Dunlop and Martill 2004	2 specimens
	SMNS 65417, 65418
see common names

Geography

Country:	Brazil	State/province:	Ceará
Coordinates:	7.1° South, 39.7° West (view map)
Paleocoordinates:	5.0° South, 2.6° East (Wright 2013)
Basis of coordinate:	estimated from map
Geographic resolution:	local area

Time

Period:	Cretaceous	Epoch:	Early Cretaceous
10 m.y. bin:	Cretaceous 2-3
Key time interval:	Late Aptian
Age range of interval:	119.57 - 113.2 m.y. ago

Stratigraphy

Geological group:	Santana	Formation:	Crato
Stratigraphic resolution:	group of beds

Lithology and environment

Primary lithology:	lithified "carbonate"
Environment:	lacustrine - large	Tectonic setting:	rift

Taphonomy

Modes of preservation:	adpression
Size of fossils:	macrofossils,mesofossils
Preservation of anatomical detail:	medium

Collection methods and comments

Collection methods:	surface (in situ)
Reason for describing collection:	taxonomic analysis
Museum repositories:	SMNS
Collection method comments: Staatliches Museum fur Naturkunde, Stuttgart. Specimens were presumably purchased or donated.

Metadata

Database number:	122557
Authorizer:	M. Clapham, D. Smith	Enterer:	M. Clapham, J. Karr, A. Benabbas, E. Leckey
Modifier:	G. Varnham	Research group:	paleoentomology
Created:	2012-01-03 06:25:15	Last modified:	2025-02-22 15:12:02
Access level:	the public	Released:	2012-01-03 06:25:15
Creative Commons license:	CC0

Reference information

Primary reference:

39073.

V. N. Makarkin and F. Menon. 2007. First record of fossil "rapismatid-like" Ithonidae (Insecta, Neuroptera) from the Lower Cretaceous Crato Formation of Brazil. Cretaceous Research 28:743-753 [M. Clapham/M. Clapham]

Secondary references:

48602		N. Barling, S. W. Heads, and D. M. Martill. 2013. A new parasitoid wasp (Hymenoptera: Chalcidoidea) from the Lower Cretaceous Crato Formation of Brazil: the first Mesozoic Pteromalidae. Cretaceous Research 45:258-264 [M. Clapham/M. Clapham]
35362		G. Bechly. 1998. New fossil dragonflies from the Lower Cretaceous Crato Formation of north-east Brazil (Insecta: Odonata). Stuttgarter Beiträge zur Naturkunde Serie B (Geologie und Paläontologie) 264:1-66 [M. Clapham/M. Clapham]
40110		G. Bechly. 2000. Two new fossil dragonfly species (Insecta: Odonata: Anisoptera: Araripegomphidae and Lindeniidae) from the Crato Limestone (Lower Cretaceous, Brazil). Stuttgarter Beiträge zur Naturkunde Serie B (Geologie und Paläontologie) 296:1-16 [M. Clapham/M. Clapham]
40764		G. Bechly. 2007. Odonata: damselflies and dragonflies. In D. M. Martill, G. Bechly, R. F. Loveridge (eds.), The Crato Fossil Beds of Brazil: Window into an Ancient World 184-222 [M. Clapham/M. Clapham]
34848		G. Bechly. 2010. Additions to the fossil dragonfly fauna from the Lower Cretaceous Crato Formation of Brazil (Insecta: Odonata). Palaeodiversity 3(Supplement):11-77 [M. Clapham/M. Clapham]
56937		G. Bechly and V. N. Makarkin. 2016. A new gigantic lacewing species (Insecta: Neuroptera) from the Lower Cretaceous of Brazil confirms the occurrence of Kalligrammatidae in the Americas. Cretaceous Research 58:135-140 [M. Clapham/M. Clapham]
35206		G. Bechly, A. Nel, X. Martínez-Declòs, E. A. Jarzembowski, R. Coram, D. Martill, G. Fleck, F. Escuillié, M. M. Wisshak and M. Maisch. 2001. A revision and phylogenetic study of Mesozoic Aeshnoptera, with description of numerous new taxa (Insecta: Odonata: Anisoptera). Neue Paläontologische Abhandlungen 4:1-219 [M. Clapham/M. Clapham/M. Clapham]
41244		X. Delclòs, A. Nel, D. Azar, G. Bechly, J. A. Dunlop, M. S. Engel, and S. W. Heads. 2008. The enigmatic Mesozoic insect taxon Chresmodidae (Polyneoptera): new palaeobiological and phylogenetic data, with the description of a new species from the Lower Cretaceous of Brazil. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 247:353-381 [M. Clapham/M. Clapham/M. Clapham]
48892		J. A. Dunlop and V. Barov. 2005. A new fossil whip spider (Arachnida: Amblypygi) from the Crato Formation of Brazil. Revista Ibérica de Aracnología 12:53-62 [M. Clapham/A. Benabbas/M. Clapham]
49675		J. A. Dunlop and D. M. Martill. 2002. The first whipspider (Arachnida: Amblypygi) and three new whipscorpions (Arachnida: Thelyphonida) from the Lower Cretaceous Crato Formation of Brazil. Transactions of the Royal Society of Edinburgh: Earth Sciences 92:325-334 [M. Clapham/M. Clapham]
47429		J. A. Dunlop and D. M. Martill. 2004. Four additional specimens of the fossil camel spider Cratosolpuga wunderlichi Selden 1996 (Arachnida: Solifugae) from the Lower Cretaceous Crato Formation of Brazil. Revista Ibérica de Aracnología 9:143-156 [M. Clapham/M. Clapham]
53579		V. G. Gratshev and A. A. Legalov. 2014. The Mesozoic stage of evolution of the family Nemonychidae (Coleoptera, Curculionoidea). Paleontological Journal 48:851-944 [M. Clapham/M. Clapham]
40188		D. Grimaldi. 2003. A revision of Cretaceous mantises and their relationships, including new taxa (Insecta: Dictyoptera: Mantodea). American Museum Novitates 3412:1-47 [M. Clapham/M. Clapham]
45425		D. A. Grimaldi, M. S. Engel, and K. Krishna. 2008. The species of Isoptera (Insecta) from the Early Cretaceous Crato Formation: a revision. American Museum Novitates 3626:1-30 [M. Clapham/M. Clapham]
26790		S. W. Heads. 2008. The first fossil Proscopiidae (Insecta, Orthoptera, Eumastacoidea) with comments on the historical biogeography and evolution of the family. Palaeontology 51(2):499-507 [D. Smith/E. Leckey]
26291		S. W. Heads, D. M. Martill, and R. F. Loveridge. 2005. An exceptionally preserved antlion (Insecta, Neuroptera) with colour pattern preservation from the Cretaceous of Brazil. Palaeontology 48(6):1409-1417 [D. Smith/E. Leckey]
39191		R. Jattiot, G. Bechly, R. Garrouste and A. Nel. 2012. An enigmatic Nepoidea from the Lower Cretaceous of Brazil (Hemiptera: Heteroptera). Cretaceous Research 34:344-347 [M. Clapham/M. Clapham]
48014		R. Jattiot, L. Krogmann, and A. Nel. 2011. Revision of Prosyntexis from the Lower Cretaceous Crato Formation of Brazil (Hymenoptera: Sepulcidae: Trematothoracinae). Zootaxa 3058:55-62 [M. Clapham/M. Clapham]
59045		J. E. Jepson and S. W. Heads. 2016. Fossil Megaloptera (Insecta: Neuropterida) from the Lower Cretaceous Crato Formation of Brazil. Zootaxa 4098:134-144 [M. Clapham/M. Clapham]
62516		W. Krzemiński, K. Kopeć, and I. Kania. 2017. New and little known species from the genus Leptotarsus Guerin-Meneville, 1831 (Diptera: Tipulidae) from the Lower Cretaceous of northern Brazil. Cretaceous Research 78:103-108 [M. Clapham/M. Clapham/M. Clapham]
52423		S. W. Lee. 2014. New Lower Cretaceous basal mantodean (Insecta) from the Crato Formation (NE Brazil). Geologica Carpathica 65 [M. Clapham/M. Clapham]
61074		S. W. Lee. 2016. Taxonomic diversity of cockroach assemblages (Blattaria, Insecta) of the Aptian Crato Formation (Cretaceous, NE Brazil). Geologica Carpathica 67:433-450 [M. Clapham/M. Clapham]
62559		V. N. Makarkin, S. W. Heads, and S. Wedmann. 2017. Taxonomic study of the Cretaceous lacewing family Babinskaiidae (Neuroptera: Myrmeleontoidea: Nymphidoidae), with description of new taxa. Cretaceous Research 78:149-160 [M. Clapham/M. Clapham]
39268		V. N. Makarkin and F. Menon. 2005. New species of the Mesochrysopidae (Insecta, Neuroptera) from the Crato Formation of Brazil (Lower Cretaceous), with taxonomic treatment of the family. Cretaceous Research 26:801-812 [M. Clapham/M. Clapham]
64452		V. N. Makarkin, S. Wedmann, and S. W. Heads. 2018. A systematic reappraisal of Araripeneuridae (Neuroptera: Myrmeleontoidea), with description of new species from the Lower Cretaceous Crato Formation of Brazil. Cretaceous Research 84:600-621 [M. Clapham/M. Clapham]
40729		D. M. Martill, G. Bechly, and R. F. Loveridge. 2007. The Crato Fossil Beds of Brazil: Window into an Ancient World 1-625 [M. Clapham/M. Clapham]
48633		F. Menon. 2005. New record of Tettigarctidae (Insecta, Hemiptera, Cicadoidea) from the Lower Cretaceous of Brazil. Zootaxa 1087:53-58 [M. Clapham/M. Clapham]
49662		F. Menon. 2007. Higher systematics of scorpions from the Crato Formation, Lower Cretaceous of Brazil. Palaeontology 50:185-195 [M. Clapham/M. Clapham]
45519		F. Menon and S. W. Heads. 2005. New species of Palaeontinidae (Insecta: Cicadomorpha) from the Lower Cretaceous Crato Formation of Brazil. Stuttgarter Beiträge zur Naturkunde Serie B (Geologie und Paläontologie) 357:1-11 [M. Clapham/M. Clapham]
38323		F. Menon, S. W. Heads, and D. M. Martill. 2005. New Palaeontinidae (Insecta: Cicadomorpha) from the Lower Cretaceous Crato Formation of Brazil. Cretaceous Research 26:837-844 [M. Clapham/J. Karr]
26282		F. Menon and V. N. Makarkin. 2008. New fossil lacewings and antlions (Insecta, Neuroptera) from the Lower Cretaceous Crato Formation of Brazil. Palaeontology 51(1):149-162 [D. Smith/E. Leckey]
38252		A. Nel, G. Bechly, R. Garrouste, B. Pohl, and F. Escuillié. 2005. A new extraordinary neuropterid family from the Lower Cretaceous Crato Formation of Brazil: a new insect order? (Insecta, Neuropterida). Cretaceous Research 26:845-852 [M. Clapham/J. Karr]
67003		R. V. Pêgas, M. E. C. Leal, and J. Damgaard. 2017. A surprisingly derived hydrometrid (Hemiptera, Gerromorpha) from the Crato Formation, Early Cretaceous of Brazil. Historical Biology 30:1094-1101 [M. Clapham/M. Clapham]
11315	ETE	P. J. Perez Goodwyn. 2002. A new genus of water measurer from the Lower Cretaceous Crato Formation in Brazil (Insecta: Heteroptera: Gerromorpha: Hydrometridae). Stuttgarter Beiträge zur Naturkunde Serie B (Geologie und Paläontologie) 316:1-9 [C. Labandeira/A. Spencer-Lee/M. Clapham]
50813		Y. A. Popov and G. Bechly. 2007. Heteroptera: bugs. In D.M. Martill, G. Bechly, R.F. Loveridge (eds.), The Crato Fossil Beds of Brazil: Window into an Ancient World 317-328 [M. Clapham/M. Clapham]
40979		A. Schomann and A. Solodovnikov. 2012. A new genus of Staphylinidae (Coleoptera) from the Lower Cretaceous: the first fossil rove beetles from the southern hemisphere. Systematic Entomology 37:379-386 [M. Clapham/M. Clapham]
74173		A. P. Storari, T. Rodrigues, A. A. F. Saraiva and F. F. Salles. 2020. Unmasking a gap: A new oligoneuriid fossil (Ephemeroptera: Insecta) from the Crato Formation (upper Aptian), Araripe Basin, NE Brazil, with comments on Colocrus McCafferty. PLoS One 15(e0240365) [M. Clapham/M. Clapham/M. Clapham]
81748		E. Tihelka, M. A. Jäch, R. Kundrata, Y. D. Li, M. S. Engel, J. Lozano-Fernandez, D. Y. Huang and C. Y. Cai. 2022. Mastigocoleidae fam. nov., a new Mesozoic beetle family and the early evolution of Dryopoidea (Coleoptera). Insect Systematics and Diversity 6(3:3) [M. Clapham/M. Clapham]
66970		P. Vršanský, G. Bechly, Q. Q. Zhang, E. A. Jarzembowski, and T. Mlynský, L., P. Barna, M. Kúdela, D. Aristov, S. Bigalk, L. Krogmann, L. Q. Li, Q. Zhang, H. C. Zhang, S. Ellenberger, P. Müller, C. Gröhn, F. Y. Xia, K. Ueda, P. Vďačný, D. Valaška, L. Vršanská, and B. Wang. 2018. Batesian insect-insect mimicry-related explosive radiation of ancient alienopterid cockroaches. Biologia 73:987-1006 [M. Clapham/M. Clapham/M. Clapham]
51026		H. M. Wilson. 2001. First Mesozoic scutigeromorph centipede, from the Lower Cretaceous of Brazil. Palaeontology 44:489-495 [M. Clapham/M. Clapham]
51027		H. M. Wilson. 2003. A new scolopendromorph centipede (Myriapoda: Chilopoda) from the Lower Cretaceous (Aptian) of Brazil. Journal of Paleontology 77:73-77 [M. Clapham/M. Clapham]
48525		H. M. Wilson and D. M. Martill. 2001. A new japygid dipluran from the Lower Cretaceous of Brazil. Palaeontology 44:1025-1031 [M. Clapham/M. Clapham]
56282		K. Wolf-Schwenninger and W. Schawaller. 2007. Coleoptera: beetles. In D. M. Martill, G. Bechly, R. F. Loveridge (eds.), The Crato Fossil Beds of Brazil: Window into an Ancient World 340-350 [M. Clapham/M. Clapham/M. Clapham]