Tsagan Khushu [HMNS-GIN]: Maastrichtian, Mongolia
collected by HMNS-GIN 1993, 1997, 2006

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Mollusca indet. (Linnaeus 1758)
Watabe and Suzuki 2000
Reptilia
Mongolochelys sp. Khosatzky 1997
Suzuki and Narmandakh 2004 4 specimens
    = Mongolochelys efremovi Khosatzky 1997
Suzuki and Tsogtbaatar 2010
MPC 25/122: peripheral fragment; 1997; Tsagan Khushu; HMNS 97-21-465 MPC 25/123: shell fragments, right scapula-coracoid; 1997; Tsagan Khushu; HMNS 97-21-455 MPC 25/124: shell fragments; 1997; Tsagan Khushu; HMNS 97-21-457; MPC 25/152: left epiplastron; 2006; Tsagan Khushu North; HMNS 2006-04-497 MPC 25/153: right distal end of humerus; 2006; Tsagan Khushu North; HMNS 2006-04-546 MPC 25/154: 3rd and 8th cervicals, 15 caudals, right distal end of humerus, right ends and left proximal end of femur, fragments of post cranial bones; 2006; Tsagan Khushu North; HMNS 2006-04-498 MPC 25/155: skull in concretion; 2006; Tsagan Khushu North; HMNS 2006-04-499 MPC 25/156: right distal end of femur; 2006; Tsagan Khushu North; HMNS 2006-04-549 MPC 25/157: 5th to 7th cervicals, pectoral girdle in concretion; 2006; Tsagan Khushu North; HMNS 2006-04-500 MPC 25/158: left humerus; 2006; Tsagan Khushu North; HMNS 2006-04-501
Reptilia - Testudines - Lindholmemydidae
Suzuki and Narmandakh 2004 6 specimens
060905 TK TSI
Reptilia - Testudines - Trionychidae
Trionychidae indet. Gray 1825
Suzuki and Narmandakh 2004 4 specimens
Reptilia
Dinosauria indet. (Owen 1842)
digits
Reptilia - Theropoda - Elongatoolithidae
Elongatoolithidae indet. Zhao 1975
Watabe et al. 2010
Reptilia - Theropoda
Oviraptorosauria indet. Barsbold 1976
Watabe et al. 2010
Reptilia - Theropoda - Tyrannosauridae
Tarbosaurus sp. Maleev 1955
1 individual
skull elements of Tarbosaurus
Reptilia - Theropoda - Ornithomimidae
Gallimimus sp. Osmólska et al. 1972
1 specimen
femur
Reptilia - Theropoda - Dendroolithidae
Dendroolithidae indet. Zhao and Li 1988
Watabe et al. 2010
Reptilia
Pachycephalosauridae indet. Sternberg 1945
1 individual
930918 TK; partial skull = MPC-D 100/1206
    = Prenocephale sp. Maryańska and Osmólska 1974
Evans et al. 2018
Reptilia - Spheroolithidae
Spheroolithidae indet. Zhao 1979
Watabe et al. 2010
Reptilia - Hadrosauridae
Saurolophus sp. Brown 1912
060903 TK TSGT
    = Saurolophus angustirostris Rozhdestvensky 1952
Tsogtbaatar and Chinzorig 2007
Reptilia
Ankylosauria indet. (Osborn 1923)
Watabe and Suzuki 2000
Sauropoda indet. (Marsh 1878)
Watabe and Suzuki 2000 1 individual
A right ilium, femur, tibia and fibula of a sauropod
Crocodylia indet. (Owen 1842)
1 specimen
scute
unclassified
Tracheophyta indet. ()
Watabe and Suzuki 2000
97076 TK Johnson; "petrified wood"
Angiospermae
Angiospermae indet.
Watabe et al. 2010
pollen
see common names

Geography
Country:Mongolia State/province:Omnogov
Coordinates: 43.5° North, 100.4° East (view map)
Paleocoordinates:41.1° North, 89.4° East
Basis of coordinate:stated in text
Altitude:1334 meters
Geographic resolution:outcrop
Time
Period: Cretaceous Epoch: Late Cretaceous
Stage: Maastrichtian 10 m.y. bin: Cretaceous 8
Key time interval: Maastrichtian
Age range of interval: 72.1 - 66 m.y. ago
Stratigraphy
Formation:Nemegt
Stratigraphic resolution:formation
Lithology and environment
Primary lithology:planar lamination,fine carbonaceous sandstone
Secondary lithology: mudstone
Includes fossils?Y
Lithology description: "The bituminous sandstone with fine horizontal lamination was found in the part of alternation of mudstone and sandstone"; "The fossiliferous beds of this area are alternation of sandstone and mudstone layers with fluvial origin. Those beds are horizontally situated. Dinosaur bones and associated skeletons are included in those layers. There is a finely stratified mudstone lens with high contents of carbon material that yield pollen and spore."
Environment:fluvial indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils,microfossils
Collection methods and comments
Collection excludes:some genera,some microfossils
Collection methods:bulk,surface (float),surface (in situ),field collection
Reason for describing collection:general faunal/floral analysis
Collectors:HMNS-GIN Collection dates:1993, 1997, 2006
Collection method comments: HMNS-Hayashibara Museum of Natural Sciences
Taxonomic list comments:JMJPE researched here in 1993 and 1997, and collected one partial skull of a pachycephalosaur, fragments of skull elements of Tarbosaurus, petrified wood, and several forms of dinosaur eggshells. "Pollen and spore fossils" found as well.
Metadata
Also known as:Tsagan Khusku, Tsagan Usu, Tsagan Hushu
Database number:127681
Authorizer:R. Benson, M. Carrano Enterer:R. Benson, M. Carrano
Modifier:M. Carrano Research group:vertebrate
Created:2012-05-22 00:56:25 Last modified:2015-05-13 15:00:29
Access level:the public Released:2012-05-22 00:56:25
Creative Commons license:CC BY
Reference information

Primary reference:
42289. M. Watabe and S. Suzuki. 2000. Report on the Japan–Mongolia Joint Paleontological Expedition to the Gobi desert, 1993. Hayashibara Museum of Natural Sciences Research Bulletin 1:17-29 [R. Benson/R. Benson/M. Carrano]

Secondary references:

86067 D. C. Evans, S. Hayashi, K. Chiba, M. Watabe, M. J. Ryan, Y.-N. Lee, P. J. Currie, K. Tsogtbaatar, and R. Barsbold. 2018. Morphology and histology of new cranial specimens of Pachycephalosauridae (Dinosauria: Ornithischia) from the Nemegt Formation, Mongolia. Palaeogeography, Palaeoclimatology, Palaeoecology 494:121-134 [M. Carrano/M. Carrano]
41670 S. Suzuki and P. Narmandakh. 2004. Change of the Cretaceous turtle faunas in Mongolia. Hayashibara Museum of Natural Sciences Research Bulletin 2:7-14 [R. Benson/R. Benson]
44038 S. Suzuki and C. Tsogtbaatar. 2010. A catalog of Mongolochelys collected by the HMNS-MPC Joint Paleontological Expedition. Hayashibara Museum of Natural Sciences Research Bulletin 3:119-131 [R. Benson/R. Benson]
80572 K. Tsogtbaatar and T. Chinzorig. 2007. Mongolian hadrosaurids: paleobiogeography, dispersal and environmental relationships. The Korean Journal of Quaternary Research 21(2):49-53 [M. Carrano/M. Carrano]
42154 M. Watabe and S. Suzuki. 2000. Cretaceous fossil localities and a list of fossils collected by the Hayashibara Museum of Natural Sciences and Mongolian Paleontological Center Joint Paleontological Expedition (JMJPE) from 1993 through 1998. Hayashibara Museum of Natural Sciences Research Bulletin 1:99-108 [R. Benson/R. Benson]
53759 M. Watabe and S. Suzuki. 2000. Report on the Japan–Mongolia Joint Paleontological Expedition to the Gobi desert, 1997. Hayashibara Museum of Natural Sciences Research Bulletin 1:69-82 [M. Carrano/M. Carrano]
55060 M. Watabe, S. Suzuki, K. Tsogtbaatar, T. Tsubamoto, and M. Saneyoshi. 2010. Report of the HMNS-MPC Joint Paleontological Expedition in 2006. Hayashibara Museum of Natural Sciences Research Bulletin 3:11-18 [M. Carrano/M. Carrano]
55298 M. Watabe, K. Tsogtbaatar, S. Suzuki and M. Saneyoshi. 2010. Geology of dinosaur-fossil-bearing localities (Jurassic and Cretaceous: Mesozoic) in the Gobi Desert: Results of the HMNS-MPC Joint Paleontological Expedition. Hayashibara Museum of Natural Sciences Research Bulletin 3:41-118 [M. Carrano/M. Carrano]