Sample A, Titusville Site (1967 excavation): Late/Upper Pleistocene, Pennsylvania
collected 1967

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Insecta - Coleoptera - Carabidae
Elaphrus lapponicus Gyllenhal 1810
Pterostichus patruelis (Dejean 1831)
recombined as Pterostichus (Phonias) patruelis
Carabus maeander Fischer von Waldheim 1822
recombined as Carabus (Homoeocarabus) maeander
Dyschirius nigricornis Motschulsky 1844
    = Dyschirius melancholicus Putzeys 1867
Bousquet 2012
Trechus crassiscapus Lindroth 1955
recombined as Trechus (Trechus) crassiscapus
Bembidion fortestriatum (Motschulsky 1845)
recombined as Bembidion (Trepanedoris) fortestriatum
Patrobus septentrionis Dejean 1828
Insecta - Coleoptera - Helophoridae
Helophorus tuberculatus Gyllenhal 1808
Insecta - Coleoptera - Staphylinidae
Holoboreaphilus nordenskioeldi (Maklin 1878)
recombined as Holoboreaphilus nordenskioldi
Pycnoglypta sp. Thomson 1858
Pycnoglypta aptera or Pycnoglypta lurida
Pycnoglypta aptera Campbell 1983
Olophrum rotundicolle (Sahlberg 1817)
Olophrum consimile (Gyllenhal 1810)
Acidota quadrata (Zetterstedt 1837)
see common names

Geography
Country:United States State/province:Pennsylvania
Coordinates: 41.6° North, 79.6° West (view map)
Paleocoordinates:41.6° North, 79.6° West
Basis of coordinate:stated in text
Geographic resolution:hand sample
Time
Period:Quaternary Epoch:Pleistocene
Stage:Late/Upper Pleistocene 10 m.y. bin:Cenozoic 6
Key time interval:Late/Upper Pleistocene
Age range of interval:0.12900 - 0.01170 m.y. ago
Stratigraphy
Formation:Titusville Till
Lithology and environment
Primary lithology: peat
Environment:mire/swamp Tectonic setting:non-subsiding area
Geology comments: Like Zone I, analogues to the Zone III fauna occur today in forest–tundra environments of North America where mean July temperatures range from 10 to 13C.
Taphonomy
Modes of preservation:body
Size of fossils:mesofossils
Temporal resolution:time-averaged
Collection methods and comments
Collection methods:selective quarrying,sieve
Reason for describing collection:paleoecologic analysis
Collection dates:1967
Collection method comments: All specimens, including those examined by Totten and Coope in 1967, were mounted onto micropaleontological slides and reposited in the Quaternary Entomology Laboratory at North Dakota State University.
Metadata
Database number:137209
Authorizer:M. Clapham Enterer:J. Karr, M. Clapham
Modifier:M. Clapham Research group:paleoentomology
Created:2012-12-12 10:56:04 Last modified:2014-08-09 14:15:07
Access level:the public Released:2012-12-12 10:56:04
Creative Commons license:CC BY
Reference information

Primary reference:

43873. S. Cong, A. C. Ashworth, D. P. Schwert and S. M. Totten. 1996. Fossil Beetle Evidence for a Short Warm Interval near 40,000 yr B.P. at Titusville, Pennsylvania. Quaternary Research 45:216-225 [M. Clapham/J. Karr/J. Karr]

Secondary references:

50873 Y. Bousquet. 2012. Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico. ZooKeys 245:1-1722 [M. Clapham/M. Clapham]