Flat Creek 5: Lancian, Montana

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Actinopteri - Amiiformes - Amiidae
Melvius thomasi Bryant 1988
Bryant 1988 151 specimens
UCMP 123289, 150+ tooth crowns; UCMP 130386, dentary fragment
Amia fragosa Jordan 1927
Database 2005 3 elements
Actinopteri - Lepisosteiformes - Lepisosteidae
Lepisosteus sp. Agassiz 1843
Database 2005 1 element
Lepisosteus occidentalis (Leidy 1856)
Database 2005 6 elements
Actinopteri - Aspidorhynchiformes - Aspidorhynchidae
Belonostomus longirostris (Lambe 1902)
Database 2005 2 elements
Actinopteri
Teleostei informal indet. U-4
Brinkman et al. 2021 104 specimens
UCMP 230626, one abdominal centrum; UCMP 230627, one abdominal centrum; UCMP 230628, two abdominal centra
Osteoglossomorpha indet. Greenwood et al. 1966
Brinkman et al. 2021 22 specimens
= Coriops/Lopadichthys; UCMP 230612, posterior abdominal centrum
Actinopteri - Hiodontiformes - Hiodontidae
Hiodontidae informal indet. small morph
Brinkman et al. 2021 19 specimens
UCMP 230617, four centra of small size, including a first abdominal centrum
Hiodontidae informal indet. large morph
Brinkman et al. 2021 2 specimens
UCMP 230616, two centra of large size, including a first abdominal centrum
Actinopteri - Osteoglossiformes
Wilsonichthys sp. Murray et al. 2016
Murray et al. 2016 1 specimen
? Wilsonichthys sp. Murray et al. 2016
Brinkman et al. 2021 7 specimens
UCMP 230619, centra type BvA
Wilsonichthys aridinsulensis Murray et al. 2016
Brinkman et al. 2021 114 specimens
Actinopteri - Esociformes - Esocidae
Estesesox foxi Wilson et al. 1992
Wilson et al. 1992 30 specimens
UCMP 230608, centrum type NvC; UCMP 230606, centrum type NvD
Actinopteri - Salmoniformes
Esocoidei indet. Berg 1936
Wilson et al. 1992
Actinopteri
Acanthomorpha informal indet. HC-1
Brinkman et al. 2021 247 specimens
UCMP 230601, five first abdominal centra; UCMP 230591, four abdominal centra; UCMP 230599, two abdominal centra
Acanthomorpha informal indet. HC-2
Brinkman et al. 2021 111 specimens
UCMP 230593, abdominal centra
Acanthomorpha informal indet. HC-4
Brinkman et al. 2021 11 specimens
Acanthomorpha informal indet. HC-5
Brinkman et al. 2021 12 specimens
UCMP 230597, first abdominal centrum
Acanthomorpha informal indet. HC-6
Brinkman et al. 2021 1 specimen
UCMP 198876, one abdominal centrum
Actinopteri - Percopsiformes
Percopsiformes indet. Berg 1940
Brinkman et al. 2021 1 specimen
UCMP 198875, dentary
Actinopteri - Moronidae
? Moronidae indet.
Brinkman et al. 2021 1 specimen
UCMP 198873, dentary
Actinopteri - Sciaenidae
Platacodon nanus Marsh 1889
Database 2005 4 elements
UCMP 198873, tooth plate; UCMP 198872, teeth
Actinopteri
Otophysi informal indet. U-3/BvD
Brinkman et al. 2021 17 specimens
UCMP 230603, one anterior Weberian centrum (centrum type U3); UCMP 230604, one anterior Weberian centrum (centrum type U3); UCMP 230620, a third Weberian centrum (centrum type BvC); UCMP 230621, a third Weberian centrum (centrum type BvC)
Actinopteri - Gonorhynchiformes - Gonorhynchidae
Notogoneus sp. Cope 1885
Brinkman et al. 2021 3 specimens
unclassified
Galagadon nordquistae
Wynd et al. 2020 3 specimens
UCMP 287558–287560
Reptilia - Globauridae
Contogenys sloani Estes 1969
Database 2005 1 specimen
UCMP 130732
Reptilia - Chamopsiidae
Haptosphenus placodon Estes 1964
Database 2005 1 specimen
Peneteius aquilonius Estes 1969
Nydam et al. 2000 2 specimens
teiid lizard; UCMP 123325 9left maxilla), 124744 (right dentary)
Leptochamops denticulatus (Gilmore 1928)
Database 2005 1 specimen
Chamops segnis Marsh 1892
Database 2005 1 specimen
Reptilia - Xenosauridae
Exostinus sp. Cope 1873
Database 2005 2 specimens
Exostinus lancensis Gilmore 1928
Database 2005 1 specimen
Reptilia - Anguidae
Odaxosaurus piger (Gilmore 1928)
Database 2005 1 specimen
Reptilia
Champsosaurus sp. Cope 1876
Database 2005 1 specimen
Champsosaurus natator Parks 1933
Database 2005 3 specimens
Reptilia - Theropoda
Theropoda indet. Marsh 1881
Carpenter 1982 3 specimens
Reptilia - Tyrannosauridae
Aublysodon mirandus Leidy 1868
Carpenter 1982 5 specimens
Reptilia
Aves indet. Linnaeus 1758
Sankey 2008 4 specimens
UCMP 186826, 214056-58
Reptilia - Troodontidae
Troodon sp. Leidy 1856
Sankey 2008 2 specimens
UCMP 186885, 214060
    = cf. Pectinodon sp. Carpenter 1982
Larson and Currie 2013
Reptilia - Dromaeosauridae
Dromaeosauridae indet. Colbert and Russell 1969
Carpenter 1982 3 specimens
Richardoestesia sp. Currie et al. 1990
Database 2005 5 specimens
Richardoestesia cf. isosceles Sankey 2001
Sankey 2008 73 specimens
UCMP 213922–214006
Richardoestesia cf. gilmorei Currie et al. 1990
Larson and Currie 2013 1 specimen
UCMP 187163
Saurornitholestinae indet. Longrich and Currie 2009
Larson and Currie 2013 2 specimens
UCMP 124987, 187041
Reptilia
Paronychodon sp. Cope 1876
Database 2005 32 specimens
Paronychodon lacustris Cope 1876
Carpenter 1982 3 specimens
cf. Paronychodon lacustris Cope 1876
Sankey 2008 5 specimens
UCMP 187159, 123342, 187156, 187162, 1871464
Reptilia - Hadrosauridae
Anatosaurus sp. Lull and Wright 1942
Database 2005 1 specimen
Reptilia
Thescelosaurus sp. Gilmore 1913
Database 2005 1 specimen
Reptilia - Crocodylia
Leidyosuchus sp. Lambe 1907
Database 2005 1 specimen
    = Crocodylia indet. Owen 1842
Alroy 2007
Reptilia
Brachychampsa sp. Gilmore 1911
Database 2005 1 specimen
Brachychampsa montana Gilmore 1911
Database 2005 1 specimen
Leidyosuchus sternbergi (Gilmore 1910)
Database 2005 1 specimen
Reptilia - Testudines - Adocidae
Adocus sp. Cope 1868
Database 2005 1 specimen
Reptilia - Testudines - Trionychidae
Database 2005 1 specimen
    = Trionychidae indet. Gray 1825
Alroy 2006
Reptilia
Compsemys victa (Leidy 1856)
Database 2005 1 specimen
Mammalia - Multituberculata
Paracimexomys priscus (Lillegraven 1969)
15 specimens
(1 measurement)
Mammalia - Multituberculata - Cimolomyidae
Meniscoessus robustus (Marsh 1889)
1 specimen
Essonodon browni Simpson 1927
2 specimens
Cimolomys gracilis Marsh 1889
2 specimens
Mammalia - Multituberculata - Neoplagiaulacidae
Mesodma cf. formosa (Marsh 1889)
18 specimens
Mesodma hensleighi Lillegraven 1969
38 specimens
(1 measurement)
Mesodma thompsoni Clemens 1964
9 specimens
? Neoplagiaulax burgessi n. sp. Archibald 1982
5 specimens
Mammalia - Multituberculata - Cimolodontidae
Cimolodon nitidus Marsh 1889
10 specimens
(1 measurement)
Mammalia - Cimolestidae
Cimolestes propalaeoryctes Lillegraven 1969
1 specimen
Batodon tenuis Marsh 1892
4 specimens
Cimolestes incisus Marsh 1889
3 specimens
Cimolestes stirtoni Clemens 1973
3 specimens
Mammalia - Gypsonictopidae
Gypsonictops cf. illuminatus Lillegraven 1969
8 specimens
Mammalia - Alphadontidae
Alphadon marshi Simpson 1927
55 specimens
includes "A. wilsoni": (32 + 23) (1 measurement)
Alphadon rhaister Clemens 1966
1 specimen
    = Turgidodon rhaister Clemens 1966
Alroy 2002
Mammalia
Alphadon jasoni Storer 1991
Johanson 1996
most of the A. marshi/wilsoni sample
Mammalia - Stagodontidae
Didelphodon vorax Marsh 1889
1 specimen
(minimum; maximum 6)
Mammalia - Glasbiidae
Glasbius twitchelli n. sp. Archibald 1982
29 specimens
Mammalia
Alphadon lulli Clemens 1966
3 specimens
    = Protalphadon foxi n. sp. Johanson 1996
Johanson 1996
Mammalia - Pediomyidae
Pediomys elegans Marsh 1889
3 specimens
Pediomys krejcii Clemens 1966
6 specimens
Pediomys cf. cooki Clemens 1966
1 specimen
Pediomys hatcheri (Osborn 1898)
1 specimen
Pediomys florencae Clemens 1966
1 specimen
Amphibia - Temnospondyli - Albanerpetontidae
Albanerpetontidae indet. Fox and Naylor 1982
Gardner and DeMar 2013
Amphibia - Temnospondyli - Batrachosauroididae
Opisthotriton kayi Auffenberg 1961
Database 2005 11 specimens
Amphibia - Temnospondyli - Urodela
Habrosaurus sp. Gilmore 1928
Database 2005 1 specimen
Habrosaurus dilatus Gilmore 1928
Database 2005 3 specimens
Lisserpeton bairdi Estes 1965
Database 2005 1 specimen
Scapherpeton tectum Cope 1876
Database 2005 8 specimens
Amphibia - Temnospondyli
Scotiophryne sp. Estes 1969
Database 2005 1 specimen
Acipenseriformes - Acipenseridae
Acipenser sp. Linnaeus 1758
Database 2005 2 elements
Chondrichthyes - Hybodontiformes - Lonchidiidae
Lonchidion selachos Estes 1964
Database 2005 52 specimens
UCMP 194918, fractured tooth; UCMP 194919, fractured tooth; UCMP 123281, 37 fragmented teeth; UCMP 128740, 13 fragmented teeth
Chondrichthyes
Cristomylus leptostemma Wynd et al. 2020
Wynd et al. 2020 18 specimens
UCMP 287576
Chondrichthyes - Orectolobiformes - Brachaeluridae
Brachaelurus estesi (Herman 1977)
Database 2005 1 element
Chondrichthyes - Orectolobiformes - Ginglymostomatidae
Cantioscyllium estesi
Wynd et al. 2020 1 specimen
UCMP 287573
Chondrichthyes - Orectolobiformes - Orectolobidae
Restesia monocuspilatus Wynd et al. 2020
Wynd et al. 2020 8 specimens
UCMP 287562, 287563, 287564, 287565, 287566, 287567, 287571, 287575
Squatirhina americana Estes 1964
Database 2005 290 specimens
UCMP 194915, one complete tooth; UCMP 194916, one complete tooth; UCMP 194917, one complete tooth; UMCP 128739, 98 complete and fragmented teeth; UCMP 123283, 187 complete and fragmented teeth
Chondrichthyes - Rajiformes - Rhinobatidae
Myledaphus bipartitus Cope 1876
Database 2005 4 elements
UCMP 128742, 287561
    = Myledaphus pustulosus Cook et al. 2014
Wynd et al. 2020
Chondrichthyes - Rajiformes
Squatirhina sp. Casier 1947
Database 2005 1 element
Chondrichthyes - Rajiformes - Sclerorhynchidae
Onchosaurus avonicolus
Database 2005 1 element
see common names

Geography
Country:United States State/province:Montana County:Garfield
Coordinates: 47.5° North, 106.4° West (view map)
Paleocoordinates:54.2° North, 79.5° West
Basis of coordinate:unpublished field data
Geographic resolution:small collection
Time
Period: Cretaceous Epoch: Late Cretaceous
Stage: Maastrichtian 10 m.y. bin: Cretaceous 8
*Epoch:Senonian
Key time interval: Lancian
Age range of interval: 72.1 - 66 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Geological group:Montana Formation:Hell Creek Member:Upper
Local section:FlatC Local bed:2
Local order:bottom to top
Stratigraphic resolution:bed
Lithology and environment
Primary lithology: silty sandstone
Includes fossils?Y
Environment:terrestrial indet.
Taphonomy
Modes of preservation:body
Lagerst�tten type:concentrate
Degree of concentration:concentrated
Size of fossils:mesofossils
Disassociated major elements:many
Disassociated minor elements:many
Collection methods and comments
Collection methods:bulk,surface (float),sieve,field collection
Collection size:217 specimens
Reason for describing collection:taxonomic analysis
Museum repositories:UCMP
Taxonomic list comments:this appears to actually post-date Archibald and Clemens 1984, so their "Flat Creek" list should be ignoredupper level of Flat Creek sectionNISP 217 (minimum; maximum 222)
Metadata
Also known as:UCMP V-73087; UWBM C1614; UCMP V73087
Database number:14549
Authorizer:J. Alroy, P. Holroyd, M. Carrano Enterer:J. Alroy, P. Holroyd, M. Carrano
Modifier:M. Carrano Research group:vertebrate
Created:1993-02-18 00:00:00 Last modified:2023-08-11 16:28:53
Access level:the public Released:1993-02-18 00:00:00
Creative Commons license:CC BY
Reference information

Primary reference:
960. J. D. Archibald. 1982. A study of Mammalia and geology across the Cretaceous-Tertiary boundary in Garfield County, Montana. University of California Publications in Geological Sciences 122:1-286 [J. Alroy/J. Alroy/M. Carrano]

Secondary references:

6294 J. Alroy. 2002. Synonymies and reidentifications of North American fossil mammals. [J. Alroy/J. Alroy/M. Kouvari]
18120 J. Alroy. 2006. Synonymies and reidentifications of North American fossil vertebrates. [J. Alroy/J. Alroy/J. Alroy]
19636 J. Alroy. 2007. Synonymies and reidentifications of North American fossil vertebrates and so forth. [J. Alroy/J. Alroy]
85648 D. B. Brinkman, J. D. Divay, D. G. DeMar and G. P. Wilson Mantilla. 2021. A systematic reappraisal and quantitative study of the nonmarine teleost fishes from the late Maastrichtian of the Western Interior of North America: evidence from vertebrate microfossil localities. Canadian Journal of Earth Sciences 58(9):936-967 [M. Carrano/M. Carrano]
39778 L. J. Bryant. 1988. A new genus and species of Amiidae (Holostei; Osteichthyes) from the Late Cretaceous of North America, with comments on the phylogeny of the Amiidae. Journal of Vertebrate Paleontology 7(4):349-361 [G. Lloyd/G. Lloyd/M. Carrano]
12202ETE K. Carpenter. 1982. Baby dinosaurs from the Late Cretaceous Lance and Hell Creek formations and a description of a new species of theropod. Contributions to Geology, University of Wyoming 20(2):123-134 [M. Carrano/M. Carrano/M. Carrano]
77911 T. D. Cook, M. G. Newbrey, D. B. Brinkman and J. I. Kirkland. 2014. Euselachians from the freshwater deposits of the Hell Creek Formation of Montana. In G. P. Wilson, W. A. Clemens, J. R. Horner, J. H. Hartman (eds.), Through the End of the Cretaceous in the Type Locality of the Hell Creek Formation in Montana and Adjacent Areas. Geological Society of America Special Paper 503:229-246 [M. Carrano/M. Carrano]
13103 U.C.M.P. Database. 2005. UCMP collections database. University of California Museum of Paleontology [M. Carrano/M. Carrano/M. Carrano]
65027 J. D. Gardner and D. G. DeMar. 2013. Mesozoic and Palaeocene lissamphibian assemblages of North America: a comprehensive review. Palaeobiodiversity and Palaeoenvironments 93:459-515 [P. Holroyd/P. Holroyd/M. Carrano]
2036 Z. Johanson. 1996. Revision of the Late Cretaceous North American marsupial genus Alphadon. Palaeontographica Abteilung A 242:127-184 [J. Alroy/J. Alroy/M. Uhen]
48570 D. W. Larson and P. J. Currie. 2013. Multivariate analyses of small theropod dinosaur teeth and implications for paleoecological turnover through time. PLoS ONE 8(1):e54329:1-14 [M. Carrano/M. Carrano/M. Carrano]
64014 A. M. Murray, M. G. Newbrey, A. G. Neuman and D. B. Brinkman. 2016. New articulated osteoglossomorph from Late Cretaceous freshwater deposits (Maastrichtian, Scollard Formation) of Alberta, Canada. Journal of Vertebrate Paleontology 1-14 [P. Holroyd/P. Holroyd/P. Holroyd]
31563ETE R. L. Nydam and B. M. Fitzpatrick. 2009. The occurrence of Contogenys-like lizards in the Late Cretaceous and Early Tertiary of the Western Interior of the U.S.A. Journal of Vertebrate Paleontology 29(3):677-701 [M. Carrano/M. Carrano/P. Mannion]
6832 R. L. Nydam, J. A. Gauthier, and J. J. Chiment. 2000. The mammal-like teeth of the Late Cretaceous lizard Peneteius aquilonius Estes 1969 (Squamata, Teiidae). Journal of Vertebrate Paleontology 20(3):628-631 [J. Alroy/J. Alroy/J. Alroy]
34479ETE J. T. Sankey. 2008. Diversity of latest Cretaceous (late Maastrichtian) small theropods and birds: teeth from the Lance and Hell Creek formations, USA. In J. T. Sankey & S. Baszio (ed.), Vertebrate Microfossil Assemblages: Their Role in Paleoecology and Paleobiogeography. Indiana University Press, Bloomington 117-134 [M. Carrano/M. Carrano]
45321 M. V. H. Wilson, D. B. Brinkman, and A. G. Neuman. 1992. Cretaceous Esocoidei (Teleostei): early radiation of the pikes in North American fresh waters. Journal of Paleontology 66(5):839-846 [M. Bell/M. Bell]
84871 B. M. Wynd, D. G. DeMar, and G. P. Wilson. 2020. Euselachian diversity through the uppermost Cretaceous Hell Creek Formation of Garfield County, Montana, USA, with implications for the Cretaceous-Paleogene mass extinction in freshwater environments. Cretaceous Research 113:104483 [M. Carrano/M. Carrano]