Swain Quarry: Torrejonian, Wyoming

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Amphibia - Temnospondyli - Batrachosauroididae
Gardner and DeMar 2013
Amphibia - Temnospondyli - Urodela
Gardner and DeMar 2013
Gardner and DeMar 2013
Reptilia - Squamata
Sullivan 1982 76 specimens
"indeterminate Sauria" AMNH 12041, ventral part of basicranium; AMNH 12046, 14301, 14322, 14323, 14324, 14325, 14326, 15895, 15896, 15897, 15903, 15904, 15906, 15921, 15922, 15924, 15927, 15930, 15941, 15942, 15944, 15948, 15968, 15997, 15998, 16005, 16009, 16013, 16016, 16018, 16020, 16154, 16158, 16160, 16163, 16166, 16167, 16168, 16171, 16172, 16173, 16174, 16177, 16178, 16179, 16180, 16181, 16183, 16188, 16189, 16190, 16192, 16193, (?) 16194, 16196, 16197, 16198, 16216, 16217, 16218, partial dentaries; AMNH 15932, 15934, 15939, (?) 15946, 15947, 15950, 15964, 16025, partial maxillae; AMNH 15943, 15954, 16195, tooth-bearing fragments; AMNH 15958, proximal end of femur; AMNH 15962, trunk vertebra; AMNH 16056, distal end of humerus; AMNH 16137, caudal vertebra.
Reptilia - Xantusiidae
Sullivan 1982
scincid
Gao and Fox 1996
'indistinguishable in both size and tooth form from Palaeoxantusia from the same locality'
Sullivan 1982 43 specimens
AMNH 12028, posterior part of right maxilla; AMNH 12036, posterior part of left spleniodentary; AMNH 12038, posterior half of left spleniodentary; AMNH 12039, anterior part of right maxilla; AMNH 12040, left spleniodentary; AMNH 12041, posterior part of right spleniodentary; AMNH 12049, medial part of left spleniodentary; AMNH 12064, nearly complete left spleniodentary; AMNH 12071, anterior part of right spleniodentary; AMNH 12072, anterior part of right spleniodentary; AMNH 12073, posterior part of right spleniodentary; AMNH 12074, posterior part of right spleniodentary; AMNH 12075, left spleniodentary; AMNH 12076, nearly complete right spleniodentary; AMNH 12077, posterior part of right spleniodentary; AMNH 12079, anterior part of right spleniodentary; AMNH 12080, posterior part of right spleniodentary; AMNH 12083 (Text-fig. 6), posterior part of left maxilla; AMNH 14302, medial part of right spleniodentary; AMNH 14312, 4 teeth; AMNH 14313, medial part of right spleniodentary; AMNH 14314, posterior part of right maxilla; AMNH 15907, medial part of left spleniodentary; AMNH 15917, medial part of left spleniodentary; AMNH 15920, greater anterior part of left spleniodentary; AMNH 15926, posterior part of left maxilla; AMNH 15928, medial part of left spleniodentary; AMNH 15 929, greater anterior part of right spleniodentary; AMNH 15938, posterior part of left maxilla; AMNH 15949, posterior part of right spleniodentary; AMNH 15952, posterior part of right maxilla; AMNH 15953, medial part of left spleniodentary; AMNH 16007, nearly complete left spleniodentary; AMNH 16156, medial part of left spleniodentary; AMNH 16157, anterior part of left spleniodentary; AMNH 16170, anterior part of left spleniodentary. Also referred tentatively: AMNH 12043, medial part of right spleniodentary; AMNH 12059, medial part of right spleniodentary; AMNH 15925, nearly complete left maxilla; AMNH 15955, incomplete medial part of right spleniodentary; AMNH 16049, spleniodentary fragment; AMNH 16159, medial part of left spleniodentary; AMNH 16182, incomplete posterior part of right maxilla.
Reptilia - Iguanidae
Sullivan 1982 5 specimens
AMNH 12082 (holotype), medial part of right dentary; 12081, medial part of right dentary; 15902, medial part of left dentary; 15933, medial part of left dentary; 12048, medial part of left maxilla
Reptilia - Xenosauridae
Sullivan 1982 4 specimens
AMNH 14315, medial part of right dentary; AMNH 14316, medial part of left dentary; AMNH 14317, medial part of left dentary; AMNH 14318, anterior part of left dentary.
Reptilia - Necrosauridae
Sullivan 1982 3 specimens
AMNH 14306, medial part of left dentary; AMNH 14307, medial part of left dentary; AMNH 14309, medial part of right dentary
Reptilia - Anguidae
Anguidae indet. Gray 1825
Sullivan 1982 6 specimens
AMNH 12052, 12053, 12056, 12057, 12058, 12093, partial dentaries.
Sullivan 1982 32 specimens
AMNH 12047, medial part of left dentary; AMNH 12063, posterior part of right maxilla; AMNH 12066, medial part of right dentary; AMNH 12068, anterior part of right maxilla; AMNH 12087, left frontal; AMNH 12094, medial part of left dentary; AMNH 12095, posterior part of right maxilla; AMNH 12097, anterior part of left maxilla; AMNH 12099, nearly complete right dentary; AMNH 14311, posterior part of right dentary; AMNH 15911, left dentary; AMNH 15914, posterior part of left dentary; AMNH 16100, nearly complete right frontal. Also tentatively referred: AMNH 12034, posterior part of left maxilla; AMNH 12050, greater part of left dentary; AMNH 12051, posterior part of right dentary; AMNH 12055, medial part of left dentary; AMNH 12092, posterior part of right dentary; AMNH 14303, greater part of right dentary; AMNH 14327, posterior part of right maxilla; AMNH 15898, anterior part of left dentary; AMNH 15899, left maxilla; AMNH 15908, posterior part of a left maxilla; AMNH 15909, greater part of right maxilla; AMNH 15913, posterior part of left maxilla; AMNH 15931, posterior part of left dentary; AMNH 15935, anterior part of right maxilla; AMNH 15936, maxilla fragment; AMNH 15951, posterior part of right maxilla; AMNH 16008, posterior part of left dentary; AMNH 16010, posterior part of right maxilla.
Odaxosaurinae indet. Sullivan 1982
Sullivan 1982 13 specimens
AMNH 12089, 21 body osteoderms; AMNH 15910, posterior part of right maxilla; AMNH 15916, right maxilla fragment with tooth; AMNH 15937, posterior part of right maxilla; AMNH 15956, posterior part of left maxilla; AMNH 16004, medial part of right dentary; AMNH 16011, medial part of right dentary; AMNH 16012, anterior part of right dentary; AMNH 16109, frontal fragment; AMNH 16124 and 16125, body osteoderms, associated with frontal fragments TEXT-FIG. 11-Glyptosaurinae, genus and species indeterminate (AMNH 12088), right pterygoid (palatal view), x10. and garscales; AMNH 16176, maxilla fragment; AMNH 16209, (?) cheek osteoderm.
Sullivan 1982 8 specimens
AMNH 12031, posterior part of right maxilla; AMNH 12042, medial part of left dentary; AMNH 12044, premaxilla; AMNH 12045, anterior part of left dentary; AMNH 12065, medial part of right dentary; AMNH 12070, anterior part of right maxilla; AMNH 12088, tooth bearing region of right pterygoid; AMNH 12091, medial part of (?) right maxilla
ElShafie 2024 1 specimen
CM 87168
Sullivan 1982 14 specimens
AMNH 12030, posterior part of right dentary; AMNH 12032, right dentary; AMNH 12033, medial part of right dentary; AMNH 12035, posterior part of right dentary; AMNH 12060, medial part of left dentary; AMNH 12084, medial part of left dentary; AMNH 12085, posterior part of left dentary; AMNH 12086, posterior part of left maxilla; AMNH 14310, medial part of left dentary; AMNH 15900, posterior part of left maxilla; AMNH 15901, posterior part of right maxilla; AMNH 15940, posterior part of right dentary. Also tentatively referred: AMNH 15905, posterior part of right dentary; AMNH 15918, medial part of right dentary.
Reptilia
Sullivan 1982 2 specimens
AMNH 15960, trunk vertebra; AMNH 16003, incomplete trunk vertebra.
Sullivan 1982 1 specimen
AMNH 15957, medial part of left dentary with 3 tooth bases.
Gao and Fox 1996
Reptilia - Baenidae
Gaffney 1972 2 specimens
AMNH 4950, anterior portion of skull roof; 4949, posterior portion of skull roof
Mammalia - Multituberculata - Ptilodontidae
Rigby 1980
Krause 1987
one specimen of "Ptilodus mediaevus"
Mammalia - Multituberculata - Neoplagiaulacidae
Sloan 1987
N. "cf. hunteri"
Rigby 1980
Sloan 1981
Rigby 1980
Sloan 1981
may include Rigby's new "Ectypodus sp.," said to be the same as the Bangtail species by Gingerich et al. 1983, who do not cite Sloan (1 measurement)
Rigby 1980
Mammalia
Rigby 1980
Johanson 1996
said to be new by Rigby
Mammalia - Condylarthra - Hyopsodontidae
Robison 1986
said to be new and not the same as "L. mantiensis", a jun. syn. of L. disjunctus
Rigby 1980
Mammalia - Deltatheridia
Deltatheridia indet. Van Valen 1965
Rigby 1980
Mammalia - Insectivora
Rigby 1980
    = Insectivora indet. Bowdich 1821
Alroy 2002
Mammalia - Primates - Picrodontidae
Rigby 1980
Mammalia - Primates - Paromomyidae
Rigby 1980
confirmed by Gunnell 1989
Paromomys maturus Gidley 1923
Rigby 1980
confirmed by Gunnell 1989
Mammalia - Primates - Palaechthonidae
Palenochtha cf. minor (Gidley 1923)
Rigby 1980
    = Palenochtha minor Gidley 1923
Gunnell 1989
Rigby 1980
(1 measurement)
Rigby 1980
(7 measurements)
Mammalia - Mixodectidae
Rigby 1980
Rigby 1980
confirmed by Gunnell 1989
Mammalia - Nyctitheriidae
Rigby 1980
Mammalia
Rigby 1980
Mammalia - Pantolestidae
Pantolestinae indet. Simpson 1937
Rigby 1980
Rigby 1980
    = Bessoecetor septentrionalis Russell 1929
Scott et al. 2002
Mammalia - Procreodi - Arctocyonidae
Rigby 1980
Rigby 1980
(1 measurement)
Chriacus pelvidens (Cope 1881)
Rigby 1980
Rigby 1980
(2 measurements)
Mammalia - Phenacodontidae
Rigby 1980
Mammalia - Periptychidae
Rigby 1980
Rigby 1980
Rigby 1980
Rigby 1980
Mammalia - Acreodi - Oxyclaenidae
Rigby 1980
Rigby 1980
Mammalia - Acreodi - Triisodontidae
Rigby 1980
Mammalia - Acreodi - Mesonychidae
Rigby 1980
Mammalia - Viverravidae
Rigby 1980
Rigby 1980
Rigby 1980
Rigby 1980
macfaddeni
Mammalia - Macroscelidea
Rigby 1980
Rigby 1980
Mammalia - Macroscelidea - Apheliscidae
Rigby 1980
Rigby 1980
Mammalia - Leptictidae
Rigby 1980
Rigby 1980
Rigby 1980
Mammalia - Apatemyidae
Rigby 1980
(1 measurement)
Mammalia - Cimolestidae
Rigby 1980
Rigby 1980
Mammalia - Proteutheria - Palaeoryctidae
Rigby 1980
Mammalia
Rigby 1980
see common names

Geography
Country:United States State/province:Wyoming County:Carbon
Coordinates: 41.3° North, 107.7° West (view map)
Paleocoordinates:51.3° North, 74.1° West (Wright 2013)
Basis of coordinate:stated in text
Geographic resolution:small collection
Time
Period: Paleogene Epoch: Paleocene
10 m.y. bin: Cenozoic 1
*Period:Early/Lower Tertiary *Epoch:Early/Lower Paleocene
*Local age/stage:Torrejonian
Key time interval: Torrejonian
Age range of interval: 63.8 - 60.9 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Fort Union
Stratigraphic resolution:group of beds
Lithology and environment
Primary lithology: not reported
Environment:terrestrial indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection methods:surface (in situ),field collection
Collection size:28000 specimens
Reason for describing collection:taxonomic analysis
Taxonomic list comments:there are seven other localities in the same section, but none of the material is described
Mesodma pygmaea, i.e. sp. P, not listed, but mentioned by Krause 1977
Metadata
Also known as:CM loc 217
Database number:14886
Authorizer:J. Alroy, R. Benson, P. Holroyd, P. Barrett Enterer:J. Alroy, R. Benson, P. Holroyd, T. Cleary
Modifier:P. Holroyd Research group:vertebrate
Created:1993-02-18 00:00:00 Last modified:2025-02-22 15:12:02
Access level:the public Released:1993-02-18 00:00:00
Creative Commons license:CC0
Reference information

Primary reference:
3591. R. M. West. 1970. Tetraclaenodon puercensis (Mammalia: Phenacodontidae), Goler Formation, Paleocene of California, and distribution of the genus. Journal of Paleontology 44(5):851-857 [J. Alroy/J. Alroy/M. Carrano]

Secondary references:

6294 J. Alroy. 2002. Synonymies and reidentifications of North American fossil mammals. [J. Alroy/J. Alroy/M. Kouvari]
87093 S. J. ElShafie. 2024. Body size estimation from isolated fossil bones reveals deep time evolutionary trends in North American lizards. PLoS ONE 19(1:e0296318):1-25 [P. Holroyd/P. Holroyd/P. Holroyd]
704 E. S. Gaffney. 1972. The systematics of the North American family Baenidae (Reptilia, Cryptodira). Bulletin of the American Museum of Natural History 147(5):245-312 [J. Alroy/M. Sommers/R. Benson]
37933 K. -Q. Gao and R. C. Fox. 1996. Taxonomy and evolution of Late Cretaceous lizards (Reptilia: Squamata) from western Canada. Bulletin of the Carnegie Museum of natural History 33:1-107 [R. Benson/R. Benson]
65027 J. D. Gardner and D. G. DeMar. 2013. Mesozoic and Palaeocene lissamphibian assemblages of North America: a comprehensive review. Palaeobiodiversity and Palaeoenvironments 93:459-515 [P. Holroyd/P. Holroyd/M. Carrano]
1730 G. F. Gunnell. 1989. Evolutionary History of Microsyopoidea (Mammalia, ?Primates) and the Relationship Between Plesiadapiformes and Primates. University of Michigan Papers on Paleontology 27:1-157 [J. Alroy/J. Alroy/P. Mannion]
2035 Z. Johanson. 1996. New marsupial from the Fort Union Formation, Swain Quarry, Wyoming. Journal of Paleontology 70(6):1023-1031 [J. Alroy/J. Alroy/J. Alroy]
2160 D. W. Krause. 1987. Baiotomeus, a New Ptilodontid Multituberculate (Mammalia) from the Middle Paleocene of Western North America . Journal of Paleontology 61(3):595-603 [J. Alroy/J. Alroy/J. Ju]
2905 J. K. Rigby, Jr. 1980. Swain Quarry of the Fort Union Formation, Middle Paleocene (Torrejonian), Carbon County, Wyoming: geologic setting and mammalian fauna. Evolutionary Monographs 3:1-178 [J. Alroy/J. Alroy/J. Marcot]
2930 S. F. Robison. 1986. Paleocene (Puercan-Torrejonian) mammalian faunas of the North Horn Formation, central Utah. Brigham Young University Geology Studies 33(1):87-133 [J. Alroy/J. Alroy]
3195 R. E. Sloan. 1981. Systematics of Paleocene multituberculates from the San Juan Basin, NewMexico. In S. G. Lucas, J. K. Rigby Jr., B. S. Kues (eds.), Advances in San Juan Basin Paleontology 127-160 [J. Alroy/J. Alroy/M. Uhen]
3196 R. E. Sloan. 1987. Paleocene and latest Cretaceous mammal ages, biozones, magnetozones, rates of sedimentation, and evolution. Geological Society of America Special Paper 209:165-200 [J. Alroy/J. Alroy/M. Uhen]
7573 R. M. Sullivan. 1982. Fossil lizards from Swain Quarry "Fort Union Formation," middle Paleocene (Torrejonian), Carbon County, Wyoming. Journal of Paleontology 56(4):996-1010 [J. Alroy/J. Alroy/J. Alroy]