Judson: Danian, North Dakota
collected by R. Holtzman 1977

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Bivalvia - Pholadida - Corbulidae
Bicorbula sp. Fischer 1887
Chondrichthyes - Lamniformes - Odontaspididae
Chondrichthyes - Rajiformes - Rhinobatidae
Actinopteri - Lepisosteiformes - Lepisosteidae
Actinopteri - Amiiformes - Amiidae
Actinopteri - Aspidorhynchiformes - Aspidorhynchidae
Bryant 1987 1 specimen
Amphibia - Temnospondyli - Batrachosauroididae
Reptilia - Crocodylia
Crocodylia indet. (Owen 1842)
"two crocodilians"
Reptilia
"birds"
Reptilia - Charadriiformes - Graculavidae
Graculavidae indet. Fürbringer 1888
Benson 1999 1 specimen
SMM P77.7.159
Reptilia
Reptilia - Squamata
"one or more small lizards"
Reptilia - Anguidae
Anguidae indet. Gray 1825
ElShafie 2024
Reptilia - Testudines
Testudines indet. Batsch 1788
"turtles"
Mammalia - Multituberculata - Ptilodontidae
Krause 1987
Mammalia - Multituberculata - Neoplagiaulacidae
Neoplagiaulax sp. Lemoine 1882
form "3"
Mammalia - Condylarthra
form "2"
Mammalia - Insectivora
Insectivora indet. Bowdich 1821
form "2"
Mammalia - Primates - Carpolestidae
Beard and Wang 1995
"more likely" this species
    = Carpodaptes cf. hobackensis Dorr 1952
Fox 2002
Carpodaptes cygneus (Russell 1967)
Beard 2000
Fox 2002
Mammalia - Primates - Plesiadapidae
Plesiadapis rex (Gidley 1923)
Kihm and Hartman 1997
Mammalia - Primates - Paromomyidae
Mammalia - Nyctitheriidae
Mammalia - Pantolestidae
Mammalia - Procreodi - Arctocyonidae
Arctocyon sp. Blainville 1841
Mammalia - Phenacodontidae
Thewissen 1990
    = Ectocion collinus Russell 1929
Alroy 2002
Mammalia - Carnivora - Miacidae
Viverravinae indet. Matthew 1909
Mammalia - Viverravidae
(1 measurement)
Mammalia - Macroscelidea
Gingerich 1983
Mammalia - Macroscelidea - Apheliscidae
    = Dorraletes diminutivus Dorr 1952
Alroy 2002
Mammalia - Apatemyidae
Mammalia
Mammalia - Proteutheria - Palaeoryctidae
Acipenseriformes - Acipenseridae
see common names

Geography
Country:United States State/province:North Dakota County:Morton
Coordinates: 46.9° North, 101.5° West (view map)
Paleocoordinates:55.0° North, 62.9° West (Wright 2013)
Basis of coordinate:based on political unit
Geographic resolution:small collection
Time
Period: Paleogene Epoch: Paleocene
Stage: Danian 10 m.y. bin: Cenozoic 1
*Period:Tertiary
Key time interval: Danian
Age range of interval: 66 - 61.66 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Fort Union Member:Tongue River
Local section:Tngue Local bed:2
Local order:bottom to top
Stratigraphy comments: Peppe et al. 2011: Paleomagnetic analyses of the Tongue River Member document a series of polarity reversals that can be correlated to C29n–C26r of the GPTS suggesting Danian age
Lithology and environment
Primary lithology: not reported
Environment:terrestrial indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Collection methods:field collection
Collection size:291 specimens
Reason for describing collection:general faunal/floral analysis
Collectors:R. Holtzman Collection dates:1977
Taxonomic list comments:"four distinct sites"
probably actually in the Slope Fm.: Hartman and Kihm 1991
Gunnell 1994 lists Ptilodus sp. "C"; not clear if this replaces P. wyomingensis
Metadata
Also known as:L6; L8
Database number:14962
Authorizer:J. Alroy, P. Holroyd, M. Carrano Enterer:J. Alroy, M. Carrano, P. Holroyd
Modifier:M. Carrano Research group:vertebrate
Created:1993-02-18 00:00:00 Last modified:2025-02-22 15:12:02
Access level:the public Released:1993-02-18 00:00:00
Creative Commons license:CC0
Reference information

Primary reference:
1929. R. C. Holtzman. 1978. Late Paleocene Mammals of the Tongue River Formation, Western North Dakota. North Dakota Geological Survey Report of Investigations 65:1-88 [J. Alroy/J. Alroy/M. Uhen]

Secondary references:

6294 J. Alroy. 2002. Synonymies and reidentifications of North American fossil mammals. [J. Alroy/J. Alroy/M. Kouvari]
1039 K. C. Beard. 2000. A new species of Carpocristes (Mammalia: Primatomorpha) from the middle Tiffanian of the Bison Basin, Wyoming, with notes on carpolestid phylogeny. Annals of Carnegie Museum 69(3):195-208 [J. Alroy/J. Alroy/K. Beard]
1047 K. C. Beard and J. Wang. 1995. The first Asian plesiadapoids. Annals of Carnegie Museum 64(1):1-33 [J. Alroy/J. Alroy/J. Alroy]
82793 R. D. Benson. 1999. Presbyornis isoni and oher late Paleocene birds from North Dakota. In S. L. Storrs, P. Wellnhofer, C. Mourer-Chauviré, D. W. Steadman, L. D. Martin (eds.), Avian Paleontology at the Close of the 20th Century: Proceedings of the 4th International Meeting of the Society of Avian Paleontology and Evolution, Washington, D.C, 4-7 June 1996. Smithsonian Contributions to Paleobiology 89:253-259 [M. Carrano/M. Carrano]
64908 L. J. Bryant. 1987. Belonostomus (Teleostei: Aspidorhynchidae) from the Late Paleocene of North Dakota. PaleoBios (43)1-2 [P. Holroyd/P. Holroyd]
87093 S. J. ElShafie. 2024. Body size estimation from isolated fossil bones reveals deep time evolutionary trends in North American lizards. PLoS ONE 19(1:e0296318):1-25 [P. Holroyd/P. Holroyd/P. Holroyd]
7175 R. C. Fox. 2002. The dentition and relationships of Carpodaptes cygneus (Russell) (Carpolestidae, Plesiadapiformes, Mammalia), from the late Paleocene of Alberta, Canada. Journal of Paleontology 76(5):864-881 [J. Alroy/J. Alroy/J. Alroy]
1652 P. D. Gingerich. 1983. New Adapisoricidae, Pentacodontidae, and Hyopsodontidae (Mammalia, Insectivora and Condylarthra) from the late Paleocene ofWyoming and Colorado. Contributions from the Museum of Paleontology, University of Michigan 26(11):227-255 [J. Alroy/J. Alroy/P. Mannion]
2083 A. J. Kihm and J. H. Hartman. 1997. A reevaluation of the biochronology of the Brisbane and Judson local faunas (Late Paleocene), North Dakota. Journal of Vertebrate Paleontology 17(3):57A [J. Alroy/J. Alroy/J. Ju]
2161 D. W. Krause. 1987. Systematic Revision of the Genus Prochetodon (Ptilodontidae, Multituberculata) from the Late Paleocene and Early Eocene of Western North America. Contributions from the Museum of Paleontology, University of Michigan 27(8):221-236 [J. Alroy/J. Alroy/P. Mannion]
3401 J. G. M. Thewissen. 1990. Evolution of Paleocene and Eocene Phenacodontidae (Mammalia, Condylarthra). University of Michigan Papers on Paleontology 29:1-107 [J. Alroy/J. Alroy/P. Mannion]