Bembridge Marls (A'Court Smith collection): Priabonian, United Kingdom
collected by E. J. A'Court Smith

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Insecta - Neuroptera
Neuroptera indet. Linnaeus 1758
1 specimen
NHML In.20557
Insecta - Neuroptera - Hemerobiidae
Hemerobiidae informal sp.B
1 specimen
NHML In.17173 (1 measurement)
Hemerobiidae informal sp.C
1 specimen
NHML In.17445
? Neuronema informal sp. A
1 specimen
NHML In.17404
Insecta - Neuroptera - Sisyridae
Sisyridae informal sp.A
1 specimen
NHML In.17287 (1 measurement)
    = Paleosisyra sp. Nel et al. 2003
Perkovsky and Makarkin 2015
Insecta - Coleoptera - Dytiscidae
Hydroporinae indet. (Aubé 1836)
Kirejtshuk et al. 2019 1 specimen
NHMUK In.17437 (?Oreodytes sp. or ?Hydrovatus sp.)
Insecta - Coleoptera - Ptilodactylidae
Paralichas striatopunctatus n. sp. Kirejtshuk 2019
Kirejtshuk et al. 2019 1 specimen
NHMUK In.17449 (1 measurement)
Insecta - Coleoptera - Scirtidae
Scirtinae indet. Fleming 1821
Kirejtshuk et al. 2019 1 specimen
NHMUK In.17432
Scirtes calcariferens Kirejtshuk and Ponomarenko 2019
Kirejtshuk et al. 2019 1 specimen
NHMUK In.17437(2) (1 measurement)
Contacyphon involutus Kirejtshuk and Ponomarenko 2019
Kirejtshuk et al. 2019 1 specimen
NHMUK In.17435 (1 measurement)
Insecta - Coleoptera - Staphylinidae
Staphylinidae indet. (Latreille 1802)
Kirejtshuk et al. 2019 1 specimen
NHMUK In.17384
Pselaphinae indet. (Latreille 1802)
Kirejtshuk et al. 2019 1 specimen
CAMSM X.50140.206
Insecta - Coleoptera - Scarabaeidae
Pentodon dorcus n. sp. Krell 2019
Kirejtshuk et al. 2019 1 specimen
NHMUK In.17431 (2 measurements)
Aphodius vectis n. sp. Krell 2019
Kirejtshuk et al. 2019 1 specimen
NHMUK In.17436 (2 measurements)
Insecta - Coleoptera - Hydrophilidae
Berosus barclayi Ponomarenko and Soriano 2019
Kirejtshuk et al. 2019 1 specimen
NHMUK In.17426 (1 measurement)
Hydrochara woodwardi n. sp. Ponomarenko and Soriano 2019
Kirejtshuk et al. 2019 1 specimen
NHMUK In.17424 (1 measurement)
Insecta - Coleoptera - Eucinetidae
Eucinetus nikolaevae n. sp. Kirejtshuk and Ponomarenko 2019
Kirejtshuk et al. 2019 1 specimen
NHMUK In.17286 (2 measurements)
Insecta - Coleoptera - Nitidulidae
Prometopia (Prometopia) europaea n. sp. Kirejtshuk and Kurochkin 2019
Kirejtshuk et al. 2019 1 specimen
NHMUK In.17438 (1 measurement)
Insecta - Coleoptera - Chrysomelidae
Plateumaris (Plateumaris) aff. wightensis Kurochkin and Kirejtshuk 2019
Kirejtshuk et al. 2019 2 specimens
NHMUK In.2580, 20552
Insecta - Lepidoptera
Lepidoptera informal sp.D
1 specimen
NHML In.17392 (1 measurement)
Lithopsyche antiqua n. gen., n. sp. Butler 1889
Butler 1889 1 specimen
NHML In.19984 I.10369 (2 measurements)
Insecta - Lepidoptera - Micropterigidae
Micropterix anglica n. sp. Jarzembowski 1980
1 specimen
NHML In.17411 (1 measurement)
Insecta - Trichoptera - Beraeidae
Beraeodes vectensis Cockerell 1921
Sukatsheva 2014 5 specimens
BMNH In.17290, 17351, 17393, 17397, 17499 (5 measurements)
Insecta - Hymenoptera - Braconidae
Eubazus flavistigma n. sp. Belokobylskij 2014
Antropov et al. 2014 1 specimen
BMNH In.17244 (1 measurement)
Homolobus rasnitsyni n. sp. Belokobylskij 2014
Antropov et al. 2014 1 specimen
BMNH In.20545 (1 measurement)
Bembracon acourtsmithi n. gen., n. sp. Belokobylskij 2014
Antropov et al. 2014 1 specimen
BMNH In.17224
Bassus magnareola n. sp. Belokobylskij 2014
Antropov et al. 2014 1 specimen
BMNH In.17127 (1 measurement)
? Hellenius kozlovi n. sp. Belokobylskij 2014
Antropov et al. 2014 1 specimen
BMNH In.17298 (1 measurement)
Ascogaster (Ascogaster) pygmaea n. sp. Belokobylskij 2014
Antropov et al. 2014 1 specimen
BMNH In.17379 (1 measurement)
Semionis wightensis n. sp. Belokobylskij 2014
Antropov et al. 2014 1 specimen
BMNH In.17081 (2 measurements)
Dacnusites sepultus n. gen., n. sp. Cockerell 1921
Cockerell 1921 1 specimen
NHML In. 17176 (1 measurement)
Insecta - Hymenoptera - Ichneumonidae
Ichneumonidae indet. (Latreille 1802)
Antropov et al. 2014 1 specimen
BMNH In.17430
Coleocentrus gurnetensis n. sp. Cockerell 1921
Cockerell 1921 1 specimen
NHML In. 17079 (1 measurement)
Insecta - Hymenoptera
Macroteleia veterna n. sp. Cockerell 1921
Cockerell 1921 1 specimen
NHML In. 17262 (1 measurement)
Lithobelyta reducta n. gen., n. sp. Cockerell 1921
Cockerell 1921 1 specimen
NHML In. 17091 (1 measurement)
Insecta - Hymenoptera - Vectevaniidae
Vectevania vetula n. gen., n. sp. Cockerell 1922
Cockerell 1922 1 specimen
BMNH In.20535 (2 measurements)
Insecta - Hymenoptera - Apidae
Oligobombus cuspidatus n. gen., n. sp. Antropov 2014
Antropov et al. 2014 1 specimen
BMNH In.17349 (1 measurement)
Insecta - Hymenoptera - Sphecidae
Sceliphron brevior n. sp. Cockerell 1921
Cockerell 1921 1 specimen
NHML In. 17472/24340 (Hooley collection) (1 measurement)
Insecta - Hymenoptera - Vespidae
Polybia oblita n. sp. Cockerell 1921
Cockerell 1921 1 specimen
BMNH In. 20530(17166) (1 measurement)
Insecta - Hymenoptera - Tiphiidae
Dryophia oculata n. gen., n. sp. Antropov 2014
Antropov et al. 2014 1 specimen
BMNH In.17086 (1 measurement)
Insecta - Hymenoptera - Formicidae
Oecophylla megarche Cockerell 1915
Donisthorpe 1920 12 specimens
Leucotaphus gurnetensis (Cockerell 1915)
Donisthorpe 1920 13 specimens
not In.17066, 17213
Camponotus cockerelli (Donisthorpe 1920)
Antropov et al. 2014 2 specimens
BMNH In.17280, 17066
Dolichoderus britannicus Cockerell 1915
Donisthorpe 1920 2 specimens
BMNH In.17315 (not 17311); In.17067/17074 in Antropov et al., 2014
Dolichoderus ovigerus Cockerell 1915
Donisthorpe 1920 1 specimen
BMNH In.17274
Emplastus hypolithus (Cockerell 1915)
Antropov et al. 2014 1 specimen
BMNH In.17311 (D. britannicus in Donisthorpe 1920)
Insecta - Blattodea - Rhinotermitidae
Reticulitermes sp. Holmgren 1913
8 specimens
(1 measurement)
Insecta - Blattodea - Mastotermitidae
Mastotermes anglicus n. sp. von Rosen 1913
von Rosen 1913 61 specimens
BMNH I 15037 (21 in von Rosen, 1913) (2 measurements)
Mastotermes batheri n. sp. von Rosen 1913
von Rosen 1913 1 specimen
Insecta - Blattodea - Ectobiidae
Ectobiidae informal indet. 6
Ross 2019 1 specimen
NHMUK In.17337
Phyllodromica protosardea n. sp. Ross 2019
Ross 2019 1 specimen
NHMUK I.15039/In.17142(2) (1 measurement)
Insecta - Mantodea
Protohierodula crabbi n. gen., n. sp. Ross 2019
Ross 2019 1 specimen
NHMUK In.17462
Insecta - Orthoptera - Gryllotalpidae
Pterotriamescaptor primus (Cockerell 1921)
Gorochov 2019 2 specimens
NHMUK In.17460a,b, 17461
Insecta - Orthoptera - Gryllidae
Ontogryllus rossi n. gen., n. sp. Gorochov 2019
Gorochov 2019 1 specimen
NHMUK In.25806 (1 measurement)
Insecta - Orthoptera - Tettigoniidae
Archepseudophylla melanospila (Cockerell 1921)
Nel et al. 2008 1 specimen
BMNH In.25803/In.17326
Insecta - Orthoptera - Acrididae
Proschistocerca oligocaenica Zeuner 1937
Zeuner 1937 8 specimens
BMNH In.17339, 17342, 17406, 17407, 20527; 17346, 17350, 20533 in Gorochov 2019
Insecta - Plecoptera - Leuctridae
Nemoura priscula n. sp. Cockerell 1922
Cockerell 1922 1 specimen
NHML In.17498 (2 measurements)
Insecta - Hemiptera - Miridae
Gurnardinia herczeki n. gen., n. sp. Popov 2019
Szwedo et al. 2019 1 specimen
NHMUK In.17231/In.17232 (1 measurement)
Insecta - Hemiptera - Lygaeidae
Gurnardobaya rossi Popov 2019
Szwedo et al. 2019 1 specimen
CAMSM X.50140.71 (TN122)
Insecta - Hemiptera - Cydnidae
Cydnidae indet. (Billberg 1820)
Szwedo et al. 2019 1 specimen
NHMUK In.17441b
Insecta - Hemiptera - Gerridae
Gerris sp. Fabricius 1794
Szwedo et al. 2019 1 specimen
CAMSM X.50140.44 (TN94)
Insecta - Hemiptera
Aphidoidea indet. Latreille 1802
Szwedo et al. 2019 4 specimens
NHMUK In.17198, In.17210(2,3), In.17214
Insecta - Hemiptera - Aphididae
? Colopha incognita Wegierek 2019
Szwedo et al. 2019 2 specimens
NHMUK In.17178, 17198
Insecta - Hemiptera - Cicadellidae
Typhlocybinae indet. Kirschbaum 1868
Szwedo et al. 2019 1 specimen
NHMUK In.17142(3)
Teniwitta andrewrossi n. gen., n. sp. Szwedo 2019
Szwedo et al. 2019 1 specimen
NHMUK In.20529 (1 measurement)
Insecta - Hemiptera - Aphrophoridae
Aphrophoridae informal indet. 2
Szwedo et al. 2019 1 specimen
CAMSM X.50140.145 (TN203)
Aphrophoridae informal indet. 3
Szwedo et al. 2019 1 specimen
NHMUK In.24560
Blenniphora skaka n. sp. Szwedo 2019
Szwedo et al. 2019 1 specimen
CAMSM X.50140.90 (TN145) (1 measurement)
Insecta - Hemiptera - Nogodinidae
Ambitaktoa stoumma n. gen., n. sp. Szwedo 2019
Szwedo et al. 2019 1 specimen
NHMUK In.17282 (1 measurement)
Phariberea gurdonika n. gen., n. sp. Szwedo 2019
Szwedo et al. 2019 1 specimen
NHMUK In.17096 (1 measurement)
Insecta - Hemiptera - Cixiidae
Margaxius sp. Szwedo 2019
Szwedo et al. 2019 1 specimen
NHMUK In.17369
Insecta - Thysanoptera - Aeolothripidae
Aeolothrips jarzembowskii Shmakov 2014
Shmakov 2014 1 specimen
BMNH In.17210
Sinaeolothrips brodiei (Cockerell 1917)
Shmakov 2014 1 specimen
BMNH In.17401
Insecta - Psocodea
Wightipsocus intasi Azar 2014
Azar 2014 1 specimen
BMNH In.17088
Awightipsocus megaloi n. sp. Azar 2014
Azar 2014 1 specimen
BMNH In.17187 (1 measurement)
Insecta - Odonata - Corduliidae
Corduliidae informal indet. B
Nel and Fleck 2014 1 specimen
BMNH In.17370/24763 (counterpart in Hooley coll.)
Corduliidae informal indet. C
Nel and Fleck 2014 1 specimen
BMNH In.17324/24771 (counterpart in Hooley coll.)
Insecta - Odonata - Urothemistidae
Eomacrodiplax incompleta Nel and Fleck 2014
Nel and Fleck 2014 1 specimen
BMNH In.17333
Insecta - Odonata - Aeshnidae
Aeschnophlebia andreasi Nel et al. 2005
Nel and Fleck 2014 2 specimens
BMNH In.17371, 17395
Anglogomphaeschna eocenica Nel and Fleck 2014
Nel and Fleck 2014 2 specimens
BMNH In.17368, 17330
Insecta - Odonata - Gomphaeschnidae
? Oligoaeschna anglica Cockerell and Andrews 1916
Nel and Fleck 2014 1 specimen
BMNH In.17353
Insecta - Odonata - Lestidae
Lestes aff. regina Theobald 1937
Nel and Fleck 2014 1 specimen
BMNH In.17408
Insecta - Odonata - Protoneuridae
Angloprotoneura emilielacroixi n. gen., n. sp. Nel and Fleck 2014
Nel and Fleck 2014 1 specimen
BMNH In.17246 (1 measurement)
Insecta - Odonata - Hypolestidae
Anglohypolestes fasciata n. gen., n. sp. Nel and Fleck 2014
Nel and Fleck 2014 1 specimen
BMNH In.17373 (1 measurement)
Insecta - Diptera - Stratiomyidae
Odontomyia brodiei (Cockerell 1915)
Krzemiński et al. 2019 4 specimens
NHMUK In.17221, 17253, 17356, 17478
Insecta - Diptera - Bombyliidae
Systropus acourti n. sp. Cockerell 1921
Cockerell 1921 1 specimen
BMNH I.17208/17293 (1 measurement)
Insecta - Diptera - Anisopodidae
Rhyphus hooleyi Cockerell 1921
Cockerell 1921 1 specimen
BMNH In.17136 (counterpart I.9063 in Brodie coll)
Insecta - Diptera - Bibionidae
Plecia acourti n. sp. Cockerell 1921
Cockerell 1921 1 specimen
BMNH In.17098 (1 measurement)
Insecta - Diptera - Cylindrotomidae
Cyttaromyia rossi n. sp. Krzemiński 2019
Krzemiński et al. 2019 1 specimen
CAMSM X.50140.14 (1 measurement)
Insecta - Diptera - Limoniidae
Limnophila (Limnophila) cyclospila (Cockerell and Haines 1921)
Krzemiński et al. 2019 1 specimen
NHMUK In.17491
Limnophila (Limnophila) deleta (Cockerell 1921)
Krzemiński et al. 2019 4 specimens
NHMUK In.17115, 17171, 24414, 17484/In.64127 (Hooley collection)
Pseudolimnophila (Pseudolimnophila) disjunctula (Cockerell and Haines 1921)
Krzemiński et al. 2019 1 specimen
NHMUK In.17383
Gymnastes fasciatipennis Cockerell 1921
Krzemiński et al. 2019 1 specimen
NHMUK In.17494
Gonomyia (Gonomyia) indecisa (Cockerell and Haines 1921)
Krzemiński et al. 2019 1 specimen
NHMUK In.17248
Gonomyia lutescens Cockerell 1921
Cockerell 1921 1 specimen
BMNH In.17099
Cheilotrichia (Empeda) szwedoi n. sp. Krzemiński 2019
Krzemiński et al. 2019 1 specimen
NHMUK In.17082 (1 measurement)
Dicranomyia (Dicranomyia) brodiei (Cockerell 1917)
Krzemiński et al. 2019 1 specimen
NHMUK In.17486
Mongoma cruciferella Cockerell 1917
Cockerell 1921 4 specimens
BMNH I.17259; In.17116, 17130, 17264/I.9200 (Brodie collection) in Krzeminski et al. 2020 (1 measurement)
Trentepohlia (Mongoma) pallescens (Cockerell 1921)
Krzemiński et al. 2019 2 specimens
NHMUK In.17117/I.9051 (Brodie collection), 17123
Limnobia spilota n. sp. Cockerell 1921
Cockerell 1921 1 specimen
BMNH In.17114 (1 measurement)
Insecta - Diptera - Tipulidae
Tipula callarche n. sp. Cockerell 1921
Cockerell 1921 1 specimen
BMNH In. 17381 (1 measurement)
Leptotarsus (Longurio) wegiereki n. sp. Krzemiński 2019
Krzemiński et al. 2019 3 specimens
NHMUK In.17493, CAMSM X.50140, X.50140.89a (1 measurement)
Megistocera gurnetensis n. sp. Cockerell 1921
Cockerell 1921 1 specimen
BMNH In.17090 (1 measurement)
Tipula gardneri Cockerell 1917
Krzemiński et al. 2019 1 specimen
NHMUK In.17429
Insecta - Diptera - Chironomidae
Chironomidae indet. Macquart 1838
Krzemiński et al. 2019 1 specimen
NHMUK In.17470
Orthocladiinae indet. Lenz 1921
Krzemiński et al. 2019 1 specimen
NHMUK In.17390(1)
Chironomini indet. Macquart 1838
Krzemiński et al. 2019 5 specimens
NHMUK In.17388
Insecta - Diptera - Psychodidae
Wightipsychoda leucospila (Cockerell 1921)
Krzemiński et al. 2019 1 specimen
NHMUK In.17204
Insecta - Diptera - Ceratopogonidae
Stilobezzia sp. Kieffer 1911
Krzemiński et al. 2019 2 specimens
NHMUK In.17390(2), 17471
Insecta - Diptera - Culicidae
Aedes protolepis (Cockerell 1915)
Edwards 1923 5 specimens
BMNH In. 17069, 17147, 17148, 17164, 20561
Culex protorhinus Cockerell 1915
Edwards 1923 2 specimens
BMNH In. 17138, 17146 (1 measurement)
Insecta - Diptera - Dixidae
Dixella priscula (Cockerell 1921)
Krzemiński et al. 2019 1 specimen
NHMUK In.17402 (1 measurement)
Arachnida - Araneae
Eoatypus woodwardii n. sp. McCook 1888
McCook 1888 1 specimen
BMNH In. 61271 (collection not specified)
Arachnida - Araneae - Dictynidae
Vectaraneus yulei Selden 2001
Selden 2001 1 specimen
BMNH In. 17151
Arachnida - Araneae - Araneidae
Araneidae indet. Simon 1895
Selden 2014 1 specimen
BMNH In.17177
see common names

Geography
Country:United Kingdom State/province:England County:Isle of Wight
Coordinates: 50.7° North, 1.4° West (view map)
Paleocoordinates:49.2° North, 5.2° West
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period: Paleogene Epoch: Eocene
Stage: Priabonian 10 m.y. bin: Cenozoic 3
*Period:Tertiary
*International age/stage:Priabonian - Rupellian *Local age/stage:Priabonian (minimum)
Key time interval: Priabonian
Age range of interval: 37.71 - 33.9 m.y. ago
Age estimate: 34 Ma (paleomagnetic)
* legacy (obsolete) database fields
Stratigraphy
Geological group:Solent Formation:Bouldnor Member:Bembridge Marls
Stratigraphy comments: Upper part of 13r chron just above 1n interval
Lithology and environment
Primary lithology: lithified argillaceous lime mudstone
Environment:lagoonal/restricted shallow subtidal Tectonic setting:passive margin
Geology comments: The Bembridge Marls were deposited in fresh-brackish water lakes and lagoons
Taphonomy
Modes of preservation:mold/impression
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Reason for describing collection:taxonomic analysis
Museum repositories:BMNH
Collectors:E. J. A'Court Smith
Collection method comments: Specimens kept in British Natural History Museum
Metadata
Also known as:ETE Locality 797
Database number:21976
Authorizer:C. Labandeira, M. Clapham Enterer:C. Labandeira, J. Sandlin, M. Clapham, J. Karr
Modifier:M. Uhen Research group:paleoentomology
Created:1998-06-08 00:00:00 Last modified:2018-12-12 10:58:10
Access level:the public Released:1998-06-08 00:00:00
Creative Commons license:CC BY
Reference information

Primary reference:
5087.ETE 799E. A. Jarzembowski. 1980. Fossil Insects from the Bembridge Marls, Palaeogene of the Isle of Wight, Southern England. Bulletin of the British Museum of Natural History (Geology) 33(4):237-293 [C. Labandeira/C. Labandeira/J. Karr]

Secondary references:

51861 A. V. Antropov, S. A. Belokobylskij, S. G. Compton, G. M. Dlussky, A. I. Khalaim, V. A. Kolyada, M. A. Kozlov, K. S. Perfilieva, and A. P. Rasnitsyn. 2014. The wasps, bees and ants (Insecta: Vespida=Hymenoptera) from the Insect Limestone (Late Eocene) of the Isle of Wight, UK. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 104:335-446 [M. Clapham/M. Clapham]
51855 D. Azar. 2014. Tertiary bark lice (Insecta: Psocodea) from the Insect Limestone (Bembridge Marls, Late Eocene) of the Isle of Wight, UK. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 104:307-316 [M. Clapham/M. Clapham]
44693 A. G. Butler. 1889. Description of a new genus of fossil moths belonging to the geometrid family Euschemidae. Proceedings of the Zoological Society of London 59:292-297 [M. Clapham/M. Clapham]
48513 T. D. A. Cockerell. 1921. Fossil arthropods in the British Museum - VII. The Annals and Magazine of Natural History, Ninth Series 8:541-545 [M. Clapham/M. Clapham/M. Clapham]
5006ETE 718T. D. A. Cockerell. 1921. Fossil arthropods in the British Museum - VI. Oligocene insects from Gurnet Bay, Isle of Wight. The Annals and Magazine of Natural History, Ninth Series 7:453-480 [C. Labandeira/C. Labandeira/M. Clapham]
5046ETE 758T. D. A. Cockerell. 1922. Two families of insects new to British Tertiary strata. The Canadian Entomologist 54:33-34 [C. Labandeira/C. Labandeira/M. Clapham]
5061ETE 773H. S.t. J. Donisthorpe. 1920. British Oligocene Ants. Annals and Magazine of Natural History, Series 9 6:81-94 [C. Labandeira/C. Labandeira/M. Clapham]
45384 F. W. Edwards. 1923. Oligocene mosquitoes in the British Museum, with a summary of our present knowledge concerning fossil Culicidae. Quarterly Journal of the Geological Society 79:139-155 [M. Clapham/M. Clapham/M. Clapham]
71360 A. V. Gorochov. 2019. Crickets and grasshoppers (Orthoptera) from the latest Eocene of the Isle of Wight, southern England, UK. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 110:317-329 [M. Clapham/M. Clapham/M. Clapham]
71414 A. G. Kirejtshuk, A. G. Ponomarenko, A. S. Kurochkin, A. V. Alexeev, V. G. Gratshev, A. Y. Solodovnikov, F. T. Krell and C. Soriano. 2019. The beetle (Coleoptera) fauna of the Insect Limestone (late Eocene), Isle of Wight, southern England. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 110:405-492 [M. Clapham/M. Clapham/M. Clapham]
71476 W. Krzemiński, V. Blagoderov, D. Azar, E. Lukashevich, R. Szadziewski, S. Wedmann, A. Nel, F. M. Collomb, A. Waller and D. B. Nicholson. 2019. True flies (Insecta: Diptera) from the late Eocene insect limestone (Bembridge Marls) of the Isle of Wight, England, UK. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 110:495-554 [M. Clapham/M. Clapham/M. Clapham]
47361 H. C. McCook. 1888. A new fossil spider, Eoatypus woodwardii. Proceedings of the Academy of Natural Sciences of Philadelphia 40:200-202 [M. Clapham/M. Clapham]
51852 A. Nel and G. Fleck. 2014. Dragonflies and damselflies (Insecta: Odonata) from the Late Eocene of the Isle of Wight. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 104:283-306 [M. Clapham/M. Clapham]
45276 A. Nel, J. Prokop, and A. J. Ross. 2008. New genus of leaf-mimicking katydids (Orthoptera: Tettigoniidae) from the Late Eocene–Early Oligocene of France and England. Comptes Rendus Palevol 7:211-216 [M. Clapham/M. Clapham/M. Clapham]
55565 E. E. Perkovsky and V. N. Makarkin. 2015. First confirmation of spongillaflies (Neuroptera: Sisyridae) from the Cretaceous. Cretaceous Research 56:363-371 [M. Clapham/M. Clapham/M. Clapham]
71354 A. J. Ross. 2019. The Blattodea (cockroaches), Mantodea (praying mantises) and Dermaptera (earwigs) of the Insect Limestone (late Eocene), Isle of Wight, including the first record of Mantodea from the UK. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 110:301-311 [M. Clapham/M. Clapham/M. Clapham]
9429 P. A. Selden. 2001. Eocene spiders from the Isle of Wight with preserved respiratory structures. Palaeontology 44(4):695-729 [D. Smith/A. Kinchloe/A. Kinchloe]
51851 P. A. Selden. 2014. Spiders (Arachnida: Araneae) from the Insect Limestone (Bembridge Marls, Late Eocene) of the Isle of Wight, southern England. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 104:275-282 [M. Clapham/M. Clapham]
51856 A. Shmakov. 2014. Thrips (Insecta: Thysanoptera) from the Insect Limestone (Bembridge Marls, Late Eocene) of the Isle of Wight, UK. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 104:317-325 [M. Clapham/M. Clapham]
51857 I. D. Sukatsheva. 2014. Caddis-flies (Insecta: Trichoptera) from the Insect Limestone (Bembridge Marls, Late Eocene) of the Isle of Wight, UK. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 104:327-333 [M. Clapham/M. Clapham]
71383 J. Szwedo, J. Drohojowska, Y. A. Popov, E. Simon, and P. Wegierek. 2019. Aphids, true hoppers, jumping plant-lice, scale insects, true bugs and whiteflies (Insecta: Hemiptera) from the Insect Limestone (latest Eocene) of the Isle of Wight, UK. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 110:331-396 [M. Clapham/M. Clapham/M. Clapham]
45339 K. von Rosen. 1913. Die Fossilen Termiten: Eine kurze Zusammenfassung der bis jetzt bekannten Funde. Transactions of the 2nd International Congress of Entomology 2:318-335 [M. Clapham/M. Clapham]
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