Lothagam-Upper Nawata Mb: Messinian - Zanclean, Kenya

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Mammalia - Tubulidentata - Orycteropodidae
Orycteropus sp. Geoffroy 1791
2 specimens
Mammalia - Proboscidea - Elephantidae
Elephantidae indet. Gray 1821
1 specimen
Stegotetrabelodon orbus Maglio 1970
2 specimens
Elephas nawataensis n. sp. Tassy 2003
2 specimens
Mammalia - Proboscidea - Gomphotheriidae
Gomphotheriidae indet. Hay 1922
1 specimen
Anancus kenyensis MacInnes 1942
2 specimens
Mammalia - Proboscidea - Deinotheriidae
Deinotherium bozasi Arambourg 1934
1 specimen
Mammalia - Perissodactyla - Rhinocerotidae
Ceratotherium praecox
5 specimens
Brachypotherium lewisi
10 specimens
Diceros bicornis (Linnaeus 1758)
4 specimens
Mammalia - Perissodactyla - Equidae
Eurygnathohippus feibeli n. sp.
24 specimens
Eurygnathohippus turkanense (Hooijer and Maglio 1974)
16 specimens
Mammalia - Bovidae
Reduncini indet. Knottnerus-Meyer 1907
5 specimens
Menelikia leakeyi Harris 1991
3 specimens
Kobus presigmoidalis n. sp. Harris 2003
13 specimens
Kobus laticornis n. sp. Harris 2003
11 specimens
Alcelaphini indet. Brookes 1876
2 specimens
Aepyceros premelampus n. sp. Harris 2003
58 specimens
Hippotragini indet. Sundevall 1845
2 specimens
Hippotragus sp. Sundevall 1845
2 specimens
Damalacra sp. Gentry 1980
23 specimens
sp. A
? Praedamalis sp. Dietrich 1950
2 specimens
Gazella sp. Blaineville 1816
2 specimens
Madoqua sp. Ogilby 1837
3 specimens
Bovini indet. Gray 1821
3 specimens
Boselaphini indet. Knottnerus-Meyer 1997
7 specimens
Tragoportax sp. Pilgrim 1937
2 specimens
sp. A
Tragoportax aff. cyrenaicus Thomas 1979
1 specimen
Tragelaphini indet. Blyth 1863
2 specimens
Tragelaphus kyaloae Harris 1991
1 specimen
Mammalia - Giraffidae
Palaeotragus sp. Gaudry 1861
4 specimens
Palaeotragus germaini Arambourg 1959
4 specimens
Mammalia - Tayassuidae
Cainochoerus cf. africanus (Hendy 1976)
1 specimen
Mammalia - Suidae
Nyanzachoerus syrticus tulotus
49 specimens
Nyanzachoerus cf. australis
4 specimens
Nyanzachoerus cf. syrticus (Leonardi 1954)
5 specimens
Nyanzachoerus devauxi (Arambourg 1968)
7 specimens
Potamochoerus sp. Gray 1854
1 specimen
Mammalia - Hippopotamidae
Hexaprotodon harvardi Coryndon 1977
63 specimens
Hexaprotodon lothagamensis n. sp. Weston 2000
1 specimen
Hexaprotodon sp. Falconer and Cautley 1836
3 specimens
Mammalia - Carnivora - Mustelidae
Mellivorinae indet. Gray 1865
1 specimen
Erokomellivora lothagamensis n. gen., n. sp.
1 specimen
Mammalia - Carnivora - Amphicyonidae
Amphicyonidae indet. Haeckel 1886
1 specimen
sp. B
Mammalia - Carnivora - Felidae
Dinofelis sp. Zdansky 1924
8 specimens
Lokotunjailurus emageritus n. gen., n. sp. Werdelin 2003
15 specimens
cf. Metailurus sp. Zdansky 1924
6 specimens
Mammalia - Carnivora - Hyaenidae
Hyaenictitherium cf. parvum Khomenko 1914
2 specimens
cf. Hyaenictis sp. Gaudry 1861
1 specimen
Mammalia - Carnivora - Viverridae
Viverridae indet. Gray 1821
1 specimen
large sp.
cf. Genetta sp. Cuvier 1816
1 specimen
sp. B
Mammalia - Rodentia - Muridae
Saidomys sp. James and Slaughter 1974
1 specimen
Mammalia - Rodentia - Thryonomyidae
Thryonomys sp. Fitzinger 1867
5 specimens
small sp.
Mammalia - Primates - Hominidae
Hominidae indet. Gray 1825
2 specimens
Mammalia - Primates - Cercopithecidae
Cercopithecidae indet. Gray 1821
2 specimens
Parapapio lothagamensis n. sp. Leakey et al. 2003
31 specimens
Colobinae indet. (Blyth 1875)
3 specimens
sp. A
Colobinae indet. (Blyth 1875)
8 specimens
sp. B
Colobinae indet. (Blyth 1875)
4 specimens
small sp.
Reptilia - Testudines - Testudinidae
cf. Geochelone indet. Fitzinger 1835
7 specimens
Reptilia - Testudines - Trionychidae
Trionychidae indet. Gray 1825
4 specimens
Cycloderma debrionae
1 specimen
Cycloderma frenatum Peters 1854
2 specimens
Reptilia - Testudines - Podocnemididae
Turkanemys pattersoni n. gen., n. sp. Wood 2003
34 specimens
Reptilia
Euthecodon brumpti (Joleaud 1920)
13 specimens
Rimasuchus n. gen. lloydi (Fourtau 1918)
9 specimens
    = Crocodylus sp. Laurenti 1768
Brochu and Storrs 2012
KNM-LT specimens
Crocodylus niloticus Laurenti 1768
3 specimens
    = Crocodylus checchiai Maccagno 1947
Brochu and Storrs 2012
KNM-LT 26618 (partial skull)
Aves indet. Linnaeus 1758
1 specimen
Reptilia - Struthioniformes - Struthionidae
cf. Struthio indet. Linnaeus 1758
12 specimens
Reptilia - Anseriformes - Anatidae
Anatidae indet. Vigors 1825
1 specimen
Reptilia - Gruiformes - Rallidae
Rallidae indet. Vigors 1825
1 specimen
Reptilia - Pelecaniformes - Pelecanidae
Pelecanus sp. Linnaeus 1758
2 specimens
Reptilia - Pelecaniformes - Ardeidae
Ardeidae indet. Leach 1820
3 specimens
Reptilia - Suliformes - Anhingidae
Anhinga cf. rufa Daudin 1802
3 specimens
Reptilia - Ciconiiformes - Ciconiidae
Leptoptilos cf. crumeniferus
1 specimen
Dipnomorpha - Lepidosirenidae
Protopterus sp. Agassiz 1843
6 specimens
Cladistia - Polypteriformes - Polypteridae
Polypterus sp. Lacépède 1803
231 specimens
Actinopteri - Siluriformes
Siluriformes indet.
134 specimens
Actinopteri - Siluriformes - Mochokidae
Synodontis sp. Cuvier 1816
4 specimens
Actinopteri - Siluriformes - Bagridae
Bagridae indet. Bleeker 1852
12 specimens
Bagrus aff. sp. Bosc 1816
2 specimens
Actinopteri - Siluriformes - Claroteidae
Clarotes sp. Kner 1855
21 specimens
Actinopteri - Characiformes - Alestidae
Hydrocynus sp. Cuvier 1816
11 specimens
Actinopteri - Perciformes
Perciformes indet. Bleeker 1859
5 specimens
Percoidei indet. (Bleeker 1859)
9 specimens
Actinopteri - Centropomidae
Lates niloticus (Linnaeus 1758)
5 specimens
Actinopteri - Osteoglossiformes - Gymnarchidae
Gymnarchus sp. Cuvier 1829
9 specimens
Actinopteri - Osteoglossiformes - Osteoglossidae
Heterotis sp. Rüppell 1828
23 specimens
unclassified
Sindacharax lothagamensis n. sp.
1 specimen
? Schilbe sp.
1 specimen
informal Clarias/Heterobranchus sp.
1420 specimens
Semlikiichthys cf. rhachirhinchus
3 specimens
Eupodotis sp.
1 specimen
Abudhabia sp.
1 specimen
Malacostraca - Decapoda - Potamonautidae
Potamonautidae indet. Bott 1970
17 specimens
see common names

Geography
Country:Kenya
Coordinates: 2.9° North, 36.1° East (view map)
Paleocoordinates:2.6° North, 35.5° East
Basis of coordinate:stated in text
Geographic resolution:local area
Time
Period: Neogene Epoch: Miocene - Pliocene
Stage: Messinian - Zanclean 10 m.y. bin: Cenozoic 6
*Period:Tertiary
*International age/stage:Tortonian - Piacenzian
Key time interval: Messinian - Zanclean
Age range of interval: 7.246 - 3.6 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Nawata or Nachukui Member:Upper Nawata and Apak
Stratigraphic resolution:member
Stratigraphy comments: K-Ar dating constrains the Upper Nawata Member to 6.54 +/- 0.04 Ma (Marker Tuff) and 4.22 +/- 0.03 Ma (pumice clasts from 35 m above the top of the Upper Nawata Purple Marker). Thus, top of the Upper Nawata is older than 4.2 Ma, and is estimated to be 5.0 Ma.
Lithology and environment
Primary lithology:current ripples,"cross stratification",grading poorly lithified sandy sandstone
Secondary lithology:tuffaceous poorly lithified mudstone
Includes fossils?Y
Lithology description: The Upper Nawata is characterized by thick, multistoried sandstone bodies, with subsidiary mudstones, and by a paucity of altered distal tephra. The type section consists of some 75% sandstones, 21% mudstones, and 4% altered tephra. The basal unit of the member is a tephra complex termed the Marker Tuff. This complex consists of a thin-bedded, relatively homogeneous lower interval (interpreted as airfall tuff), overlain by a massive, poorly sorted sandy tuff. Sandstones of the upper Nawata are typically characterized by well-developed epsilon cross-stratification and trough cross-beds in the coarser units, grading into small-scale troughs and ripple marks in the overlying finer intervals, with abrupt transitions back to coarser sands. Mudstones are subordinate, but where present they show characteristics of paleosol development. (ref # 8636 p. 22-23).
Environment:fluvial indet. Tectonic setting:rift
Geology comments: Upper Nawata strata indicated that subsidence rates decreased dramatically, allowing the fluvial system to rework overbank deposits- recycling mud into clay to produce the characteristic multistoried sandstones. Probably decrased chanel flow, perhaps to a seasonal state. Environmental conditions suggested to be generally drier than previous or subsequent times.
Taphonomy
Modes of preservation:body
Degree of concentration:dispersed
Size of fossils:macrofossils,mesofossils
Spatial orientation:random
Preservation of anatomical detail:good
Articulated whole bodies:none
Associated major elements:some
Disassociated major elements:many
Disassociated minor elements:many
Size sorting:poor
Fragmentation:frequent
Feeding/predation traces:fractures
Temporal resolution:time-averaged
Collection methods and comments
Collection methods:bulk,surface (in situ),mechanical,sieve,field collection
Collection size:2463 specimens
Reason for describing collection:paleoecologic analysis
Collection method comments: Held at the Kenya National Museums, Nairobi (KNM-LT-). Surface collection.
Metadata
Also known as:ETE Locality 1065, Lothagam 1C, Lothagam 1, Lothagam, Lothagam-Upper Nawata
Database number:22242
Authorizer:A. Behrensmeyer, P. Mannion Enterer:A. Behrensmeyer, P. Mannion, S. Cote
Modifier:G. Varnham Research group:vertebrate
Created:1999-06-30 00:00:00 Last modified:2022-04-27 05:30:36
Access level:the public Released:1999-06-30 00:00:00
Creative Commons license:CC BY
Reference information

Primary reference:

8636.ETE M. G. Leakey and J. M. Harris. 2003. Lothagam: the dawn of humanity in eastern Africa. Lothagam: the dawn of humanity in eastern Africa 678 [A. Behrensmeyer/R. Bobe/R. Bobe]

Secondary references:

46593 C. A. Brochu and G. W. Storrs. 2012. A giant crocodile from the Plio-Pleistocene of Kenya, the phylogenetic relationships of Neogene African crocodylines, and the antiquity of Crocodylus in Africa. Journal of Vertebrate Paleontology 32(3):587-602 [P. Mannion/P. Mannion]
5448ETE 1183M. G. Leakey and C. Feibel. 1996. Lothagam: a record of faunal change in the late Miocene of East Africa. Journal of Vertebrate Paleontology 16(3):556-570 [A. Behrensmeyer/A. Behrensmeyer/J. Alroy]