Bajo de Santa Rosa locality (MACN): Late Campanian - Early Maastrichtian, Argentina
collected 1990–1994

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Actinopteri - Amiiformes - Amiidae
Amia sp. Linnaeus 1766
Bogan et al. 2010 1 specimen
MML 449, an incomplete anterior portion of right dentary
Actinopteri - Lepisosteiformes - Lepisosteidae
Lepisosteidae indet. Cuvier 1825
Martinelli and Forasiepi 2004 6 elements
MACN-PV RN 1071, six vertebral centra
Actinopteri
Teleostei indet. Müller 1846
Martinelli and Forasiepi 2004 10 elements
MACN-PV RN 1077, ten hemispherical teeth of different sizes
Actinopteri - Percichthyidae
cf. Percichthyidae indet. Jordan and Eigenman 1890
Martinelli and Forasiepi 2004 19 elements
MACN-PV RN 1081, nineteen fragmentary vertebrae
Actinopteri - Siluriformes
Siluriformes indet.
Martinelli and Forasiepi 2004 5 elements
MACN-PV RN 1074, four very small incomplete pectoral spines; MACN-PV RN 1084, incomplete pectoral spine lacking its proxinial portion
Actinopteri - Siluriformes - Diplomystidae
Diplomystidae indet. Eigenman 1890
Martinelli and Forasiepi 2004 4 elements
MACN-PV RN 1072, four incomplete pectoral spines
Amphibia - Temnospondyli - Calyptocephalellidae
Leptodactylidae indet. Stirton 1953
Martinelli and Forasiepi 2004 25 elements
MACN-PV RN 1063, fragment of right maxilla; MACN-PV RN 1066, complete right humerus; MACN-PV RN 1069, twenty three fragments of ornamented cranial hones including incomplete maxillae, and frontoparietals
    = Calyptocephalella satan Agnolin 2012
Agnolin 2012
    = Calyptocephalellidae indet. Reig 1960
Suazo Lara and Gómez 2022
Anura indet. (Fischer von Waldheim 1813)
Martinelli and Forasiepi 2004 11 elements
MACN-PV RN 1067, five incomplete vertebrae; MACN-PV RN 1068, five incomplete angulosplenials; MACN-PV RN 1070, fused radio and ulna
    = Calyptocephalella satan Agnolin 2012
Agnolin 2012
    = Calyptocephalellidae indet. Reig 1960
Suazo Lara and Gómez 2022
Amphibia - Temnospondyli - Pipidae
Pipidae indet. Gray 1825
Martinelli and Forasiepi 2004 3 specimens
MACN-PV RN 1064, almost complete sphenethmoid; MACN-PV RN 1065, sphenethmoid
    = Kuruleufenia xenopoides n. gen., n. sp. Gómez 2016
Gómez 2016
Reptilia - Aniliidae
? Madtsoiidae indet. (Hoffstetter 1961)
Martinelli and Forasiepi 2004 9 specimens
MACN-PV RN 1059: almost complete trunk vertebra
    = Australophis anilioides Gómez et al. 2008
Gómez et al. 2008
MML-PV182-190
Reptilia - Madtsoiidae
Madtsoiidae indet. (Hoffstetter 1961)
Martinelli and Forasiepi 2004 3 specimens
MACN-PV RN 1051-52, MACN-PV RN 1054: isolated trunk vertebrae
Alamitophis argentinus Albino 1986
Martinelli and Forasiepi 2004 1 specimen
MACN-PV RN 1053: incomplete trunk vertebra.
Patagoniophis parvus Albino 1986
Martinelli and Forasiepi 2004 2 specimens
MACN-PV RN 1049 and MACN-PV RN 1056: incomplete trunk vertebrae
Reptilia - Testudines - Chelidae
Chelidae informal indet. 1
Martinelli and Forasiepi 2004 1 specimen
MACN-PV RN 1089: isolated fragments of posterior peripheral caparace
Chelidae informal indet. 2
Martinelli and Forasiepi 2004 1 specimen
MACN-PV RN 1090: isolated fragments of shell
Chelidae informal indet. 3
Martinelli and Forasiepi 2004 1 specimen
MACN-PV RN 1091: isolated fragments of shell
Reptilia - Faveoloolithidae
Sphaerovum erbeni Mones 1980
Martinelli and Forasiepi 2004 7 specimens
Reptilia
Bonatitan reigi n. gen., n. sp. Martinelli and Forasiepi 2004
Martinelli and Forasiepi 2004 2 individuals
MACN-PV RN 821 (holotype); MACN-PV RN 1061
Reptilia - Titanosauridae
Megaloolithidae indet. Zhao 1979
Martinelli and Forasiepi 2004 8 specimens
MACN-PV RN 1096: fragments of eggshells and eight partial eggs
Reptilia - Theropoda - Abelisauridae
Abelisauridae indet. Bonaparte and Novas 1985
Meso et al. 2021 1 specimen
MACN-Pv-RN 1085, tooth
Reptilia - Theropoda
cf. Carcharodontosauridae indet. Stromer 1931
Martinelli and Forasiepi 2004 1 specimen
MACN-PV RN 1086: an isolated tooth
    = Tetanurae indet. Gauthier 1986
Carrano et al. 2012
cf. Neornithes sp. (Gadow 1893)
Agnolin and Novas 2012 1 specimen
MML 206; left carpometacarpus
Reptilia - Theropoda - Dromaeosauridae
? Dromaeosauridae indet. Colbert and Russell 1969
1 individual
MML-195 (3 measurements)
    = Deinonychosauria indet. Colbert and Russell 1969
Novas et al. 2004
    = Austroraptor cabazai n. gen., n. sp. Novas et al. 2008
Novas et al. 2008
MML-195
Reptilia - Hadrosauridae
Hadrosauridae indet. Cope 1869
Martinelli and Forasiepi 2004 7 specimens
MRPV 431/P to 437/P, fragmentary dentaries
Reptilia
Ankylosauria indet. (Osborn 1923)
Murray et al. 2019 5 specimens
MACN Pv RN1149, five isolated osteoderms
Reptilia - Plesiosauria
Plesiosauria indet. de Blainville 1835
O'Gorman et al. 2011 1 element
MACN-PV RN 1087d (cervical centra)
Reptilia - Plesiosauria - Elasmosauridae
Elasmosauridae indet. Cope 1869
Martinelli and Forasiepi 2004 2 elements
MACN-PV RN 1087a-b (cervical centra)
Reptilia - Plesiosauria - Polycotylidae
Polycotylidae indet. Williston 1908
O'Gorman et al. 2011 1 element
MACN-PV RN 1087c (cervical centra)
Reptilia - Sphenodontidae
Sphenodontidae indet. Cope 1871
Martinelli and Forasiepi 2004 1 specimen
MACN-PV RN 1062, middle portion of the right lower jaw
    = Sphenodontinae indet. Cope 1871
Apesteguia and Jones 2012
MACN-PV 1062 (partial right maxilla and two isolated teeth)
Dipnoi - Ceratodontidae
Ceratodontidae indet. Gill 1872
Martinelli and Forasiepi 2004 3 elements
MACN-PV RN 1079, left tooth plate; MACN-PV RN 1080, two tooth plates
    = Atlantoceratodus patagonicus Agnolín 2010
Agnolín 2010
Chondrichthyes
Chondrichthyes indet. Huxley 1880
Martinelli and Forasiepi 2004 11 elements
MACN-PV RN 1076: eleven complete vertebral centra
see common names

Geography
Country:Argentina State/province:Río Negro
Coordinates: 40.1° South, 66.8° West (view map)
Paleocoordinates:42.9° South, 52.5° West
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period: Cretaceous Epoch: Late Cretaceous
Stage: Campanian - Maastrichtian 10 m.y. bin: Cretaceous 7 - Cretaceous 8
Key time interval: Late Campanian - Early Maastrichtian
Age range of interval: 83.6 - 66 m.y. ago
Stratigraphy
Geological group:Malargüe Formation:Allen Member:Lower
Stratigraphic resolution:group of beds
Stratigraphy comments: lower member of formation; possibly equivalent to "egg level 5"
Lithology and environment
Primary lithology:fine,yellow poorly lithified sandstone
Secondary lithology: mudstone
Includes fossils?Y
Includes fossils?Y
Lithology description: "alternated sandstones and mudstones with small lents of conglomerates." "The fossil-bearing bed consists of friable, yellowish, fine-grained clean sandstone enclosed between siltstone layers."
Environment:fluvial indet.
Geology comments: "a fluvial environment in the basal section"
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Preservation of anatomical detail:good
Associated major elements:some
Disassociated major elements:many
Disassociated minor elements:many
Fragmentation:occasional
Collection methods and comments
Collection methods:selective quarrying,surface (float),surface (in situ),mechanical,field collection
Reason for describing collection:general faunal/floral analysis
Museum repositories:MACN
Collection dates:1990–1994
Metadata
Also known as:Salitral de Santa Rosa, El Matuasto, "Cerro Bonaparte", Lamarque
Database number:49557
Authorizer:M. Carrano, R. Benson, E. Vlachos, P. Mannion Enterer:M. Carrano, P. Mannion, M. Jones, F. Aspromonte
Modifier:F. Aspromonte Research group:vertebrate
Created:2005-04-19 14:57:44 Last modified:2024-01-08 10:12:25
Access level:the public Released:2005-04-19 14:57:44
Creative Commons license:CC BY
Reference information

Primary reference:
76469. F. E. Novas, J. I. Canale, and M. P. Isas. 2003. Un terópodo maniraptor del Campaniano–Maastrichtiano del norte patagónico [A maniraptoran theropod from the Campanian–Maastrichtian of the Patagonian north]. Ameghiniana 40(4 (suppl.)):63R [M. Carrano/M. Carrano]

Secondary references:

86689 F. Agnolín. 2010. A new species of the genus Atlantoceratodus (Dipnoiformes: Ceratodontoidei) from the Uppermost Cretaceous of Patagonia and a brief overview of fossil dipnoans from the Cretaceous and Paleogene of South America. Brazilian Geographical Journal 1(2):162-210 [E. Vlachos/F. Aspromonte]
73340 F. Agnolin. 2012. A new Calyptocephalellidae (Anura, Neobatrachia) from the Upper Cretaceous of Patagonia, Argentina, with comments on its systematic position. Studia Geologica Salmanticensia 48(2):129-178 [P. Mannion/P. Mannion]
38457 F. L. Agnolin and F. E. Novas. 2012. A carpometacarpus from the Upper Cretaceous of Patagonia sheds light on the Ornithurine bird radiation. Palaeontologische Zeitschrift 86(1):85-89 [M. Carrano/M. Carrano/P. Mannion]
38874 S. Apesteguia and M. E. H. Jones. 2012. A Late Cretaceous "tuatara" (Lepidosauria; Sphenodontinae) from South America. Cretaceous Research 34:154-160 [R. Benson/M. Jones/P. Mannion]
86230 S. Bogan, L. Taverne, and F. L. Agnolín. 2010. First fossil record of an amiid fish (Halecomorphi, Amiidae) from the Latest Cretaceous of Patagonia, Argentina, and comments on the status of Pappichthys patagonica Ameghino, 1906 (Teleostei, Osteoglossidae). Bulletin de l'Institut royal del Sciences naturelles de Belgique, Sciences de la Terre 80:163-170 [E. Vlachos/F. Aspromonte]
41617 M. T. Carrano, R. B. J. Benson, and S. D. Sampson. 2012. The phylogeny of Tetanurae (Dinosauria: Theropoda). Journal of Systematic Palaeontology 10(2):211-300 [M. Carrano/M. Carrano]
76416 P. Cruzado-Caballero. 2017. New hadrosaurid remains from the Late Cretaceous of Río Negro Province (Argentina, Late Cretaceous). Journal of Iberian Geology 43:307-318 [M. Carrano/M. Carrano/M. Carrano]
75729 P. Cruzado-Caballero, L. S. Filippi, A. H. Méndez, A. C. Garrido, and I. Díaz-Martínez. 2018. First ornithopod remains from the Bajo de la Carpa Formation (Santonian, Upper Cretaceous), northern Patagonia, Argentina. Cretaceous Research 83:182-193 [M. Carrano/M. Carrano/M. Carrano]
46024 P. J. Currie and A. Paulina Carabajal. 2012. A new specimen of Austroraptor cabazai, Pol, Canale, Porfiri and Calvo, 2008 (Dinosauria, Theropoda, Unenlagiidae) from the latest Cretaceous (Maastrichtian) of Río Negro, Argentina. Ameghiniana 49(4):662-667 [P. Mannion/J. Tennant/M. Carrano]
76256 M. S. Fernández. 2013. Análisis de cáscaras de huevos de dinosaurios de La Formación Allen, Cretácico Superior de Río Negro (Campaniano-Maastrichtiano): utilidad de los macrocaracteres de interés parataxonómico [Analysis of dinosaur eggshells from the Allen Formation, Upper Cretaceous of Río Negro (Campanian–Maastrichtian): usefulness of macrocharacters of parataxonomic interest]. Ameghiniana 50(1):79-97 [M. Carrano/M. Carrano]
29611ETE M. S. Fernández and S. D. Matheos. 2008. Estudio petrográfico de cáscaras anomrlaes de huevos de dinosaurios del Cretácico Superior (Río Negro, Argentina) [Petrographic study of abnormal dinosaur eggshells from the Upper Cretaceous (Río Negro, Argentina)]. Ameghiniana 45(4, suppl.):26R [M. Carrano/M. Carrano]
46152 Z. Gasparini, L. Salgado, and A. Parras. 2007. Late Cretaceous plesiosaurs from northern Patagonia, Argentina. Geological Journal 42:185-202 [P. Mannion/J. Tennant/F. Aspromonte]
49276 F. A. Gianechini, P. J. Makovicky, and S. Apesteguía. 2011. The teeth of the unenlagiine theropod Buitreraptor from the Cretaceous of Patagonia, Argentina, and the unusual dentition of the Gondwanan dromaeosaurids. Acta Palaeontologica Polonica 56(2):279-290 [M. Carrano/M. Carrano]
78231 F. A. Goianechini and S. Apesteguía. 2011. Unenlagiinae revisited: dromaeosaurid theropods from South America. Anais da Academia Brasileira de Ciências 83(1):163-195 [M. Carrano/M. Carrano]
62828 R. O. Gómez. 2016. A new pipid frog from the Upper Cretaceous of Patagonia and early evolution of crown-group Pipidae. Cretaceous Research 62:52-64 [P. Mannion/P. Mannion/F. Aspromonte]
29129ETE R. O. Gómez, A. M. Báez, and G. W. Rougier. 2008. An anilioid snake from the Upper Cretaceous of northern Patagonia. Cretaceous Research 29:481-488 [M. Carrano/M. Carrano/R. Benson]
40526 G. Grellet-Tinner, L. E. Fiorelli, and R. B. Salvador. 2012. Water vapor conductance of the Lower Cretaceous dinosaurian eggs from Sanagasta, La Rioja, Argentina: paleobiological and paleoecological implications for South American faveoloolithid and megaloolithid eggs. Palaios 27:35-47 [M. Carrano/M. Carrano]
34689ETE R. D. Juárez Valieri, J. O. Calvo, and S. D. Rios Díaz. 2011. Sauropods crossing formations: biostratigraphical implications for Patagonian faunal assemblages. In J. O. Calvo, J. D. Porfiri, B. J. González Riga, D. Dos Santos (eds.), Dinosaurios y Paleontología desde América Latina. Anales del III Congreso Latinoamericano de Paleontología. Editorial de la Universidad Nacional de Cuyo, Neuquén 153-160 [M. Carrano/M. Carrano]
13269ETE A. G. Martinelli and A. M. Forasiepi. 2004. Late Cretaceous vertebrates from Bajo de Santa Rosa (Allen Formation), Río Negro province, Argentina, with the description of a new sauropod dinosaur (Titanosauridae). Revista del Museo Argentino de Ciencias Naturales, nuevo serie 6(2):257-305 [M. Carrano/M. Carrano/F. Aspromonte]
86551 J. G. Meso, R. D. Juárez Valieri, J. D. Porfiri, S. A. S. Correa, A. G. Martinelli, G. A. Casal, J. I. Canudo, F. Poblete, and D. Dos Santos. 2021. Testing the persistence of Carcharodontosauridae (Theropoda) in the Upper Cretaceous of Patagonia based on dental evidence. Cretaceous Research 125:104875 [M. Carrano/M. Carrano]
85173 A. Murray, F. Riguetti, and S. Rozadilla. 2019. New ankylosaur (Thyreophora, Ornithischia) remains from the Upper Cretaceous of Patagonia. Journal of South American Earth Sciences 96:102320:1-5 [E. Vlachos/F. Aspromonte/M. Carrano]
16795ETE F. E. Novas, J. Canale, and M. Isasi. 2004. Giant deinonychosaurian theropod from the Late Cretaceous of Patagonia. Journal of Vertebrate Paleontology 24(3):98A [M. Carrano/M. Carrano]
28884ETE F. E. Novas, D. Pol, J. I. Canale, J. D. Porfiri, and J. O. Calvo. 2008. A bizarre Cretaceous theropod dinosaur from Patagonia and the evolution of Gondwanan dromaeosaurids. Proceedings of the Royal Society B [M. Carrano/M. Carrano]
86335 J. P. O'Gorman, L. Salgado, and Z. Gasparini. 2011. Plesiosaurios de la Formación Allen (Campaniano-Maastrichtiano) en el área del Salitral de Santa Rosa (Provincia de Río Negro, Argentina). Ameghiniana 48(1):129-135 [E. Vlachos/F. Aspromonte]
85526 F. Suazo Lara and R. O. Gómez. 2022. In the shadow of dinosaurs: Late Cretaceous frogs are distinct components of a widespread tetrapod assemblage across Argentinean and Chilean Patagonia. Cretaceous Research 131:105085 [P. Mannion/P. Mannion]