Los Cayos A tracksite: Aptian, Spain
collected 1985
List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
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Reptilia
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| Hernán Martínez 2018 | ||||||||||
| Sanz et al. 1990 | 36 individuals | |||||||||
| trackways (+ 232 isolated tracks) | ||||||||||
| Pérez-Lorente 2015 | ||||||||||
| referred to Bueckeburgichnus maximus by Moratalla 1993 (PhD thesis) | ||||||||||
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Reptilia
- Pterosauria
- Pteraichnidae
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| Moratalla et al. 2003 | 4 specimens | |||||||||
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Reptilia
- Testudines
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Testudines indet.
Batsch 1788
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Moratalla and Hernan 2009 | |||||||||
| Emydiphus sp. (trackways) | ||||||||||
| see common names | ||||||||||
Geography
| Country: | Spain | State/province: | La Rioja | County: | Cornago |
| Coordinates: | 42.1° North, 2.1° West (view map) | ||||
| Paleocoordinates: | 32.1° North, 23.3° East (Wright 2013) | ||||
| Basis of coordinate: | stated in text | ||||
| Altitude: | 755 meters | ||||
| Geographic resolution: | outcrop |
Time
| Period: | Cretaceous | Epoch: | Early Cretaceous |
| Stage: | Aptian | 10 m.y. bin: | Cretaceous 3 |
| Key time interval: | Aptian | ||
| Age range of interval: | 121.4 - 113.2 m.y. ago |
Stratigraphy
| Geological group: | Enciso | ||||
| Stratigraphic resolution: | group of beds | ||||
| Stratigraphy comments: 8 track-bearing layers (A1-A8) | |||||
Lithology and environment
| Primary lithology: | desiccation cracks,wave ripples,"cross stratification",planar lamination,bioturbation,ferruginous claystone |
| Secondary lithology: | wave ripples,fine,quartzose sandstone |
| Includes fossils? | Y |
| Includes fossils? | Y |
| Lithology description: "fine-grained sandstones" A1–A2: theropod tracks; "fine quartzarenites with clear evidence of surf action" A3: pterosaur and theropod tracks; "a dark claystone with symmetrical ripples, cross-bedded lamination, horizontal lamination, Fe crusts, a few fossil fragments, and bioturbation." A4: possible dinosaur tracks; "a sandy siltstone with abundant carbonate cement, dessication cracks, ostracods, and bioturbation." A5: theropod tracks; "quartzose fine-grained sandstone (locally graywacke), 20 cm thickness...symmetrical ripple-marks with straight crests" A6: theropod tracks; "a fine-grained graywacke, 20 cm thickness, with ferruginous crusts, bioturbation, and a few ostracods." A7: hundreds of tracks; "a very fine-grained graywacke (locally a sandy siltstone), with horizontal lamination, cross-bedded lamination, dessication cracks, iron crusts, and intense bioturbation...many symmetrical ripple-marks with straight crests" A8: theropod tracks; "a dark siltstone with horizontal lamination, cross-bedded lamination, siltstone nodules, dessication cracks, Fe crusts, bioturbation, and symmetrical ripple-marks with straight crests" | |
| Environment: | lacustrine delta plain |
Taphonomy
| Modes of preservation: | mold/impression,trace |
| Degree of concentration: | concentrated |
| Size of fossils: | macrofossils |
| Spatial orientation: | life position |
| Preservation of anatomical detail: | good |
| Associated major elements: | many |
| Disassociated major elements: | some |
| Spatial resolution: | autochthonous |
Collection methods and comments
| Collection methods: | surface (in situ),observed (not collected) |
| Reason for describing collection: | general faunal/floral analysis | Collection dates: | 1985 |
Metadata
| Also known as: | LCA | ||
| Database number: | 68448 | ||
| Authorizer: | M. Carrano, P. Mannion | Enterer: | M. Carrano, J. Tennant |
| Modifier: | M. Carrano | Research group: | vertebrate |
| Created: | 2007-01-22 13:41:38 | Last modified: | 2025-02-22 15:12:02 |
| Access level: | the public | Released: | 2007-01-22 13:41:38 |
| Creative Commons license: | CC0 | ||
Reference information
Primary reference:
| 28579. | ETE | J. J. Moratalla, J. L. Sanz, and S. Jiménez. 1988. Nueva evidencia icnologica de dinosaurios en el Cretácico Inferior de La Rioja (España) [New ichnological evidence of dinosaurs in the Lower Cretaceous of La Rioja (Spain)]. Estudios Geológicos 44:119-131 [M. Carrano/M. Carrano] |
Secondary references:
| 14199 | ETE | M. T. Carrano. 2025. Taxonomic opinions on the Dinosauria. [M. Carrano/M. Carrano/M. Carrano] |
| 55394 | E. García-Ortiz and F. Pérez-Lorente. 2014. Palaeoecological inferences about dinosaur gregarious behaviour based on the study of tracksites from La Rioja area in the Cameros Basin (Lower Cretaceous, Spain). Journal of Iberian Geology 40(1):113-127 [M. Carrano/M. Carrano] | |
| 28419 | ETE | E. García-Ortiz de Landaluce and I. Díaz-Martínez. 2008. Aportaciones de algunos yacimientos representativos la La Rioja al estudio del comportamiento de los dinosaurios [Contributions of somerepresentative localities of La Rioja to the study of dinosaur behavior]. Palaeontologica Nova 8:207-219 [M. Carrano/M. Carrano] |
| 82548 | F. J. Hernán Martínez. 2018. Estratigrafía y Sedimentología de las Formaciones con Icnitas de Dinosaurios del Grupo Enciso (Cameros, La RIoja, Aptiense). Memoria [Stratigraphy and Sedimentology of the Formations with Dinosaur Footprints from the Enciso Group (Cameros, La Rioja, Aptian). Memoir]. 1-490 [M. Carrano/M. Carrano/M. Uhen] | |
| 78077 | C. Martín Escorza. 2001. Orientación de la huellas de dinosaurios en la Sierra de Cameros [Orientation of the dinosaur tracks in the Sierra de Cameros]. Zubía 19:139-163 [M. Carrano/M. Carrano] | |
| 28670 | ETE | J. J. Moratalla, J. Hernán, and S. Jiménez. 2003. Los Cayos dinosaur tracksite: an overview on the Lower Cretaceous ichno-diversity of the Cameros Basin (Cornago, La Rioja province, Spain). Ichnos 10:229-240 [M. Carrano/M. Carrano/M. Carrano] |
| 45205 | J. J. Moratalla and J. Hernan. 2009. Turtle and pterosaur tracks from the Los Cayos dinosaur tracksite, Cameros Basin (Cornago, La Rioja, Spain): tracking the Lower Cretaceous Biodiversity. Revista Española de Paleontología 24(1):59-77 [P. Mannion/J. Tennant] | |
| 78647 | J. J. Moratalla and J. Hernán. 2010. Probable palaeogeographic influences of the Lower Cretaceous Iberian rifting phase in the Eastern Cameros Basin (Spain) on dinosaur trackway orientations. Palaeogeography, Palaeoclimatology, Palaeoecology 295(1-2):116-130 [M. Carrano/M. Carrano] | |
| 78697 | J. J. Moratalla, J. L. Sanz, and S. Jimenez. 1988. Multivariate analysis on Lower Cretaceous dinosaur footprints: discrimination between ornithopods and theropods. Geobios 21(4):395-408 [M. Carrano/M. Carrano] | |
| 55588 | F. Pérez-Lorente. 2002. La distribución de yacimientos y de tipos de huellas de dinosaurios en la Cuenca de Cameroa (La Rioja, Burgos, Soria, España) [The distribution of localities and types of dinosaur footprints in the Cameros Basin (La Rioja, Burgos, Soria, Spain)]. Zubia Monográfico 14:191-210 [M. Carrano/M. Carrano] | |
| 82174 | F. Pérez-Lorente. 2015. Dinosaur Footprints and Trackways of La Rioja 1-363 [M. Carrano/M. Carrano] | |
| 70303 | J. L. Sanz. 2000. Yacimientos ibéricos de dinosaurios [Iberian dinosaur localities]. Dinosaurios: Los Señores del Pasado [Dinosaurs: The Lords of the Land] 267-299 [M. Carrano/M. Carrano] | |
| 19339 | ETE | J. L. Sanz, A. D. Buscalioni, J. J. Moratalla, V. Francés, and M. Antón. 1990. Los reptiles Mesozoicos del registro español [The Mesozoic reptiles from the Spanish record]. Monografias del Museo Nacional de Ciencias Naturales 2:1-79 [M. Carrano/M. Carrano] |