Khermeen Tsav, Red Beds (PIN coll. 3142) [SMPE]: Middle Campanian - Late Campanian, Mongolia
collected by V. Reshetov, R. Barsbold 1972, 2008

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Mammalia - Deltatheroida - Deltatheridiidae
Mammalia - Asioryctidae
Mammalia - Zalambdalestidae
Kielan-Jaworowska and Trofimov 1980 1 specimen
PIN 3142/710 (skull)
Mammalia - Multituberculata
Reptilia - Testudines
Testudines indet. Batsch 1788
'tortoises'
Reptilia - Testudines - Lindholmemydidae
Mongolemys barsboldi n. sp. Sukhanov and Narmandakh 1974
Danilov 2003 1 specimen
IG unnumbered
Reptilia - Hoplocercidae
Alifanov 1996 1 specimen
PIN 3142/319 (holotype skull)
Reptilia - Priscagamidae
Alifanov 1996 1 specimen
PIN 3142/317 (holotype fragmentary skull and vertebrae)
Alifanov 1996 1 specimen
PIN 3142/318 (holotype skull)
Alifanov 1996 2 specimens
PIN 3142/345 (holotype skull)
Reptilia - Polyglyphanodontidae
Alifanov 1993 1 specimen
PIN 3142/309 (holotype mandible)
Alifanov 1993 1 specimen
PIN 3142/308 (holotype skull)
Reptilia
Alifanov 1993 1 specimen
PIN 3142/324 (holotype partial skull)
Alifanov 1993 1 specimen
PIN 3142/310 (holotype partial skull)
Alifanov 1988 1 specimen
PIN 3142/306 (holotype partial snout)
'dinosaurs'
Reptilia - Oviraptoridae
4 specimens
GIN 100/30-100/33 (4 measurements)
    = Ingenia yanshini n. gen., n. sp. Barsbold 1981
Barsbold 1981
Reptilia - Gobioolithidae
Mikhailov et al. 1994 1 individual
eggs
    = Gobioolithus minor n. gen., n. sp. Mikhailov 1996
Mikhailov 1996
PIN 3142/422
Reptilia - Dromaeosauridae
Reptilia - Avetheropoda
Reptilia - Oblongoolithidae
Mikhailov 1996 1 specimen
PIN 3142/500-1
Reptilia - Ankylosauridae
Park et al. 2021 1 individual
MPC-D 100/1359 - first discovered and uncovered by the Joint Soviet-Mongolian Paleontological Expedition team in 1972 or 1973, but only collected in 2008 by members of Korea-Mongolia International Dinosaur Expedition
1 individual
PIN 3142/251
Reptilia - Protoceratopsidae
Kurzanov 1990 4 individuals
PIN 3142/1, 2, 3, 5
Czepiński 2019
1 individual
PIN 3142/4 (1 measurement)
Kurzanov 1990
    = Platyceratops tatarinovi n. gen., n. sp. Alifanov 2003
Alifanov 2003
Alifanov 2008 1 specimen
PIN 3142/299
Reptilia - Prismatoolithidae
Mikhailov et al. 1994 1 specimen
PIN 3142/496; = P2 Group ("?Breviceratops kozlowskii"); + P1 and P3 groups; PIN 3142/415 (group P2 eggs)
Mikhailov 1994
Mikhailov 1994
Reptilia - Crocodylia
Crocodylia indet. (Owen 1842)
Gubin 1999
'crocodiles'
Amphibia - Temnospondyli - Gobiatidae
Spinar and Tatarinov 1986 3 individuals
three partial skulls (PIN 3142/1, 2, and 3)
Gobiates leptocolaptus (Borsuk-Bialynicka 1978)
Gubin 1999 1 individual
Ostracoda - Podocopida - Limnocytheridae
Ostracoda - Podocopida - Cyprididae
Ostracoda - Podocopida
"ex. gr."
"ex. gr."
see common names

Geography
Country:Mongolia State/province:Omnogov
Coordinates: 43.3° North, 99.6° East (view map)
Paleocoordinates:37.5° North, 95.4° East (Wright 2013)
Basis of coordinate:estimated from map
Geographic resolution:outcrop
Time
Period: Cretaceous Epoch: Late Cretaceous
Stage: Campanian 10 m.y. bin: Cretaceous 7
*Period:Late/Upper Cretaceous - Late/Upper Cretaceous *Epoch:Senonian
*International age/stage:Campanian - Maastrichtian
Key time interval: Middle Campanian - Late Campanian
Age range of interval: 83.6 - 72.2 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Baruungoyot
Stratigraphic resolution:group of beds
Stratigraphy comments: Red Beds of Khermin Tsav
Lithology and environment
Primary lithology: not reported
Environment:terrestrial indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Preservation of anatomical detail:excellent
Collection methods and comments
Collection excludes:some macrofossils
Collection methods:surface (in situ),field collection
Reason for describing collection:taxonomic analysis
Collectors:V. Reshetov, R. Barsbold Collection dates:1972, 2008
Collection method comments: A broad collection that might include the collection #1157, but necessary because of nomenclatural variation from authors to authors, and also because the stratigraphic position of Gobiates khermeentsavi in the locality section is unknown. Distinct from PMPE and other collections from the same general site. Collected via Joint Soviet-Mongolian Paleontological Expedition
Metadata
Also known as:Khermin Tsav; Khermin-Tsab; Hermiin Tsav, Soviet-Mongolian Paleontological Expedition
Database number:11582
Authorizer:E. Fara, R. Benson, P. Mannion, M. Carrano Enterer:E. Fara, M. Carrano, P. Mannion, R. Benson
Modifier:M. Carrano Research group:vertebrate
Collections that are a subset of this one:11576
Created:2001-08-22 11:26:41 Last modified:2025-02-22 15:12:02
Access level:group members Released:2001-11-22 11:26:41
Creative Commons license:CC0
Reference information

Primary reference:
59136. R. Gradzinski, Z. Kielan-Jaworowska, and T. Maryanska. 1977. Upper Cretaceous Djadokhta, Barun Goyot and Nemegt formations of Mongolia, including remarks on previous subdivisions. Acta Geologica Polonica 27(3):281-318 [M. Carrano/M. Carrano]

Secondary references:

37919 V. R. Alifanov. 1988. The new lizards (Lacertilia: Teiidae) from the Upper Cretaceous of Mongolia. In F. Kurochkin (ed.), Fossil reptiles and birds of Mongolia 90-100 [R. Benson/R. Benson]
37931 V. R. Alifanov. 1989. [New priscagamas (Lacertilia) from the Upper Cretaceous of Mongolia and their systematic position in the Iguania]. Paleontologiceskij Zhurnal 4:73-87 [R. Benson/R. Benson]
38090 V. R. Alifanov. 1993. New lizards of the family Macrocephalosauridae (Sauria) from the Upper Cretaceous of Mongolia, critical remarks on the systematics of the Teiidae. Paleontological Journal 27(1):70-90 [R. Benson/R. Benson]
38117 V. R. Alifanov. 1996. Lizards of the families Priscagamidae and Hoplocercidae (Sauria, Iguania): phylogenetic position and new representatives from the Late Cretaceous of Mongolia. Paleontological Journal 30(4):466-483 [R. Benson/R. Benson]
38112 V. R. Alifanov. 2000. The fossil record of Cretaceous lizards from Mongolia. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 368-389 [R. Benson/R. Benson/M. Carrano]
11744ETE V. R. Alifanov. 2003. Two new dinosaurs of the infraorder Neoceratopsia (Ornithischia) from the Upper Cretaceous of the Nemegt Depression, Mongolian People's Republic. Paleontological Journal 37(5):524-534 [M. Carrano/M. Carrano/M. Carrano]
29118ETE V. R. Alifanov. 2008. The tiny horned dinosaur Gobiceratops minutus gen. et sp. nov. (Bagaceratopidae, Neoceratopsia) from the Upper Cretaceous of Mongolia. Paleontological Journal 42(6):621-633 [M. Carrano/M. Carrano/R. Butler]
71135 V. M. Arbour and P. J. Currie. 2011. Tail and pelvis pathologies of ankylosaurian dinosaurs. Historical Biology 23(4):375-390 [M. Carrano/M. Carrano]
66054 V. M. Arbour, P. J. Currie, and D. Badamgarav. 2014. The ankylosaurid dinosaurs of the Upper Cretaceous Baruungoyot and Nemegt formations of Mongolia. Zoological Journal of the Linnean Society 172(3):631-652 [G. Lloyd/G. Lloyd/M. Carrano]
64040 V. M. Arbour and P. J. Currie. 2016. Systematics, phylogeny and palaeobiogeography of the ankylosaurid dinosaurs. Journal of Systematic Palaeontology 14(5):385-444 [G. Lloyd/G. Lloyd/M. Carrano]
29489ETE R. Barsbold. 1981. Bezzubyye khishchnyye dinozavry Mongolii [Toothless carnivorous dinosaurs of Mongolia]. Sovmestnaia Sovetsko-Mongol’skaia Paleontologicheskaia Ekspeditsiia Trudy 15:28-39 [M. Carrano/M. Carrano]
15004ETE R. Barsbold. 1983. Khishchnye dinosavry mela Mongoliy [Carnivorous dinosaurs from the Cretaceous of Mongolia]. Transactions of the Joint Soviet-Mongolian Paleontological Expedition 19:1-117 [M. Carrano/M. Carrano]
86269 Ł Czepiński. 2019. Ontogeny and variation of a protoceratopsid dinosaur Bagaceratops rozhdestvenskyi from the Late Cretaceous of the Gobi Desert. Historical Biology 32(10):1394-1421 [M. Carrano/M. Carrano]
39832 I. Danilov. 2003. Gravemys Sukhanov and Narmandakh, 1983 (Testudinoidea: Lindholmemydidae) from the Late Cretaceous of Asia: new data. PaleoBios 23(3):9-19 [M. Carrano/M. Carrano]
41735% 1280Y. M. Gubin. 1999. Gobiatidy (Anura) iz verkhnemelovogo mestonakhozhdeniya Khermin-Tsav; Gobi, Mongoliya (= Gobiatids (Anura) from the Upper Cretaceous locality Khermeen-Tsav (Gobi Desert, Mongolia)). Paleontologicheskii Zhurnal 1:76-87 [E. Fara/E. Fara/J. Alroy]
39263 Z. Kielan-Jaworowska and B. A. Trofimov. 1980. Cranial morphology of the Cretaceous eutherian mammal Barunlestes. Acta Palaeontologica Polonica 25(2):167-185 [R. Benson/R. Benson]
69233 E. N. Kurochkin and R. Barsbold. 2000. The Russian-Mongolian expeditions and research in vertebrate palaeontology. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 235-255 [M. Carrano/M. Carrano]
86270 S. M. Kurzanov. 1990. A new Late Cretaceous protoceratopsid genus from Mongolia. Paleontological Journal 1990(4):85-91 [M. Carrano/M. Carrano]
14217ETE K. Mikhailov, K. Sabath, and S. Kurzanov. 1994. Eggs and nests from the Cretaceous of Mongolia. In K. Carpenter, K. F. Hirsch, and J. R. Horner (eds.), Dinosaur Eggs and Babies, Cambridge University Press, Cambridge 88-115 [M. Carrano/M. Carrano/M. Carrano]
44374 K. E. Mikhailov. 1994. Theropod and protoceratopsian dinosaur eggs from the Cretaceous of Mongolia and Kazakhstan. Paleontological Journal 28:101-120 [J. Alroy/A. Garcia Selles/M. Carrano]
69240 K. E. Mikhailov. 1996. Bird eggs in the Upper Cretaceous of Mongolia. Paleontological Journal 30(1):114-116 [M. Carrano/M. Carrano/M. Carrano]
69241 K. E. Mikhailov. 1996. New genera of fossil eggs from the Upper Cretaceous of Mongolia. Paleontological Journal 30(2):246-248 [M. Carrano/M. Carrano]
18353ETE K. E. Mikhailov. 2000. Eggs and eggshells of dinosaurs and birds from the Cretaceous of Mongolia. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 560-572 [M. Carrano/M. Carrano/M. Carrano]
82395 J.-Y. Park, Y.-N. Lee, Y. Kobayashi, L. L. Jacobs, R. Barsbold, H.-J. Lee, N. Kim, K.-Y. Song, and M. J. Polcyn. 2021. A new ankylosaurid from the Upper Cretaceous Nemegt Formation of Mongolia and implications for paleoecology of armoured dinosaurs. Scientific Reports 11:22928 [P. Mannion/P. Mannion]
75923 J.-Y. Park, Y.-N. Lee, P. J. Currie, M. J. Ryan, P. Bell, R. Sissons, E. B. Koppelhus, R. Barsbold, S. Lee and S.-H. Kim. 2021. A new ankylosaurid skeleton from the Upper Cretaceous Baruungoyot Formation of Mongolia: its implications for ankylosaurid postcranial evolution. Scientific Reports 11:4101:1-10 [P. Mannion/P. Mannion]
69215 M. A. Shishkin. 2000. Mesozoic amphibians from Mongolia and the Central Asiatic republics. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 297-308 [M. Carrano/M. Carrano]
62149 V. F. Shuvalov. 2000. The Cretaceous stratigraphy and palaeobiogeography of Mongolia. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 256-278 [P. Barrett/T. Cleary/M. Carrano]
30994 Z. V. Spinar and L. P. Tatarinov. 1986. A new genus and species of discoglossid frog from the Upper Cretaceous of the Gobi Desert. Journal of Vertebrate Paleontology 6(2):113-122 [J. Alroy/J. Alroy]
41695 V. B. Sukhanov. 2000. Mesozoic turtles of Middle and Central Asia. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 309-367 [R. Benson/R. Benson/M. Carrano]
52600 T. A. Tumanova. 1977. Novyye dannyye ob ankilozavre Tarchia gigantea [New data on the ankylosaur Tarchia gigantea]. Paleontologicheskii Zhurnal 1977(4):92-100 [M. Carrano/M. Carrano/M. Carrano]
52601 T. A. Tumanova. 1978. New data on the ankylosaur Tarchia gigantea. Paleontological Journal 1977(4):480-486 [M. Carrano/M. Carrano/M. Carrano]
26796ETE T. A. Tumanova. 1987. Pantsirnyye dinozavry Mongolii [The armored dinosaurs of Mongolia]. Trudy Sovmestnaya Sovetsko-Mongol'skaya Paleontologicheskaya Ekspeditsiya 32:1-80 [M. Carrano/M. Carrano/M. Carrano]
3793 R. J. Zakrzewski. 1985.  A New Species of Woodrat (Cricetidae) from the Pleistocene (Irvingtonian) of South Dakota. Journal of Mammalogy 66(4):770-773 [J. Alroy/J. Alroy/J. Ju]