Oise amber, Le Quesnoy: Ypresian, France

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Insecta - Thysanoptera - Uzelothripidae
Nel et al. 2013 2 specimens
MNHN-F.A38530 (PA 6816), A38531 (PA 1794)
Insecta - Thysanoptera - Melanthripidae
Nel et al. 2020 1 specimen
MNHN.F.A71309 (PA 3917) (2 measurements)
Insecta - Hemiptera - Dipsocoridae
Hartung et al. 2017 1 specimen
MNHN.F.A53793 (PA15518)
Insecta - Hemiptera - Tingidae
Nel et al. 2004 1 specimen
PA 2443
Insecta - Hemiptera - Miridae
Vernoux et al. 2010 4 specimens
MNHN F.A30851 (PA 8543), A30852 (PA 6524), A30853 (PA 8584), A30854 (PA 9493) (2 measurements)
Insecta - Hemiptera - Thaumastocoridae
Nel et al. 2004 2 specimens
PA 86, PA 7024
Insecta - Hemiptera - Blissidae
Garrouste et al. 2019 1 specimen
MNHN.F.A70574 (LQ192) (2 measurements)
Insecta - Hemiptera - Malcidae
Camier et al. 2019 1 specimen
MNHN.F.A71302 (PA 2495) (2 measurements)
Insecta - Hemiptera - Piesmatidae
Nel et al. 2004 1 specimen
PA 2306
Insecta - Hemiptera - Aradidae
Marchal et al. 2011 1 specimen
MNHN.F.A33749 (PA 12511) (1 measurement)
Insecta - Hemiptera - Aleyrodidae
Drohojowska and Szwedo 2013 10 specimens
MNHN PA 1297, 2287, 2474, 42, 154, 420, 1607, 5072, 1796, 5109
Aleurodicinae indet. Quaintance and Baker 1913
Drohojowska and Szwedo 2013 1 specimen
MNHN PA 17081
Drohojowska and Szwedo 2013 1 specimen
MNHN PA 1671
Drohojowska and Szwedo 2013 1 specimen
MNHN PA 2394 (3 measurements)
Drohojowska and Szwedo 2013 1 specimen
MNHN PA 5149 (3 measurements)
Drohojowska and Szwedo 2013 1 specimen
MNHN PA 16445 (3 measurements)
Drohojowska and Szwedo 2013 1 specimen
MNHN PA 2493 (3 measurements)
Insecta - Hemiptera - Cixiidae
Szwedo et al. 2006 1 specimen
MNHN PA 2427 1/2 (1 measurement)
Szwedo et al. 2006 1 specimen
MNHN PA 2770 (1 measurement)
Szwedo et al. 2006 1 specimen
MNHN PA 1781 1/15 (1 measurement)
Insecta - Hemiptera - Ricaniidae
Stroinski and Szwedo 2011 1 specimen
PA 5155 (1 measurement)
Insecta - Hemiptera - Lophopidae
Szwedo 2011 1 specimen
MNHN PA 2378 (1 measurement)
Insecta - Psocodea - Psoquillidae
Nel et al. 2005 6 specimens
PA 427, 6832, 3908, 3048, 350
Insecta - Psocodea - Empheriidae
Nel et al. 2005 13 specimens
PA 724, 226, 1733, 1712, 1004, 4182, 5615, 5111, 2793; 4 in Alvarez-Parra et al. (2024) MNHN.F.A94330 (PA 9252), 94359 (PA 12544), 94370 (PA 15159), 94342 (PA 15170)
Álvarez‐Parra et al. 2024 1 specimen
MNHN.F.A71305 (PA 2945) (2 measurements)
Insecta - Psocodea - Lepidopsocidae
Álvarez-Parra and Nel 2022 1 specimen
MNHN-F.A71357 (2 measurements)
Nel et al. 2005 15 specimens
PA 1478, 57, LQ 72, PA 2787, 6850, 4967, 4407, 4947, 8110; 5 in Alvarez-Parra et al. (2024) MNHN.F.A94336 (PA 5289), 94364 (LQ 70), 71357 (RI 118b), 94346 (PA PSO.01)
Insecta - Psocodea - Psyllipsocidae
Nel et al. 2005 2 specimens
PA1732; 1 in Alvarez-Parra et al. (2024) MNHN.F.A94352 (PA 742)
Insecta - Psocodea - Lachesillidae
Nel et al. 2005 40 specimens
PA 2785, 447, 363, 1100, 1542, 1641, 1749, 5187, 12340, 1913, 6166; 26 in Alvarez-Parra et al. (2024)
Insecta - Psocodea - Archipsocidae
Nel et al. 2005 4 specimens
PA 1705, 363; 2 in Alvarez-Parra et al. (2024) MNHN.F.A94355 (PA 15665), 94351 (LQ 95)
Insecta - Psocodea - Amphientomidae
Nel et al. 2005 3 specimens
PA 4125, LQ 228, PA 2419
Insecta - Psocodea - Manicapsocidae
Nel et al. 2005 3 specimens
PA 4996; 2 in Alvarez-Parra et al. (2024) MNHN.F.A94340 (PA 1816), 94339 (PA 9543)
Nel et al. 2005 11 specimens
PA 2380, 2462, 7873, 233, 1518, 2382, 5452, 1837, 4845; 2 in Alvarez-Parra et al. (2024) MNHN.F.A94345 (PA 15541), 94368 (LQ 4903)
Insecta - Psocodea - Pachytroctidae
Nel et al. 2005 6 specimens
PA 2468 3/3, 3 1/3, 1804 1/5; 2468 1/3; 2 in Alvarez-Parra et al. (2024) MNHN.F.A94378 (PA 180 1/2), 94338 (PA 4087)
Insecta - Psocodea - Liposcelididae
Liposcelis sp. Motschulsky 1852
Álvarez‐Parra et al. 2024 2 specimens
Nel et al. 2005 2 specimens
PA 4151, 6135; 1 in Alvarez-Parra et al. (2024) MNHN.F.A94371 (PA 11381)
Insecta - Hymenoptera - Ichneumonidae
Viertler et al. 2022 1 specimen
MNHN.F.A71346 (PA-349) (2 measurements)
Menier et al. 2004 1 specimen
MNHN PA 2439 (3 measurements)
Viertler et al. 2022 1 specimen
MNHN.F.A71347 (PA2189) (2 measurements)
Insecta - Hymenoptera - Braconidae
Belokobylskij et al. 2010 1 specimen
MNHN A32944 (2 measurements)
Belokobylskij et al. 2010 1 specimen
MNHN A32945 (2 measurements)
Insecta - Hymenoptera
Nel and Prokop 2005 1 specimen
MNHN PA 139 (2 measurements)
Falières and Nel 2019 3 specimens
MNHN.F.A71232 (PA 2993 1/3, 2/3, 3/3) (3 measurements)
Insecta - Hymenoptera - Pirenidae
van de Kamp et al. 2014 1 specimen
MNHN PA 16311
Insecta - Hymenoptera - Megalyridae
Perrichot 2009 1 specimen
MNHN PA-48
Insecta - Hymenoptera - Vespidae
Eumeninae indet. (Leach 1815)
van de Kamp et al. 2014 1 specimen
MNHN PA 3145
Insecta - Hymenoptera - Megachilidae
Michez et al. 2007 1 specimen
MNHN PA 3190
Insecta - Hymenoptera - Crabronidae
Eopison menieri n. gen., n. sp. Nel 2005
Nel 2005 5 specimens
PA 340, 2388, 1931, 2384(6/7), 2502
Nel 2005 7 specimens
PA 12238(1/4), 1129, 2360(1/4), 2370, 2527, 2363, 2470
Insecta - Hymenoptera - Scolebythidae
Lacau et al. 2000 2 specimens
PA 2395, PA 1293
Insecta - Hymenoptera - Dryinidae
Peinado et al. 2006 1 specimen
MNHN PA 2378 1/2
Insecta - Hymenoptera - Embolemidae
Chény et al. 2020 1 specimen
MNHN F.A58662 (1 measurement)
Insecta - Hymenoptera - Bethylidae
Perrichot and Nel 2008 1 specimen
MNHN PA 6470
Epyrini indet. Kieffer 1914
Perrichot and Nel 2008 1 specimen
MNHN PA 6744 1/3
Epyris moulyi n. sp. Falieres and Nel 2018
Colombo et al. 2021 1 specimen
MNHN.F.A70620 (PA 16149)
Perrichot and Nel 2008 1 specimen
MNHN PA 186
Falières and Nel 2020 1 specimen
MNHN.F.A71308 (PA 16948) (2 measurements)
Falières and Nel 2019 1 specimen
MNHN.F.A71187 (PA 16799)
Falières and Nel 2019 1 specimen
MNHN.F.A71156 (PA 6176) (2 measurements)
Falières and Nel 2019 1 specimen
MNHN.F.A70916 (PA 3389) (1 measurement)
Falières and Nel 2019 1 specimen
MNHN.F.A70917 (PA 8508) (2 measurements)
Falières and Nel 2019 1 specimen
MNHN.F.A71123 (PA 3673) (2 measurements)
De Ploëg and Nel 2004 1 specimen
MNHN PA 2436 (3 measurements)
Insecta - Hymenoptera - Formicidae
Aria et al. 2011
(1 measurement)
Aria et al. 2023 4 specimens
MNHN.F.A32941, 32938, 32939, 32940 (2 measurements)
Insecta - Hymenoptera - Aulacidae
Nel et al. 2004 1 specimen
PA 2523
Insecta - Hymenoptera - Evaniidae
Nel et al. 2002 4 specimens
MNHN PA 2494, PA 13(1/2, PA 221(1/2), PA 2818
Insecta - Coleoptera - Buprestidae
Bílý and Kirejtshuk 2007 1 specimen
(1 measurement)
Insecta - Coleoptera - Throscidae
Kovalev et al. 2012 1 specimen
MNHN-F.A46040 (PA 1673)
Insecta - Coleoptera - Dermestidae
Kirejtshuk et al. 2010 9 specimens
MNHN PA 2307, PA 160, PA 135, PA 15552, PA 12256, PA 12222, PA 438
Kirejtshuk et al. 2010 4 specimens
MNHN PA 336, PA 382, PA 12377, PA 2262
Kirejtshuk et al. 2010 2 specimens
MNHN PA 2501, PA 1581
Kirejtshuk et al. 2010 3 specimens
MNHN PA 8813, PA 1842, PA 164
Kirejtshuk et al. 2010 1 specimen
MNHN PA 6520
Kirejtshuk et al. 2010 1 specimen
MNHN PA 1387
Insecta - Coleoptera - Boleopsidae
Kirejtshuk and Nel 2013 1 specimen
MNHN 1389 (1 measurement)
Insecta - Coleoptera - Scirtidae
Kirejtshuk and Nel 2013 1 specimen
MNHN 7448 (1 measurement)
Kirejtshuk and Nel 2008 2 specimens
PA 3404 (2 measurements)
Kirejtshuk and Nel 2008 1 specimen
PA 5605 (1 measurement)
Insecta - Coleoptera - Melyridae
Kirejtshuk and Nel 2008 2 specimens
PA 890, 9094 (2 measurements)
Kirejtshuk and Nel 2008 3 specimens
PA 9530, 2764(3/6), 1781(10/15) (3 measurements)
Insecta - Coleoptera - Salpingidae
Kirejtshuk and Nel 2009 1 specimen
MNHN PA 891/3
Insecta - Coleoptera - Anthicidae
Kirejtshuk and Nel 2008 1 specimen
PA 1154, 1781(11/15) (1 measurement)
Insecta - Coleoptera - Mycteridae
Kirejtshuk and Nel 2009 1 specimen
MNHN PA 326
Insecta - Coleoptera - Tenebrionidae
Kirejtshuk et al. 2010
(1 measurement)
Insecta - Coleoptera - Ripiphoridae
Batelka et al. 2006 2 specimens
MNHN PA 2519, PA 1745 (4 measurements)
Insecta - Coleoptera - Silvanidae
Kirejtshuk and Nel 2013 1 specimen
MNHN PA-15066 (1 measurement)
Insecta - Coleoptera - Smicripidae
Kirejtshuk and Nel 2008 1 specimen
PA 4167 (1 measurement)
Insecta - Coleoptera - Nitidulidae
Kirejtshuk and Nel 2008 4 specimens
PA 5132, 2451, 2545, 1032 (4 measurements)
Kirejtshuk and Nel 2008 1 specimen
PA 8980 (1 measurement)
Kirejtshuk and Nel 2008 1 specimen
PA 602 (1 measurement)
Insecta - Coleoptera - Kateretidae
Kirejtshuk and Nel 2008 1 specimen
PA 9167 (1 measurement)
Kirejtshuk and Nel 2008 1 specimen
PA 1937 (1 measurement)
Insecta - Coleoptera - Brentidae
Kirejtshuk et al. 2015 7 specimens
MNHN A52061 (PA-15854); PA 9881, PA 12221, PA 533 1/2, PA 202 2/2, PA 5663, PA 268 2/2 in Legalov et al. 2019 (2 measurements)
Insecta - Coleoptera - Anthribidae
Legalov et al. 2019 1 specimen
8277
Legalov et al. 2019 1 specimen
MNHN.F.A66685 (PA 429) (2 measurements)
Insecta - Coleoptera - Curculionidae
Legalov et al. 2019 1 specimen
MNHN.F.A66690 (15247) (1 measurement)
Legalov et al. 2019 1 specimen
PA 1218
Legalov et al. 2019 1 specimen
7587
Legalov et al. 2019 1 specimen
MNHN.F.A66691 (PA 7600) (1 measurement)
Legalov et al. 2019 1 specimen
MNHN.F.A66687 (12213) (1 measurement)
Legalov et al. 2019 1 specimen
MNHN.F.A66688 (PA 2524) (1 measurement)
Legalov et al. 2019 1 specimen
MNHN.F.A66689 (PA 2469) (1 measurement)
Legalov et al. 2019 1 specimen
MNHN.F.A66686 (PA 1339) (1 measurement)
Insecta - Coleoptera - Chrysomelidae
Moseyko et al. 2010 1 specimen
PA 2437
Moseyko et al. 2010 1 specimen
PA 2364
Moseyko et al. 2010 1 specimen
PA 2802
Insecta - Coleoptera - Coccinellidae
Kirejtshuk and Nel 2012 6 specimens
MNHN PA 11862, 972, 245, 5388, 4840, 5162
Kirejtshuk and Nel 2012 1 specimen
MNHN PA 1290
Kirejtshuk and Nel 2012 1 specimen
MNHN PA 1047
Insecta - Coleoptera - Latridiidae
Kirejtshuk and Nel 2009 1 specimen
MNHN PA 394
Kirejtshuk and Nel 2009 1 specimen
MNHN PA 2381
Insecta - Coleoptera - Endomychidae
Kirejtshuk and Nel 2009 1 specimen
MNHN PA 1052
Insecta - Coleoptera - Micromalthidae
Kirejtshuk et al. 2010 1 specimen
PA 7870 (1 measurement)
Insecta - Coleoptera - Cupedidae
Kirejtshuk et al. 2010 1 specimen
PA 6457 (2 measurements)
Insecta - Megaloptera - Sialidae
Nel et al. 2002 1 specimen
PA 2285
Insecta - Neuroptera - Coniopterygidae
Nel et al. 2005 1 specimen
PA 28 (4/13)
Coniopteryx sp. Curtis 1834
Sziráki and Gröhn 2015 1 specimen
MNHN PA 8980 (3/3) (paratype of Gallosemidalis eocenica)
Insecta - Neuroptera - Rhachiberothidae
Nel et al. 2005 1 specimen
MNHN PA 346(1)
Insecta - Neuroptera - Sisyridae
Nel et al. 2003 16 specimens
PA 2789, 156, 11, 52, 157, 2329, 2478/1, 2478/2, 2493, 2812, 3921, 4144, 4864, 5368, 5374, 5603
Insecta - Diptera - Bombyliidae
Nel and De Ploeg 2004 1 specimen
PA 8334
Nel 2006 1 specimen
PA 208
Maheu and Nel 2020 1 specimen
MNHN.F.A71306 (1 measurement)
Insecta - Diptera - Mythicomyiidae
Nel 2006 8 specimens
PA 629, 154, 2-7/7
Myskowiak et al. 2016 1 specimen
MNHN.F.A57338 (2 measurements)
Nel and De Ploeg 2004 1 specimen
PA 2358
Insecta - Diptera - Scenopinidae
Garrouste et al. 2016 1 specimen
MNHN PA 16841 (2 measurements)
Insecta - Diptera - Hybotidae
Myskowiak et al. 2018 1 specimen
MNHN F.A57316
Insecta - Diptera - Dolichopodidae
Bramuzzo and Nel 2017 1 specimen
MNHN PA 979 (2 measurements)
Ngô-Muller et al. 2020 1 specimen
MNHN.F.A71320 (1 measurement)
Insecta - Diptera - Rhagionidae
Nel et al. 2016 1 specimen
MNHN.F.A57537 (PA 2496) (2 measurements)
Insecta - Diptera - Athericidae
Myskowiak and Nel 2014 4 specimens
MNHN.F.A51153 (PA 9055 1/6), A51152 (PA 106 1/2), PA 9055 2/6, 9055 3/6 (2 measurements)
Insecta - Diptera - Sciaridae
Camier and Nel 2019 1 specimen
MNHN.F.A71243 (LQ 105) (2 measurements)
Insecta - Diptera - Keroplatidae
Blagoderov et al. 2010 3 specimens
MNHN A32914 (2 measurements)
Insecta - Diptera - Cecidomyiidae
Nel and Prokop 2006 3 specimens
PA 3004, 5468
Nel 2021 1 specimen
MNHN.F.A71373 (PA 12413 1/2) (1 measurement)
Nel and Prokop 2006 1 specimen
PA 1678
Nel and Prokop 2006 2 specimens
PA 1599
Nel and Prokop 2006 7 specimens
PA 5314, 3296, 2540, 1303, 5135, 8106, 157
Nel and Prokop 2006 2 specimens
PA 1842, 1608
Nel 2021 1 specimen
MNHN.F.A71372 (PA 128 1/2) (1 measurement)
Insecta - Diptera - Mycetophilidae
Camier and Nel 2020 2 specimens
MNHN.F.A71245 (PA 3071) (2 measurements)
Insecta - Diptera - Anisopodidae
Camier and Nel 2019 1 specimen
MNHN.F.A71235 (PA 2890 2/2) (1 measurement)
Camier and Nel 2019 2 specimens
MNHN.F.A71231, MNHN.F.A.71234 (PA 8503) (3 measurements)
Insecta - Diptera - Bibionidae
Gee et al. 2001 4 specimens
PA 74 2/5, PA 74 3/5, PA 74 4/5, PA 2549 1/2
Insecta - Diptera - Psychodidae
Nel et al. 2002 4 specimens
MNHN PA 355, 353, 73, 74
Insecta - Diptera - Scatopsidae
Nel and Prokop 2004 3 specimens
MNHN PA 2359, 2440, 69-2/3 (2 measurements)
Bramuzzo et al. 2017 1 specimen
MNHN PA 16627 1–2/3
Insecta - Diptera - Limoniidae
Kopeć et al. 2025 2 specimens
MNHN.F.A97853, 97854 (2 measurements)
Kopeć et al. 2025 3 specimens
MNHN.F.A97848, 97849, 97850 (2 measurements)
Kopeć et al. 2025 2 specimens
MNHN.F.A97852, 97851 (2 measurements)
Insecta - Diptera - Chironomidae
Doitteau and Nel 2007 1 specimen
MNHN PA 1077 (1 measurement)
Doitteau and Nel 2007 1 specimen
MNHN PA 1161 (1 measurement)
Doitteau and Nel 2007 2 specimens
MNHN PA 2267 2/2, 7546 (1 measurement)
Doitteau and Nel 2007 1 specimen
MNHN PA 9182 (1 measurement)
Doitteau and Nel 2007 2 specimens
MNHN PA 12425, 4321 (1 measurement)
Doitteau and Nel 2007 2 specimens
MNHN PA 12709, 692
Doitteau and Nel 2007 1 specimen
MNHN PA 1683 (1 measurement)
Doitteau and Nel 2007 1 specimen
MNHN PA 4273 (1 measurement)
Doitteau and Nel 2007 1 specimen
MNHN PA 709
Doitteau and Nel 2007 1 specimen
MNHN PA 461
Doitteau and Nel 2007 1 specimen
MNHN PA 11432
Doitteau and Nel 2007 1 specimen
MNHN PA 15016 (1 measurement)
Doitteau and Nel 2007 5 specimens
MNHN PA 132 (3/12) and two female paratypes; PA 9 (1/6), 2894 (1 measurement)
Doitteau and Nel 2007 1 specimen
MNHN PA 685 (2 measurements)
Doitteau and Nel 2007 1 specimen
MNHN PA 11435 (1 measurement)
Doitteau and Nel 2007 1 specimen
MNHN PA 9563 (2 measurements)
Doitteau and Nel 2007 2 specimens
MNHN PA 1413, 318 (3/3) (1 measurement)
Doitteau and Nel 2007 1 specimen
MNHN PA 2818 (1 measurement)
Doitteau and Nel 2007 2 specimens
MNHN PA 2355, 1742 (4/6) (1 measurement)
Doitteau and Nel 2007 153 specimens
(1 measurement)
Doitteau and Nel 2007 6 specimens
MNHN PA 1652, 2964, 944 (1,2/2), 132 (2/12), 944 (1/2) (1 measurement)
Doitteau and Nel 2007 3 specimens
MNHN PA 5502, 7945, 8127 (1 measurement)
Doitteau and Nel 2007 1 specimen
MNHN PA 6543 (1 measurement)
Doitteau and Nel 2007 1 specimen
MNHN PA 276 1/2
Doitteau and Nel 2007 1 specimen
MNHN PA 3813 (2 measurements)
Doitteau and Nel 2007 2 specimens
MNHN PA 2505, 501(1/2) (2 measurements)
Doitteau and Nel 2007 3 specimens
MNHN PA 15044, 145 1/3, 15001 (2 measurements)
Doitteau and Nel 2007 1 specimen
MNHN PA 534 (1 measurement)
Doitteau and Nel 2007 1 specimen
MNHN PA 428 1/2 (1 measurement)
Doitteau and Nel 2007 17 specimens
PA 2347, 10550, 9761, 1263, 6575, 454, 15030, 1182, 5997, 2709, 8177, 6469, 171 2/2, 3798, 4246, 2871, 2752 (1 measurement)
Insecta - Gallophasmatidae
Nel et al. 2010 3 specimens
MNHN A30855 (PA 16000), A30856 (PA 1574), A30857 (PA 16095)
Insecta - Orthoptera - Tridactylidae
Azar and Nel 2008 1 specimen
PA 15270
Insecta - Embioptera - Embiidae
Falières et al. 2021 1 specimen
MNHN.F.A71313 (PA 17338) (1 measurement)
Insecta - Dermaptera
Nel et al. 2003 2 specimens
PA 29, 205
Insecta - Mantodea - Mantoididae
Schubnel and Nel 2019 2 specimens
MNHN-F.A71141 (PA 2378 2/2), with doubt MNHN-F.A71140
Insecta - Mantodea - Chaeteessidae
Chaeteessidae indet. Handlirsch 1925
Schubnel and Nel 2019 2 specimens
MNHN-F.A71139 (PA 3309), MNHN-F.A71142 (PA 2420)
Insecta - Blattodea - Mastotermitidae
Nel and Bourguet 2006 3 specimens
MNHN PA 2405, 3177, 3997 (1 measurement)
Insecta - Blattodea - Kalotermitidae
Nel and Bourguet 2006 4 specimens
MNHN PA 2542, 2404, 2383, 3141 (3 measurements)
Insecta - Odonata - Euphaeidae
Nel 2022 1 specimen
MNHN.F.A71361
Insecta - Odonata
Fleck et al. 2000 1 specimen
MNHN PA 2410
Arachnida - Araneae - Archaeidae
Carbuccia et al. 2020 1 specimen
MNHN.F.A71310 (PA 12262)
Arachnida - Araneae - Pholcidae
Penney 2007 1 specimen
MNHN PA 3148
Arachnida - Araneae - Oonopidae
Penney 2007 2 specimens
PA 759, 1909
Arachnida - Araneae - Anapidae
Penney et al. 2007 1 specimen
MNHN PA 327
Arachnida - Araneae - Selenopidae
Selenops sp. Latreille 1819
Penney 2007 1 specimen
MNHN PA 2375
see common names

Geography
Country:France State/province:Oise
Coordinates: 49.3° North, 2.7° East (view map)
Paleocoordinates:40.4° North, 9.6° East (Wright 2013)
Basis of coordinate:based on nearby landmark
Time
Period: Paleogene Epoch: Eocene
Stage: Ypresian 10 m.y. bin: Cenozoic 2
Key time interval: Ypresian
Age range of interval: 56 - 48.07 m.y. ago
Stratigraphy
Formation:Argiles d'lignite du Soissonnais
Stratigraphic resolution:group of beds
Stratigraphy comments: Lowermost Eocene, Sparnacian, level MP7 of the mammal fauna of Dormaal, early Ypresian
Lithology and environment
Primary lithology: amber
Includes fossils?Y
Environment:terrestrial indet.
Taphonomy
Modes of preservation:body,soft parts,amber
Size of fossils:macrofossils,microfossils
Collection methods and comments
Collection methods:bulk,field collection
Reason for describing collection:taxonomic analysis
Museum repositories:MNHN
Collection method comments: Includes fossil material from amber in the Le Quesnoy outcrop, representing a different source fauna than the fish/amphibians/etc.
Metadata
Database number:123850
Authorizer:M. Clapham, J. Alroy Enterer:M. Clapham, J. Alroy, A. Delelli, A. Noshirvan, J. Karr
Modifier:G. Varnham Research group:paleoentomology
Created:2012-02-01 04:27:57 Last modified:2025-02-22 15:12:02
Access level:the public Released:2012-02-01 04:27:57
Creative Commons license:CC0
Reference information

Primary reference:
19606. A. Nel, G. de Plöeg, J. Dejax, D. Duthiel, D. Franceschi, E. Gheerbrant, M. Godinot, S. Hervet, J.-J. Menier, M. Augé, G. Bignot, C. Cavagnetto, S. Duffaud, J. Gaudant, S. Hua, A. Jossang, F. Lapparent Broin, J.-P. Pozzi, J.-C. Paicheler, F. Beuchet, and J.-C. Rage. 1999. Un gisement sparnacien exceptionnel à plantes, arthropodes et vertébrés (Éocène basal, MP7): Le Quesnoy (Oise, France). Comptes Rendus de l'Académie des Sciences - Series IIA - Earth and Planetary Science 329(1):65-72 [P. Holroyd/P. Holroyd]

Secondary references:

85104 S. Álvarez-Parra and A. Nel. 2022. A new genus of setose-winged barklice (Psocodea: Trogiomorpha: Lepidopsocidae) from the Eocene amber of Oise with notes on the biogeography of Thylacellinae. Historical Biology 35:1136-1145 [M. Clapham/M. Clapham]
84261 C. Aria, C. Jouault, V. Perrichot and A. Nel. 2023. The megathermal ant genus Gesomyrmex (Formicidae: Formicinae), palaeoindicator of wide latitudinal biome homogeneity during the PETM. Geological Magazine 160:187-197 [M. Clapham/M. Clapham]
36397 C. Aria, V. Perrichot, and A. Nel. 2011. Fossil Ponerinae (Hymenoptera: Formicidae) in Early Eocene amber of France. Zootaxa 2870:53-62 [M. Clapham/M. Clapham]
39981 D. Azar and A. Nel. 2008. First Tridactylidae from the Eocene French amber (Insecta: Orthoptera). Alavesia 2:169-175 [M. Clapham/A. Delelli]
90212 S. Álvarez‐Parra, A. Nel, and D. Azar. 2024. Overview of Psocodea in Eocene Oise amber and implications for the transition to modern‐day barklice communities. Papers in Palaeontology 10(e1609) [M. Clapham/M. Clapham]
40659 J. Batelka, F. Collomb, and A. Nel. 2006. Macrosiagon deuvei n. sp. (Coleoptera: Ripiphoridae) from the French Eocene amber. Annales de la Société Entomologique de France 42(1):75-78 [M. Clapham/A. Delelli]
40580 S. A. Belokobylskij, A. Nel, A. Waller and G. Plöeg. 2010. New fossil non-cyclostome braconid wasps from the lowermost Eocene amber of Paris Basin. Acta Palaeontologica Polonica 55(3):519-527 [M. Clapham/A. Noshirvan]
48352 S. Bílý and A. G. Kirejtshuk. 2007. Philanthaxoides gallicus gen. nov. sp. nov. from the lowermost Eocene French amber (Coleoptera: Buprestidae). Folia Heyrovskyana, Series A 14:181-186 [M. Clapham/M. Clapham/M. Clapham]
40653 V. A. Blagoderov, H. Hippa, and A. Nel. 2010. Parisognoriste, a new genus of Lygistorrhinidae (Diptera: Sciaroidea) from the Oise amber with redescription of Palaeognoriste Meunier. ZooKeys 50:79-90 [M. Clapham/A. Delelli/J. Karr]
66255 S. Bramuzzo, D. Coty, and A. Nel. 2017. A new species of Ferneiella from the Eocene French amber (Diptera: Scatopsidae). Zootaxa 4350:177-184 [M. Clapham/M. Clapham]
61601 S. Bramuzzo and A. Nel. 2017. Youngest representative of the extinct genus Microphorites in the Eocene amber of France (Diptera: Dolichopodidae: Microphorinae). Zootaxa 4231:590-594 [M. Clapham/M. Clapham]
77644 M. Camier, A. Logghe, A. Nel and R. Garrouste. 2019. The first fossil Malcidae (Hemiptera, Heteroptera) discovered in the earliest Eocene Oise amber. Historical Biology 33:1208-1211 [M. Clapham/M. Clapham]
77418 M. Camier, A. Logghe, A. Nel and R. Garrouste. 2021. The first Malcidae (Hemiptera, Heteroptera) from the earliest Eocene Oise amber. Historical Biology 33:1024-1028 [M. Clapham/M. Clapham]
82531 M. Camier and A. Nel. 2019. First Mycetobiinae of the lowermost Eocene Oise amber (France) (Diptera: Anisopodidae). Palaeoentomology 2:245-250 [M. Clapham/M. Clapham]
82536 M. Camier and A. Nel. 2019. A fossil black fungus gnat from the lowermost Eocene amber of France (Diptera: Sciaridae). Palaeoentomology 2:316-321 [M. Clapham/M. Clapham]
78974 M. Camier and A. Nel. 2020. The oldest fungus gnat of the tribe Exechiini in the lowermost Eocene Oise amber (Diptera: Mycetophilidae). Zootaxa 4722:91-98 [M. Clapham/M. Clapham]
73867 B. Carbuccia, H. M. Wood, C. Rollad, A. Nel, and R. Garrouste. 2020. A new Myrmecarchaea (Araneae: Archaeidae) species from Oise amber (earliest Eocene, France). BSGF - Earth Sciences Bulletin 191(24):1-5 [M. Clapham/M. Clapham/M. Clapham]
73108 C. Chény, E. Guillam, A. Nel and V. Perrichot. 2020. A new species of Ampulicomorpha Ashmead from Eocene French amber, with a list of fossil and extant Embolemidae (Insecta: Hymenoptera) of the world. BSGF - Earth Sciences Bulletin 191(20):1-7 [M. Clapham/M. Clapham/M. Clapham]
76008 W. D. Colombo, E. E. Perkovsky, C. Waichert and C. O. Azevedo. 2021. Synopsis of the fossil flat wasps Epyrinae (Hymenoptera, Bethylidae), with description of three new genera and 10 new species. Journal of Systematic Palaeontology 19:39-89 [M. Clapham/M. Clapham]
44483 G. De Ploëg and A. Nel. 2004. A new bethylid wasp from the Lowermost Eocene amber of France (Hymenoptera: Bethylidae: Bethylinae). Geologica Acta 2:75-82 [M. Clapham/M. Clapham]
48464 G. Doitteau and A. Nel. 2007. Chironomid midges from early Eocene amber of France (Diptera: Chironomidae). Zootaxa 1404:1-66 [M. Clapham/A. Delelli]
46157 J. Drohojowska and J. Szwedo. 2013. The first Aleyrodidae from the lowermost Eocene Oise amber (Hemiptera: Sternorrhyncha). Zootaxa 3636:319-347 [M. Clapham/M. Clapham]
78306 E. Falières, M. S. Engel, and A. Nel. 2021. Earliest occurrence of Embiidae: A new genus from earliest Eocene Oise amber (Insecta: Embiodea). Geodiversitas 20:799-805 [M. Clapham/M. Clapham]
70185 E. Falières and A. Nel. 2019. The first sclerodermine and pristocerine flat wasps in Lowermost Eocene amber of France (Hymenoptera: Bethylidae). Comptes Rendus Palevol 18:509-515 [M. Clapham/M. Clapham/M. Clapham]
82537 E. Falières and A. Nel. 2019. The first representative of the extinct flat wasp subfamily Protopristocerinae (Insecta: Hymenoptera, Bethylidae). Palaeoentomology 2:322-326 [M. Clapham/M. Clapham]
82544 E. Falières and A. Nel. 2019. A new scelionine genus from the earliest Eocene Oise amber (Hymenoptera: Platygastroidea: Scelionidae). Palaeoentomology 2:418-424 [M. Clapham/M. Clapham]
82525 E. Falières and A. Nel. 2019. A new bethylid wasp from the Lowermost Eocene Oise amber, France (Insecta: Hymenoptera). Palaeoentomology 2:211-214 [M. Clapham/M. Clapham]
74993 E. Falières and A. Nel. 2020. First fossil of the extant flat wasp genus Dissomphalus Ashmead, 1893 (Hymenoptera: Bethylidae: Pristocerinae). Zoosystema 42:207-213 [M. Clapham/M. Clapham]
46743 G. Fleck, A. Nel, G. De Plöeg and G. Masselot. 2000. A fossil dragonfly from the Paris Basin amber of France (Lowermost Eocene) (Insecta, Odonata, Anisoptera). Acta Geologica Hispanica 35(1-2):131-134 [M. Clapham/J. Karr]
58593 R. Garrouste, D. Azar, and A. Nel. 2016. The oldest accurate record of Scenopinidae in the Lowermost Eocene amber of France (Diptera: Brachycera). Zootaxa 4093:444-450 [M. Clapham/M. Clapham]
82516 R. Garrouste, T. Schubnel, and A. Nel. 2019. The first chinch bug Blissidae (Hemiptera, Heteroptera) from the Eocene Oise amber. Palaeoentomology 2:85-93 [M. Clapham/M. Clapham]
40803 J. Gee, A. Nel, J. Menier and G. De Ploëg. 2001. A new lovebug fly (Insecta, Diptera) from the lowermost Eocene amber of the Paris Basi. Geodiversitas 23(3):341-348 [M. Clapham/A. Noshirvan]
63322 V. Hartung, R. Garrouste, and A. Nel. 2017. The first fossil Dipsocoridae found in the early Eocene amber of France (Hemiptera: Heteroptera). Comptes Rendus Palevol 16:715-720 [M. Clapham/M. Clapham]
40703 M. L. I. Judson and J. Makol. 2009. A mite of the family Tanaupodidae (Arachnida, Acari, Parasitengona) from the Lower Cretaceous of France. Geodiversitas 31(1):41-47 [M. Clapham/A. Delelli/A. Delelli]
40662 A. G. Kirejtshuk, J. Háva, and A. Nel. 2010. New genus and species of subfamily Trinodinae (Coleoptera, Polyphaga, Dermestidae) from Lowermost Eocene French amber. Zoosystematica Rossica 19(1):54-69 [M. Clapham/A. Delelli/M. Clapham]
57542 A. G. Kirejtshuk, A. A. Legalov, and A. Nel. 2015. A new genus of the subfamily Apioninae (Coleoptera: Brentidae) from the Lower Eocene Oise amber. Paleontological Journal 49:1436-1441 [M. Clapham/M. Clapham]
36717 A. G. Kirejtshuk, M. V. Nabozhenko, and A. Nel. 2010. New genus and species of the tribe Opatrini (Coleoptera, Tenebrionidae, Tenebrioninae) from the lowermost Eocene amber of Paris Basin. Proceedings of the Zoological Institute RAS 314(2):191-196 [J. Alroy/J. Alroy]
40603 A. G. Kirejtshuk and A. Nel. 2008. New beetles of the suborder Polyphaga from the lowermost Eocene French amber (Insecta: Coleoptera). Annales de la Société Entomologique de France 44:419-442 [M. Clapham/M. Clapham]
40657 A. G. Kirejtshuk and A. Nel. 2009. New genera and species of Cucujiformia (Coleoptera, Polyphaga) from lowermost Eocene French amber. Denisia 26:103-118 [M. Clapham/A. Delelli/M. Clapham]
36979 A. G. Kirejtshuk, A. Nel, and F. Collomb. 2010. New Archostemata (Insecta: Coleoptera) from the French Paleocene and Early Eocene, with a note on the composition of the suborder. Annales de la Société Entomologique de France 46(1-2):216-227 [M. Clapham/J. Karr]
44119 A. G. Kirejtshuk and A. Nel. 2012. The oldest representatives of the family Coccinellidae (Coleoptera: Polyphaga) from the lowermost Eocene Oise amber (France). Zoosystematica Rossica 21:131-144 [M. Clapham/M. Clapham]
51506 A. G. Kirejtshuk and A. Nel. 2013. Current knowledge of Coleoptera (Insecta) from the lowermost Eocene Oise amber. Insect Systematics & Evolution 44:175-201 [M. Clapham/M. Clapham/M. Clapham]
91895 K. Kopeć, W. Krzemiński, I. Kania-Kłosok, M. Syratt, A. Nel and A. Soszyńska. 2025. First evidence of Limoniidae (Diptera: Nematocera) in French amber from Oise. Scientific Reports 15(14472) [M. Clapham/M. Clapham]
44115 A. V. Kovalev, A. G. Kirejtshuk, and A. Nel. 2012. New species of the genus Trixagus Kugelann, 1794 (Coleoptera: Throscidae) from the lowermost Eocene amber of Oise (France). Proceedings of the Zoological Institute RAS 316:83-88 [M. Clapham/M. Clapham/M. Clapham]
40471 S. Lacau, A. Nel, C. Villemant, J. J. Menier, M. J. Orliac and G. De Plöeg. 2000. A fossil Scolebythidae from the lowermost Eocene amber of France (Insecta: Hymenoptera). Annals of the Entomological Society of America 93:701-706 [M. Clapham/M. Clapham]
71833 A. A. Legalov, A. G. Kirejtshuk, and A. Nel. 2019. New weevils (Coleoptera, Curculionoidea) from the earlymost Eocene Oise amber. Paleontological Journal 53:729-751 [M. Clapham/M. Clapham/M. Clapham]
71882 A. Maheu and A. Nel. 2020. The oldest representative of the bombyliid bee fly in the earliest Eocene French amber. Acta Palaeontologica Polonica 65:81-83 [M. Clapham/M. Clapham/M. Clapham]
56447 L. Marchal, E. Guilbert, P. Brisac and A. Nel. 2011. A new record and a new species of Aradidae fossils (Hemiptera: Heteroptera). Zootaxa 2832:56-62 [M. Clapham/M. Clapham/M. Clapham]
46762 J. J. Menier, A. Nel, A. Waller and G. De Ploëg. 2004. A new fossil ichneumon wasp from the Lowermost Eocene amber of Paris Basin (France), with a checklist of fossil Ichneumonoidea s. l. (Insecta: Hymenoptera: Ichneumonidae: Metopiinae). Geologica Acta 2(1):83-94 [M. Clapham/J. Karr]
40707 D. Michez, A. Nel, J.J. Menier and P. Rasmont. 2007. The oldest fossil of a melittid bee (Hymenoptera: Apiformes) from the early Eocene of Oise (France). Zoological Journal of the Linnean Society 150:701-709 [M. Clapham/A. Delelli]
39648 A. G. Moseyko, A. G. Kirejtshuk, and A. Nel. 2010. New genera and new species of leaf beetles (Coleoptera: Polyphaga: Chrysomelidae) from lowermost Eocene French amber. Annales de la Société Entomologique de France 46:116-123 [M. Clapham/M. Clapham]
59987 J. Myskowiak, R. Garrouste, and A. Nel. 2016. A new genus and species of micro bee fly from the Earliest Eocene French amber (Diptera: Mythicomyiidae: Psiloderoidinae). Zootaxa 4114:583-586 [M. Clapham/M. Clapham]
66256 J. Myskowiak, R. Garrouste, and A. Nel. 2018. Eodromyia pumilio gen. et sp. nov., a new empidoid fly from the earliest Eocene amber of France (Diptera: Hybotidae: Tachydromiinae). Zootaxa 4379:279-286 [M. Clapham/M. Clapham]
54491 J. Myskowiak and A. Nel. 2014. A new genus and species of ibis fly in the Lowermost Eocene amber of Oise (France) (Diptera: Athericidae). Zootaxa 3869:372-382 [M. Clapham/M. Clapham]
41259 A. Nel. 2005. Oldest representatives of the Sphecidae: Trypoxylini in the early Eocene French amber (Insecta: Hymenoptera). Comptes Rendus Palevol 4:17-24 [M. Clapham/M. Clapham/M. Clapham]
39549 A. Nel. 2006. Oldest records of Bombyliidae: Phthiriinae and Mythicomyiidae: Glabellulinae from the Lowermost Eocene amber of France (Diptera: Bombylioidea). European Journal of Entomology 103(1):109-114 [M. Clapham/J. Karr]
84037 A. Nel. 2021. New Cecidomyiidae from the lowermost Eocene French amber (Diptera). Palaeoentomology 4:620-628 [M. Clapham/M. Clapham]
83479 A. Nel. 2022. The second European representative of the epallagid genus Labandeiraia in the lowermost Eocene Oise amber (Odonata, Zygoptera). Palaeoentomology 5:520-523 [M. Clapham/M. Clapham]
50784 A. Nel and E. Bourguet. 2006. Termite of the early Eocene amber of France (Isoptera: Mastotermitidae, Kalotermitidae). Neues Jahrbuch für Geologie und Paläontologie, Monatshefte 2006:101-115 [M. Clapham/M. Clapham]
40845 A. Nel and G. De Ploeg. 2004. New fossil bee flies (Diptera: Bombylioidea) in the Lowermost Eocene amber of the Paris Basin. Geologica Acta 2(1):57-65 [M. Clapham/A. Delelli]
41260 A. Nel, E. Delfosse, T. Robillard and J. F. Petrulevičius. 2010. An early winged crown group stick insect from the Early Eocene amber of France (Insecta, Phasmatodea). Systematic Entomology 35:340-346 [M. Clapham/M. Clapham/M. Clapham]
48716 A. Nel, J. J. Menier, and G. De Plöeg. 2002. The oldest representative of Trichomyiinae from the lowermost Eocene amber of the Paris basin (France) (Diptera: Psychodidae). Annales de la Société Entomologique de France 38:247-252 [M. Clapham/M. Clapham]
39909 A. Nel, J. Menier, G. De Ploeg, G. Hodebert, and L. Danvin. 2002. Eosialis, a new alderfly genus in French Lowermost Eocene amber (Insecta, Megaloptera, Sialidae). Geobios 35:313-319 [M. Clapham/A. Delelli/M. Clapham]
39865 A. Nel, J. J. Menier, A. Waller, G. Hodebert, and G. Ploëg. 2003. New fossil spongilla-flies from the lowermost Eocene amber of France (Insecta, Neuroptera, Sisyridae). Geodiversitas 25:109-117 [M. Clapham/M. Clapham]
60865 A. Nel, Z. Perreau, and G. Doitteau. 2016. The oldest representative of the modern snipe fly genus Symphoromyia (Diptera: Rhagionidae). Zootaxa 4196:144-150 [M. Clapham/M. Clapham]
39100 A. Nel, V. Perrichot, and D. Azar. 2005. New and poorly known fossil Coniopterygidae in Cretaceous and Cenozoic ambers (Insecta: Neuroptera). Annales Zoologici 55:1-7 [M. Clapham/M. Clapham]
47992 A. Nel, V. Perrichot, D. Azar and D. Néraudeau. 2005. New Rhachiberothidae (Insecta: Neuroptera) in Early Cretaceous and Early Eocene ambers from France and Lebanon. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 235:51-85 [M. Clapham/M. Clapham/M. Clapham]
51009 A. Nel and J. Prokop. 2004. New Paleogene Scatopsidae (Diptera: Nematocera) in amber from the Czech Republic and France. Acta Societatis Zoologicae Bohemicae 68:91-98 [M. Clapham/M. Clapham]
50150 A. Nel and J. Prokop. 2005. New fossil Scelionidae (Insecta: Hymenoptera) in early Paleogene amber from eastern Moravia (Czech Republic) and northern France. Polskie Pismo Entomologiczne 74:339-347 [M. Clapham/M. Clapham]
39913 A. Nel, J. Prokop, G. De Ploeg and J. Millet. 2005. New Psocoptera (Insecta) from the lowermost Eocene Amber of Oise, France. Journal of Systematic Palaeontology 3(4):371-391 [M. Clapham/A. Delelli/M. Clapham]
40584 A. Nel and J. Prokop. 2006. New fossil gall midges from the earliest Eocene French amber (Insecta, Diptera, Cecidomyiidae). Geodiversitas 28(1):37-54 [M. Clapham/A. Delelli]
48715 A. Nel, A. Waller, G. Hodebert and G. De Plöeg. 2002. An ensign-fly in the lowermost Eocene amber from the Paris basin (Hymenoptera: Evaniidae). Annales de la Société Entomologique de France 38:263-269 [M. Clapham/M. Clapham/M. Clapham]
39936 A. Nel, A. Waller, V. Albouy, J.-J. Menier, and G. De Plöeg. 2003. New fossil earwigs from the lowermost Eocene amber of Paris basin (France) (Insecta, Dermaptera, family incertae sedis). Geodiversitas 25(1):119-129 [M. Clapham/A. Delelli/P. Wagner]
39910 A. Nel, A. Waller, and G. De Ploeg. 2004. The oldest fossil piesmatid bug in the Lowermost Eocene amber of the Paris Basin (Heteroptera: Lygaeoidea: Piesmatidae). Geologica Acta 2(1):45-50 [M. Clapham/A. Delelli/A. Delelli]
39493 A. Nel, A. Waller, and G. Ploég. 2004. The oldest fossil Tingidae from the lowermost Eocene amber of the Paris Basin (Heteroptera: Cimicomorpha: Tingoidea). Geologica Acta 2:37-43 [M. Clapham/M. Clapham]
39911 A. Nel, A. Waller, and G. De Ploeg. 2004. The oldest palm bug in the Lowermost Eocene amber of the Paris Basin (Heteroptera: Cimicomorpha: Thaumastocoridae). Geologica Acta 2(1):51-55 [M. Clapham/A. Delelli]
39912 A. Nel, A. Waller, and G. De Ploeg. 2004. An aulacid wasp in the Lowermost Eocene amber from the Paris Basin (Hymenoptera: Aulacidae). Geologica Acta 2(1):67-74 [M. Clapham/A. Delelli]
41338 P. Nel, A. R. Schmidt, C. Bässler and A. Nel. 2013. Fossil thrips of the family Uzelothripidae suggest 53 million years of morphological and ecological stability. Acta Palaeontologica Polonica 58:609-614 [M. Clapham/M. Clapham/M. Clapham]
76884 P. Nel, T. Schubnel, V. Perrichot and A. Nel. 2020. Ankothrips, the most ancient extant thrips genus (Thysanoptera, Melanthripidae). Papers in Palaeontology 7:825-837 [M. Clapham/M. Clapham]
83541 V. Ngô-Muller, R. Garrouste, and A. Nel. 2020. The oldest long-legged fly of the subfamily Medeterinae (Diptera: Dolichopodidae) from the Early Eocene of France. Palaeoentomology 3:173-179 [M. Clapham/M. Clapham]
48636 J. Peinado, A. Nel, and A. Waller. 2006. A dryinid wasp in Early Eocene amber from the Paris basin (Hymenoptera: Dryinidae). Zootaxa 1168:31-41 [M. Clapham/M. Clapham]
40588 D. Penney. 2007. A new fossil oonopid spider in lowermost Eocene amber from the Paris Basin, with comments on the fossil spider assemblage. African Invertebrates 48(1):71-75 [M. Clapham/A. Delelli]
47387 D. Penney. 2007. The oldest fossil pholcid and selenopid spiders (Araneae) in lowermost Eocene amber from the Paris basin, France. The Journal of Arachnology 34:592-598 [M. Clapham/M. Clapham]
47390 D. Penney, M. Dierick, V. Cnudde, B. Masschaele, J. Vlassenbroeck, L. Van Hoorebeke, and P. Jacobs. 2007. First fossil Micropholcommatidae (Araneae), imaged in Eocene Paris amber using X-ray computed tomography. Zootaxa 1623:47-53 [M. Clapham/M. Clapham]
42084 V. Perrichot. 2009. Long-tailed wasps (Hymenoptera: Megalyridae) from Cretaceous and Paleogene European amber. Paleontological Contributions 1:1-35 [M. Clapham/M. Clapham]
47989 V. Perrichot and A. Nel. 2008. Eocene bethylid wasps from French amber (Hymenoptera: Bethylidae). Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 248:91-101 [M. Clapham/M. Clapham/M. Clapham]
71152 T. Schubnel and A. Nel. 2019. New Paleogene mantises from the Oise amber and their evolutionary importance. Acta Palaeontologica Polonica 64:779-786 [M. Clapham/M. Clapham/M. Clapham]
40509 A. Stroinski and J. Szwedo. 2011. Abraracourcix curvivenatus n. gen. n. sp. from the Lowermost Eocene Oise amber (Hemiptera: Fulgoromorpha: Ricaniidae). Annales de la Société Entomologique de France 47:480-486 [M. Clapham/M. Clapham]
60353 G. Sziráki and C. Gröhn. 2015. Presence of two extant genera of dusty lacewings (Neuroptera: Coniopterygidae) in Baltic amber, with remarks on some earlier described fossil taxa. Folia Historico-Naturalia Musei Matraensis 39:63-71 [M. Clapham/M. Clapham]
61283 J. Szwedo. 2011. Ordralfabetix sirophatanis gen. et sp. n. - the first Lophopidae from the lowermost Eocene Oise amber, Paris Basin, France (Hemiptera: Fulgoromorpha). Zootaxa 2822:52-60 [M. Clapham/M. Clapham]
48637 J. Szwedo, T. Bourgoin, and F. Lefebvre. 2006. New Mnemosynini taxa (Hemiptera, Fulgoromorpha: Cixiidae) from the Palaeogene of France with notes on their early association with host plants. Zootaxa 1122:25-45 [M. Clapham/M. Clapham]
61602 T. van de Kamp, T. Dos Santos Rolo, T. Baumbach and L. Krogmann. 2014. Scanning the past - synchrotron X-Ray microtomography of fossil wasps in amber. Entomologie Heute 26:151-160 [M. Clapham/M. Clapham/M. Clapham]
61977 J. Vernoux, R. Garrouste, and A. Nel. 2010. The first psallopinous bug from Lowermost Eocene French amber (Hemiptera: Heteroptera: Miridae). Zootaxa 2499:63-68 [M. Clapham/M. Clapham]
80359 A. Viertler, S. Klopfstein, C. Jouault and T. Spasojevic. 2022. Darwin wasps (Hymenoptera, Ichneumonidae) in Lower Eocene amber from the Paris basin. Journal of Hymenoptera Research 89:19-45 [M. Clapham/M. Clapham]