Humaca tracksite (SM14): Campanian, Bolivia
collected by D. Keremba Mamami, V. M. Rial Cez, C. Meyer, M. Lockley 1995, 1998

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia - Titanosauridae
11 individuals
trackways (306 total tracks)
Lockley et al. 2002
Reptilia - Theropoda
21 specimens
tracks
see common names

Geography
Country:Bolivia State/province:Chuquisaca
Coordinates: 19.1° South, 65.5° West (view map)
Paleocoordinates:20.9° South, 42.4° West (Wright 2013)
Basis of coordinate:stated in text
Altitude:2856 meters
Geographic resolution:outcrop
Time
Period: Cretaceous Epoch: Late Cretaceous
Stage: Campanian 10 m.y. bin: Cretaceous 7
Key time interval: Campanian
Age range of interval: 83.6 - 72.2 m.y. ago
Stratigraphy
Geological group:Puca Formation:Chaunaca
Stratigraphic resolution:bed
Stratigraphy comments: upper part of formation
Lithology and environment
Primary lithology:fine,brown silty sandstone
Includes fossils?Y
Lithology description: "The track-bearing layer is a fine grained, brownish sandstone"
Environment:fluvial indet.
Geology comments: "Perennial to ephemeral lacustrine shorelines"; "fluvial origin" for the track layer (Meyer et al., 2021)
Taphonomy
Modes of preservation:trace
Degree of concentration:-single event
Size of fossils:macrofossils
Spatial orientation:preferred
Preservation of anatomical detail:medium
Associated major elements:many
Disassociated major elements:some
Temporal resolution:snapshot
Spatial resolution:autochthonous
Collection methods and comments
Collection methods:surface (in situ),observed (not collected)
Reason for describing collection:taxonomic analysis
Collectors:D. Keremba Mamami, V. M. Rial Cez, C. Meyer, M. Lockley Collection dates:1995, 1998
Collection method comments: Site was discovered in 1995 by a group of amateurs including David Keremba Mamani and Vicente Manuel Rial Cez. The track-bearing part that has been mapped has a surface of about 750 m2 and yields a total of 327 individual footprints.
Metadata
Also known as:Ore-Pájla
Database number:36232
Authorizer:M. Carrano Enterer:M. Carrano
Modifier:M. Carrano Research group:vertebrate
Created:2004-01-19 15:09:44 Last modified:2025-02-22 15:12:02
Access level:the public Released:2004-01-19 15:09:44
Creative Commons license:CC0
Reference information

Primary reference:

82631. C. A. Meyer, D. Hippler, and M. G. Lockley. 2001. The Late Cretaceous vertebrate ichnofacies of Bolivia – facts and implications. VII International Symposium on Mesozoic Terrestrial Ecosystems. Asociación Paleontológica Argentina Publicación Especial 7:133-138 [M. Carrano/M. Carrano]

Secondary references:

76344 C. Cónsole-Gonella, S. de Valais, R. A. Marquillas and M. Cristina Sánchez. 2017. The Maastrichtian–Danian Maimará tracksite (Yacoraite Formation, Salta Group), Quebrada de Humahuaca, Argentina: environments and ichnofacies implications. Palaeogeography, Palaeoclimatology, Palaeoecology 468:327-350 [M. Carrano/M. Carrano]
9715 M. G. Lockley, A. S. Schulp, C. A. Meyer, G. Leonardi, and D. K. Mamani. 2002. Titanosaurid trackways from the Upper Cretaceous of Bolivia: evidence for large manus, wide-gauge locomotion and gregarious behaviour. Cretaceous Research 23:383-400 [M. Carrano/M. Carrano/M. Carrano]
76343 C. A. Meyer, D. Marty, and M. Belvedere. 2018. Titanosaur trackways from the Late Cretaceous El Molino Formation of Bolivia (Cal Orck’o, Sucre). Annales Societatis Geologorum Poloniae 88:223-241 [M. Carrano/M. Carrano]
76342 C. A. Meyer, D. Marty, B. Thüring, S. Thüring, and M. Belvedere. 2021. The Late Cretaceous dinosaur track record of Bolivia – review and perspective. Journal of South American Earth Sciences 106:102992 [M. Carrano/M. Carrano]
83929 K. Niu and L. Xing. 2023. The first dinosaur track assemblages from the Upper Cretaceous Shaxian Formation, Fujian Province, southeastern China. Cretaceous Research 146 [M. Carrano/M. Carrano]
55488 M. Romano, M. A. Whyte, and S. J. Jackson. 2007. Trackway ratio: A new look at trackway gauge in the analysis of quadrupedal dinosaur trackways and its implications for ichnotaxonomy. Ichnos 14(3–4):257-270 [M. Carrano/M. Carrano]
76345 M. Suárez Riglos, R. Céspedes Paz, and O. Medina Ramirez. 2019. Ruta de Migracíon de los Dinosaurios en Bolivia [Migration Route of Dinosaurs in Bolivia] 1-104 [M. Carrano/M. Carrano]