PBDB Collection

Shestakovo-1: Aptian, Russian Federation
collected by A. Mossakovsky, I. Lebedev, E. Mashchenko 1953–1960, 1994–1999, 2014–

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

unclassified

	Ginkgocycadophytus sp. Samoilovitch 1953	Bugdaeva et al. 2017

	Taurocusporites reduncus Stover 1962	Bugdaeva et al. 2017
Pinopsida - Pinales - Pinaceae

	Pinaceae indet. Rudolphi 1830	Bugdaeva et al. 2017
unclassified

	Pilosisporites setiferus	Bugdaeva et al. 2017
Polypodiopsida - Gleicheniaceae

	Gleicheniidites senonicus Ross 1949	Bugdaeva et al. 2017
Pteridopsida

	Osmundacidites wellmanii Couper 1953	Bugdaeva et al. 2017
Amphibia - Temnospondyli

	Caudata indet. (Scopoli 1777)	Ivantsov et al. 2016
Amphibia - Temnospondyli - Urodela

	Kiyatriton leshchinskiyi n. gen., n. sp. Averianov and Voronkevich 2002	Averianov and Voronkevich 2002	6 specimens
Reptilia - Pterosauria

	Pterodactyloidea indet. (Plieninger 1901)	Ivantsov et al. 2016
Reptilia - Pterosauria - Ornithocheiridae

	Ornithocheiridae indet. Seeley 1870	Skutschas 2006	3 elements
	collection of PM TGU, three teeth
Reptilia

	Coelurosauria informal indet. A	Averianov et al. 2019	31 specimens
	LMCCE 004/5, 53, 86, 125 (Dromaeosauridae or Tyrannosauroidea)

	Coelurosauria informal indet. B	Averianov et al. 2019	10 specimens
	LMCCE 004/74, 83 (Dromaeosauridae or Tyrannosauroidea)

	Coelurosauria informal indet. D	Averianov et al. 2019	7 specimens
	LMCCE 004/13, 26, 51, 52, 116 (Dromaeosauridae or Tyrannosauroidea)

	Deinonychosauria informal indet. C	Averianov et al. 2019	14 specimens
	LMCCE 004/32, 44, 61, 62 (Microraptorinae or Troodontidae)
Reptilia - Troodontidae

	Troodontidae indet. Gilmore 1924	Averianov and Sues 2007	3 specimens
	PM TGU 16/5-124

	cf. Urbacodon sp. Averianov and Sues 2007	Averianov et al. 2019	5 specimens
	LMCCE 004/1, 29, 78 (morphotype E)
Reptilia - Dromaeosauridae

	Dromaeosauridae indet. Colbert and Russell 1969	Averianov et al. 2002

	cf. Dromaeosauridae indet. Colbert and Russell 1969	Averianov et al. 2004
Reptilia

	Aves indet. Linnaeus 1758	Skutschas 2006
Reptilia - Mystiornithiformes - Mystiornithidae

	Mystiornis cyrili n. gen., n. sp. Kurochkin et al. 2011	Kurochkin et al. 2011	1 specimen
	PM TSU 16/5–45 - holotype (left metatarsus)
Reptilia

	Therizinosauroidea indet. Russell and Dong 1994	Ivantsov et al. 2016

	cf. Prodeinodon sp. Osborn 1924	Averianov et al. 2004
Reptilia - Noasauridae

	Coelurosauria indet. (Huene 1914)	Averianov and Lopatin 2023	1 specimen
	PIN 329/16, incomplete anterior or middle cervical vertebra
	= Kiyacursor longipes Averianov et al. 2024	Averianov et al. 2024

	Kiyacursor longipes n. gen., n. sp. Averianov et al. 2024	Averianov et al. 2024	1 specimen
	KOKM 5542 (holotype), partial skeleton
Reptilia

	? Sauropoda indet. Marsh 1878	Averianov et al. 2002
	now lost

	Titanosauriformes indet. Salgado et al. 1997	Averianov et al. 2002	1 individual
	partial pes
	= Sibirotitan astrosacralis n. gen., n. sp. Averianov et al. 2018	Averianov et al. 2018
	PM TGU 120/10-Sh1-22 -holotype (middle dorsal vertebra); PM TGU 16/0 - 81-88 (associated pedal elements)

	Brachiosauridae indet. Riggs 1904	Averianov et al. 2002	2 specimens
	"Pleurocoelus-like" teeth
	= Sibirotitan astrosacralis Averianov et al. 2018	Averianov et al. 2018
	LMCCE 105/4 and 108/4-115/4 (9 teeth)

	Titanosauridae indet. Lydekker 1885	Averianov et al. 2002	1 specimen
	peg-like tooth
	= Sibirotitan astrosacralis Averianov et al. 2018	Averianov et al. 2018
	LMCCE 105/4 and 108/4-115/4 (9 teeth)

	Stegosauria indet. Marsh 1877	Averianov et al. 2004

	Psittacosaurus sp. Osborn 1923	Rozhdestvensky 1960
	(7 measurements)
	= Psittacosaurus mongoliensis Osborn 1923	Rozhdestvensky 1973
	= Psittacosaurus sibiricus Voronkevich and Averianov 2000	Averianov et al. 2006
Reptilia - Hypsilophodontidae

	Hypsilophodontidae informal indet. 1	Averianov et al. 2004
	"sp. 1"

	Hypsilophodontidae informal indet. 2	Averianov et al. 2004
Reptilia

	Tagarosuchus kulemzini n. gen., n. sp. Alifanov et al. 1999	Averianov et al. 2002

	Choristodera indet. (Cope 1876)	Ivantsov et al. 2016	10 specimens
	fragmentary dentaries LMCCE 2-3/4, cervical vertebra LMCCE 4/4, dorsal vertebrae LMCCE 5-8/4, proximal rib fragments LMCCE 9-10/4, dorsal vertebra LMCCE 11/4
	= ? Neochoristodera indet. Evans and Hecht 1993	Skutschas and Vitenko 2017
	"Shestakovo choristodere"
Reptilia - Testudines

	Kirgizemys sp. Nessov and Khozatskiy 1973	Leshchinskiy et al. 2000
Reptilia - Xenosauridae

	Xenosauridae indet. Cope 1866	Podobina et al. 2002
	"gen. et spec. nov."
Reptilia

	Gekkota indet. Cuvier 1817	Skutschas 2006
	"gen. et sp. nov."
Reptilia - Paramacellodidae

	Paramacellodidae indet. Estes 1983	Podobina et al. 2002
	"gen. et sp. nov."
	= Paramacellodus sp. Hoffstetter 1967	Ivantsov et al. 2016
Reptilia

	Scincomorpha indet. Camp 1923	Podobina et al. 2002
Tritylodontidae

	Tritylodontidae indet. Cope 1884	Podobina et al. 2002

	Xenocretosuchus sibiricus n. gen., n. sp. Tatarinov and Matschenko 1999	Leshchinskiy et al. 2000
Mammalia

	Mammalia indet. Linnaeus 1758	Averianov et al. 2002
Mammalia - Eutriconodonta - Gobiconodontidae

	Gobiconodon sp. Trofimov 1978	Leshchinskiy et al. 2000
	"sp. nov"

	Gobiconodon borissiaki Trofimov 1978	Maschenko and Lopatin 1998	1 specimen
	PIN N 4463/1

	Gobiconodon hoburensis (Trofimov 1978)	Leshchinskiy et al. 2000
Mammalia - Symmetrodonta - Amphidontidae

	Acinacodus tagaricus n. gen., n. sp. Lopatin et al. 2010	Lopatin et al. 2010	1 specimen
	PIN, no. 4463/5, holotype partial right dentary
Mammalia - Amphilestidae

	Amphilestidae indet. Osborn 1888	Averianov et al. 2002
	= Amphilestinae indet. Osborn 1888	Lopatin and Averianov 2006
Mammalia - Tinodontidae

	? Tinodontidae indet. Marsh 1887	Gambaryan et al. 2004	1 specimen
	PM TGU 16/5-12, distal left humerus fragment

	Yermakia domitor n. gen., n. sp.	Lopatin et al. 2005	9 specimens
	PM TGU, no. 16/6-16, holotype right dentary fragment; PM TGU, no. 16/7-22, PM TGU, no. 16/4-8, PM TGU, no. 120/9-25, PM TGU, no. 16/6-18, PM TGU, no. 16/9-28, PM TGU, no. 16/6-6, PM TGU, no. 16/7-6; PM TGU 120/5-Sh1-1, left upper molar (possibly M2; reported by Averianov & Lopatin 2008)
Mammalia - Zhangheotheriidae

	Kiyatherium cardiodens Maschenko and Lopatin 2002	Maschenko and Lopatin 2002	1 specimen
	P5
Docodonta - Tegotheriidae

	Sibirotherium rossicus Maschenko et al. 2002	Lopatin and Averianov 2006
	= Sibirotherium rossicum n. gen., n. sp. Maschenko et al. 2002	Lopatin et al. 2009
	PM TGU 16/5-22, left dentary fragment
Palaeonisciformes

	Palaeonisciformes indet. Hay 1902	Leshchinskiy et al. 2000
Sinamiidae

	Sinamiidae indet. Berg 1940	Leshchinskiy et al. 2000
see common names

Geography

Country:	Russian Federation	State/province:	Kemerovo	County:	Chebulinsky
Coordinates:	55.9° North, 88.0° East (view map)
Paleocoordinates:	49.5° North, 97.4° East (Wright 2013)
Basis of coordinate:	stated in text
Altitude:	154 meters
Geographic resolution:	outcrop

Time

Period:	Cretaceous	Epoch:	Early Cretaceous
Stage:	Aptian	10 m.y. bin:	Cretaceous 3
Key time interval:	Aptian
Age range of interval:	121.4 - 113.2 m.y. ago

Stratigraphy

Formation:	Ilek
Stratigraphic resolution:	group of beds
Stratigraphy comments: "Ilekskaya Suite" Recently the fossiliferous level at Shestakovo 1 locality has been dated as Aptian by pollen (Bugdaeva et al., 2017).

Lithology and environment

Primary lithology:	concretionary,gray,green,yellow calcareous sandstone
Secondary lithology:	planar lamination,brown,gray,green,red siltstone
Includes fossils?	Y
Lithology description: "greenish-grey and yellowish-green sands, with calcareous sandstone concretions, interbedded rhythmically with packets of thin-laminated greenish-grey and brown-red siltstones and clay marls." Most bones from sandstone layer; mammal from siltstone just above
Environment:	terrestrial indet.

Taphonomy

Modes of preservation:	body
Size of fossils:	macrofossils,mesofossils,microfossils
Preservation of anatomical detail:	good
Articulated whole bodies:	some
Associated major elements:	some
Disassociated major elements:	many
Disassociated minor elements:	many
Fragmentation:	occasional

Collection methods and comments

Collection methods:	bulk,selective quarrying,surface (float),surface (in situ),mechanical,sieve,field collection,observed (not collected)
Reason for describing collection:	taxonomic analysis
Museum repositories:	PIN
Collectors:	A. Mossakovsky, I. Lebedev, E. Mashchenko	Collection dates:	1953–1960, 1994–1999, 2014–
Collection method comments: discovered 1953 by A. A. Mossakovsky & I. V. Lebedev; work resumed 1994

Metadata

Also known as:	LMCCE 004
Database number:	39680
Authorizer:	M. Carrano, E. Vlachos, P. Mannion, R. Benson	Enterer:	M. Carrano, P. Mannion, M. Oreska, R. Benson, J. Tennant, F. Aspromonte
Modifier:	M. Carrano	Research group:	vertebrate
Created:	2004-06-03 06:01:58	Last modified:	2025-02-22 15:12:02
Access level:	the public	Released:	2004-06-03 06:01:58
Creative Commons license:	CC0

Reference information

Primary reference:

68327.

K. B. Yuryev. 1954. Kratkiy obzor nakhodok dinozavrov na territorii SSSR [A brief reivew of dinosaur finds in the USSR]. Uchenyye Zapiski, Leningradskogo Ordena Lenina Gosudarstvennogo Universiteta. Seriya Biologicheskikh Nauk 181(38):183-197 [M. Carrano/M. Carrano]

Secondary references:

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65771		A. Averianov, S. Ivantsov, P. Skutschas, A. Faingertz, and S. Leshchinskiy. 2018. A new sauropod dinosaur from the Lower Cretaceous Ilek Formation, Western Siberia, Russia. Geobios 51:1-14 [P. Mannion/P. Mannion]
31142		A. O. Averianov. 2007. Mid-Cretaceous ornithocheirids (Pterosauria, Ornithocheiridae) from Russia and Uzbekistan. Paleontological Journal 41:79-86 [R. Butler/R. Butler/R. Butler]
71017		A. O. Averianov, S. V. Ivantsov, and P. P. Skutschas. 2019. Theropod teeth from the Lower Cretaceous Ilek Formation of Western Siberia, Russia. Proceedings of the Zoological Institute, Russian Academy of Sciences 323(2):65-84 [M. Carrano/M. Carrano]
14367	ETE	A. O. Averianov, S. Leshchinskiy, P. Skutschas, A. Fayngertz, and A. Rezvyi. 2004. Dinosaurs from the Early Cretaceous Ilek Formation in West Siberia, Russia. Second European Association of Vertebrate Palaeontologists Meeting, Moravian Museum, Brno. Abstracts of Papers 6 [M. Carrano/M. Carrano/M. Carrano]
39704		A. O. Averianov and A. Lopatin. 2008. “Protocone” in a pretribosphenic mammal and upper dentition Of tinodontid “symmetrodontans. Journal of Vertebrate Paleontology 28(2):548-552 [R. Benson/R. Benson]
86461		A. O. Averianov and A. V. Lopatin. 2023. Dinosaurs of Russia: a review of the localities. Herald of the Russian Academy of Sciences 93(4) [M. Carrano/M. Carrano/M. Carrano]
85489		A. O. Averianov and A. V. Lopatin. 2023. A long-necked theropod from the Lower Cretaceous of western Siberia, Russia. Journal of Vertebrate Paleontology e2216761:1-4 [M. Carrano/M. Carrano]
71292		A. O. Averianov, P. P. Skutschas, R. Schellhorn, A. V. Lopatin, P. N. Kolosov, V. V. Kolchanov, D. D. Vitenko, D. V. Grigoriev, and T. Martin. 2019. The northernmost sauropod record in the Northern Hemisphere. Lethaia 53:362-368 [P. Mannion/P. Mannion/M. Carrano]
88468		A. O. Averianov, P. P. Skutschas, A. A. Atuchin, D. A. Slobodin, O. A. Feofanova and O. N. Vladimirova. 2024. The last ceratosaur of Asia: a new noasaurid from the Early Cretaceous Great Siberian Refugium. Proceedings of the Royal Society B: Biological Sciences 291(2023):20240537:1-9 [E. Vlachos/F. Aspromonte/M. Carrano]
23626	ETE	A. O. Averianov and H.-D. Sues. 2007. A new troodontid (Dinosauria: Theropoda) from the Cenomanian of Uzbekistan, with a review of troodontid records from the territories of the former Soviet Union. Journal of Vertebrate Paleontology 27(1):87-98 [M. Carrano/M. Carrano]
14369	ETE	A. O. Averianov and A. V. Voronkevich. 2002. A new crown-group salamander from the Early Cretaceous of western Siberia. Russian Journal of Herpetology 9(3):209-214 [M. Carrano/M. Carrano/M. Carrano]
8702		A. O. Averianov, A. V. Voronkevich, E. N. Maschenko, S. V. Leshchinskiy, and A. V. Fayngertz. 2002. A sauropod foot from the Early Cretaceous of western Siberia, Russia. Acta Palaeontologica Polonica 47(1):117-124 [M. Carrano/M. Carrano/M. Carrano]
19092	ETE	A. O. Averianov, A. V. Voronkevich, S. V. Lechchinskiy and A. Fayngertz. 2006. A ceratopsian dinosaur Psittacosaurus sibiricus from the Early Cretaceous of west Siberia, Russia and its phylogenetic relationships. Journal of Systematic Palaeontology 4(4):359-395 [M. Carrano/M. Carrano]
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14468	ETE	M. B. Efimov and S. V. Leshchinskiy. 2000. [First discovery of a fossil crocodile skull in Siberia]. In A.V. Komarov (ed.), Materialy regionalanoj konferencii geologov Sibiri, Dalshchnego Vostoka i Severoâ ’Vostoka Rossii. Gala Press, Tomsk 2:361-363 [M. Carrano/M. Carrano/M. Carrano]
34817	ETE	P. P. Gambaryan, A. O. Averianov, E. N. Maschenko and S. V. Leshchinskiy. 2004. Mammalian humerus from the Early Cretaceous of West Siberia. Russian Journal of Theriology 3(2):51-58 [M. Carrano/M. Oreska]
86245		S. V. Ivantsov, A. V. Fayngerts, S.V. Leshchinskiy, A.S. Rezviy, P.P. Skutschas and A.O. Averianov. 2016. The Lower Cretaceous continental vertebrate fauna from the Shestakovo locality (West Siberia): Results of 20-year research. In O. S. Dzyuba, E. B. Pestchevitskaya, & B. N. Shurygin (eds.), Cretaceous Ecosystems and Their Responses to Paleoenvironmental Changes in Asia and the Western Pacific. Short papers for the Fourth International Symposium of International Geoscience Programme, IGCP Project 608 121-123 [M. Carrano/M. Carrano]
86246		A. E. Kostyunin. 2016. The history of discovery and research of the Shestakovo locality of Early Cretaceous vertebrates. In O. S. Dzyuba, E. B. Pestchevitskaya, & B. N. Shurygin (eds.), Cretaceous Ecosystems and Their Responses to Paleoenvironmental Changes in Asia and the Western Pacific. Short papers for the Fourth International Symposium of International Geoscience Programme, IGCP Project 608 124-126 [M. Carrano/M. Carrano]
38196		E. N. Kurochkin, N. V. Zelenkov, A. O. Averianov and S. V. Leshchinskiy. 2011. A new taxon of birds (Aves) from the Early Cretaceous of Western Siberia, Russia. Journal of Systematic Palaeontology 9(1):109-117 [P. Mannion/P. Mannion]
40059		S. V. Leshchinskiy, A. V. Faingerts, A. V. Voronkevich, E. N. Mashchensko, and A. O. Averianov. 2000. Predvaritel'nye rezul'taty izucheniy mestonakhozhdenii Shestakovskogo kompleksa rannemelovykh pozvonochnykh [Preliminary results of the investigation of the Shestakovo localities of Early Cretaceous vertebrates]. In A.V. Komarov (ed.), Materialy Regional'noy Konferentsii Geologov Sibiri Dal'nego Vostoka i Severo-Vostoka Rossii. Gala Press, Tomsk 2:363-366 [M. Carrano/M. Carrano/M. Carrano]
17882	ETE	A. V. Lopatin and A. O. Averianov. 2006. Mesozoic mammals of Russia. 9th International Symposium on Mesozoic Terrestrial Ecosystems and Biota, Abstracts and Proceedings Volume 67-70 [M. Carrano/M. Carrano/M. Carrano]
45970		A. V. Lopatin, A. O. Averianov, E. N. Maschenko and S. V. Leschinskiy. 2009. Early Cretaceous mammals of Western Siberia: 2. Tegotheriidae. Paleontological Journal 43(4):92-100 [P. Mannion/J. Tennant]
39702		A. V. Lopatin, E. N. Maschenko, A. O. Averianov, A. S. Rezvyi, P. P. Skutschas and S. V. Leschinskiy. 2005. Early Cretaceous Mammals from Western Siberia: 1. Tinodontidae. Paleontological Journal 39(5):523-534 [R. Benson/R. Benson]
39755		A. V. Lopatin, E. N. Maschenko, and A. O. Averianov. 2010. A new genus of triconodont mammals from the Early Cretaceous of western Siberia. Doklady biological Sciences 433:282-285 [R. Benson/R. Benson/R. Benson]
59918		A. V. Lopatin, E. N. Mashchenko, K. K. Tarasenko, A. V. Podlesnov, N. V. Demidenko and E. A. Kuzmina. 2015. A unique burial site of Early Cretaceous vertebrates in western Siberia (the Shestakovo 3 locality, Kemerovo Province, Russia). Doklady Biological Sciences 462:148-151 [M. Carrano/M. Carrano]
36988		E. N. Maschenko and A. V. Lopatin. 1998. First record of an Early Cretaceous triconodont mammal from Siberia. Bulletin de l'Institute Royal des Sciences Naturelles de Belgique, Sciences de la Terre 68:233-236 [M. Carrano/M. Carrano]
45980		E. N. Maschenko, A. V. Lopatin, and A. V. Voronkevich. 2002. A new genus of the tegotheriid docodonts (Docodonta, Tegotheriidae) from the Early Cretaceous of West Siberia. Russian Journal of Theriology 1(2):75-81 [P. Mannion/J. Tennant/J. Tennant]
39703		E. N. Maschenko and A. V. Lopatin. 2002. A new Early Cretaceous mammal from Western Siberia. Doklady Biological Sciences 386:475-477 [R. Benson/R. Benson/R. Benson]
16510	ETE	L. A. Nessov. 1995. Dinozavri severnoi Yevrazii: Novye dannye o sostave kompleksov, ekologii i paleobiogeografii [Dinosaurs of northern Eurasia: new data about assemblages, ecology, and paleobiogeography]. Institute for Scientific Research on the Earth's Crust, St. Petersburg State University, St. Petersburg 1-156 [M. Carrano/K. Maguire/M. Carrano]
14209	ETE	V. M. Podobina, G. M. Tatyanin, S. V. Leschchinskiy, A. V. Voronkevich, and A. V. Faingerts. 2002. A review of the Early Cretaceous fossil vertebrates of the Sehstakovo assemblage (south-east western Siberia, Russia). M. Wagreich (ed.), Aspects of Cretaceous Stratigraphy and Palaeobiogeography, Proceedings of the 6th International Cretaceous Symposium. Osterreichische Akademie der Wissenschaften Schriftenreihe der Erdwissenschaftlichen Kommissionen 15:239-247 [M. Carrano/M. Carrano/M. Carrano]
62664		A. K. Rozhdestvensky. 1960. Mestonakhozhdeniye nizhnemelovykh dinozavrov v Kuzbasse [The location of Lower Cretaceous dinosaurs in Kuzbas]. Paleontologicheskii Zhurnal 3(2):165 [M. Carrano/M. Carrano]
14374	ETE	A. K. Rozhdestvensky. 1973. The study of Cretaceous reptiles in Russia. Paleontological Journal 1973(2):90-99 [M. Carrano/M. Carrano/M. Carrano]
85555		A. K. Rozhdestvensky and L. I. Khozatsky. 1967. [Late Mesozoic terrestrial vertebrates from the Asiatic part of the USSR]. In G. G. Martinson (ed.), [Stratigraphy and Paleontology of the Mesozoic and Paleogene-Neogene continental deposits of the Asiatic part of the USSR] 82-92 [M. Carrano/M. Carrano]
17903	ETE	P. Skutschas. 2006. Mesozoic amphibians from Siberia, Russia. 9th International Symposium on Mesozoic Terrestrial Ecosystems and Biota, Abstracts and Proceedings Volume 123-126 [M. Carrano/M. Carrano]
54019		P. P. Skutschas. 2014. Kiyatriton leshchinskiyi Averianov et Voronkevich, 2001, a crown-group salamander from the Lower Cretaceous of Western Siberia, Russia. Cretaceous Research 51:88-94 [P. Mannion/J. Tennant/J. Tennant]
89217		P. P. Skutschas and D. D. Vitenko. 2017. Early Cretaceous choristoderes (Diapsida, Choristodera) from Siberia, Russia. Cretaceous Research 77(78):79–92 [M. Carrano/M. Carrano]
67961		T. A. Tumanova and V. R. Alifanov. 2018. First record of stegosaur (Ornithischia, Dinosauria) from the Aptian–Albian of Mongolia. Paleontological Journal 52(14):1771-1779 [P. Holroyd/P. Holroyd/M. Carrano]