Shestakovo-1: Aptian, Russian Federation
collected by A. Mossakovsky, I. Lebedev, E. Mashchenko 1953–1960, 1994–1999, 2014–

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Tritylodontidae
Tritylodontidae indet. Cope 1884
Podobina et al. 2002
Xenocretosuchus sibiricus n. gen., n. sp. Tatarinov and Matschenko 1999
Leshchinskiy et al. 2000
Mammalia
Mammalia indet. Linnaeus 1758
Averianov et al. 2002
Mammalia - Zhangheotheriidae
Kiyatherium cardiodens Maschenko and Lopatin 2002
Maschenko and Lopatin 2002 1 specimen
P5
Mammalia - Tinodontidae
? Tinodontidae indet. Marsh 1887
Gambaryan et al. 2004 1 specimen
PM TGU 16/5-12, distal left humerus fragment
Yermakia domitor n. gen., n. sp.
Lopatin et al. 2005 9 specimens
PM TGU, no. 16/6-16, holotype right dentary fragment; PM TGU, no. 16/7-22, PM TGU, no. 16/4-8, PM TGU, no. 120/9-25, PM TGU, no. 16/6-18, PM TGU, no. 16/9-28, PM TGU, no. 16/6-6, PM TGU, no. 16/7-6; PM TGU 120/5-Sh1-1, left upper molar (possibly M2; reported by Averianov & Lopatin 2008)
Mammalia - Amphilestidae
Amphilestidae indet. Osborn 1888
Averianov et al. 2002
    = Amphilestinae indet. Osborn 1888
Lopatin and Averianov 2006
spelled with current rank as Amphilestidae
Mammalia - Eutriconodonta - Gobiconodontidae
Gobiconodon sp. Trofimov 1978
Leshchinskiy et al. 2000
"sp. nov"
Gobiconodon hoburensis (Trofimov 1978)
Leshchinskiy et al. 2000
Gobiconodon borissiaki Trofimov 1978
Maschenko and Lopatin 1998 1 specimen
PIN N 4463/1
Mammalia - Symmetrodonta - Amphidontidae
Acinacodus tagaricus n. gen., n. sp. Lopatin et al. 2010
Lopatin et al. 2010 1 specimen
PIN, no. 4463/5, holotype partial right dentary
Docodonta - Tegotheriidae
Sibirotherium rossicus Maschenko et al. 2002
Lopatin and Averianov 2006
    = Sibirotherium rossicum n. gen., n. sp. Maschenko et al. 2002
Lopatin et al. 2009
original and current combination Sibirotherium rossicus
PM TGU 16/5-22, left dentary fragment
Reptilia
Tagarosuchus kulemzini n. gen., n. sp. Alifanov et al. 1999
Averianov et al. 2002
Pterodactyloidea indet. (Plieninger 1901)
Ivantsov et al. 2016
Reptilia - Ornithocheiridae
Ornithocheiridae indet. Seeley 1870
Skutschas 2006 3 elements
collection of PM TGU, three teeth
Reptilia
Stegosauria indet. Marsh 1877
Averianov et al. 2004
Reptilia - Hypsilophodontidae
Hypsilophodontidae informal indet. 1 Dollo 1882
Averianov et al. 2004
"sp. 1"
Hypsilophodontidae informal indet. 2 Dollo 1882
Averianov et al. 2004
Reptilia
Psittacosaurus sp. Osborn 1923
Rozhdestvensky 1960
(7 measurements)
    = Psittacosaurus mongoliensis Osborn 1923
Rozhdestvensky 1973
    = Psittacosaurus sibiricus Voronkevich and Averianov 2000
Averianov et al. 2006
? Sauropoda indet. Marsh 1878
Averianov et al. 2002
now lost
Titanosauriformes indet. Salgado et al. 1997
Averianov et al. 2002 1 individual
partial pes
    = Sibirotitan astrosacralis n. gen., n. sp. Averianov et al. 2018
Averianov et al. 2018
PM TGU 120/10-Sh1-22 -holotype (middle dorsal vertebra); PM TGU 16/0 - 81-88 (associated pedal elements)
Brachiosauridae indet. Riggs 1904
Averianov et al. 2002 2 specimens
"Pleurocoelus-like" teeth
    = Sibirotitan astrosacralis Averianov et al. 2018
Averianov et al. 2018
LMCCE 105/4 and 108/4-115/4 (9 teeth)
Titanosauridae indet. Lydekker 1885
Averianov et al. 2002 1 specimen
peg-like tooth
    = Sibirotitan astrosacralis Averianov et al. 2018
Averianov et al. 2018
LMCCE 105/4 and 108/4-115/4 (9 teeth)
cf. Prodeinodon sp. Osborn 1924
Averianov et al. 2004
nomen dubium belonging to Tetanurae
Coelurosauria informal indet. A (Huene 1914)
Averianov et al. 2019 31 specimens
LMCCE 004/5, 53, 86, 125 (Dromaeosauridae or Tyrannosauroidea)
Coelurosauria informal indet. B (Huene 1914)
Averianov et al. 2019 10 specimens
LMCCE 004/74, 83 (Dromaeosauridae or Tyrannosauroidea)
Coelurosauria informal indet. D (Huene 1914)
Averianov et al. 2019 7 specimens
LMCCE 004/13, 26, 51, 52, 116 (Dromaeosauridae or Tyrannosauroidea)
Coelurosauria indet. (Huene 1914)
Averianov and Lopatin 2023 1 specimen
PIN 329/16, incomplete anterior or middle cervical vertebra
Aves indet. Linnaeus 1758
Skutschas 2006
Reptilia - Mystiornithiformes - Mystiornithidae
Mystiornis cyrili n. gen., n. sp. Kurochkin et al. 2011
Kurochkin et al. 2011
PM TSU 16/5–45 - holotype (left metatarsus)
Reptilia
Deinonychosauria informal indet. C Colbert and Russell 1969
Averianov et al. 2019 14 specimens
LMCCE 004/32, 44, 61, 62 (Microraptorinae or Troodontidae)
Reptilia - Troodontidae
Troodontidae indet. Gilmore 1924
Averianov and Sues 2007 3 specimens
PM TGU 16/5-124
cf. Urbacodon sp. Averianov and Sues 2007
Averianov et al. 2019 5 specimens
LMCCE 004/1, 29, 78 (morphotype E)
Reptilia - Dromaeosauridae
Dromaeosauridae indet. Colbert and Russell 1969
Averianov et al. 2002
cf. Dromaeosauridae indet. Colbert and Russell 1969
Averianov et al. 2004
Reptilia
Therizinosauroidea indet. Russell and Dong 1994
Ivantsov et al. 2016
Choristodera indet. (Cope 1876)
Ivantsov et al. 2016
Reptilia - Paramacellodidae
Paramacellodidae indet. Estes 1983
Podobina et al. 2002
"gen. et sp. nov."
    = Paramacellodus sp. Hoffstetter 1967
Ivantsov et al. 2016
Reptilia - Xenosauridae
Xenosauridae indet. Cope 1866
Podobina et al. 2002
"gen. et spec. nov."
Reptilia
Scincomorpha indet. Camp 1923
Podobina et al. 2002
Gekkota indet. Cuvier 1817
Skutschas 2006
"gen. et sp. nov."
Reptilia - Testudines
Kirgizemys sp. Nessov and Khozatsky 1973
Leshchinskiy et al. 2000
Amphibia - Temnospondyli
Caudata indet. (Scopoli 1777)
Ivantsov et al. 2016
Amphibia - Temnospondyli - Urodela
Kiyatriton leshchinskiyi n. gen., n. sp. Averianov and Voronkevich 2002
Averianov and Voronkevich 2002 6 specimens
Sinamiidae
Sinamiidae indet. Berg 1940
Leshchinskiy et al. 2000
Palaeonisciformes
Palaeonisciformes indet. Hay 1902
Leshchinskiy et al. 2000
unclassified
Ginkgocycadophytus sp. Samoilovitch 1953
Bugdaeva et al. 2017
Pteridopsida
Osmundacidites wellmanii Couper 1953
Bugdaeva et al. 2017
Polypodiopsida - Gleicheniaceae
Gleicheniidites senonicus Ross 1949
Bugdaeva et al. 2017
unclassified
Pilosisporites setiferus
Bugdaeva et al. 2017
Pinopsida - Pinales - Pinaceae
Pinaceae indet. Rudolphi 1830
Bugdaeva et al. 2017
unclassified
Taurocusporites reduncus Stover 1962
Bugdaeva et al. 2017
see common names

Geography
Country:Russian Federation State/province:Kemerovo County:Chebulinsky
Coordinates: 55.9° North, 88.0° East (view map)
Paleocoordinates:54.2° North, 83.9° East
Basis of coordinate:stated in text
Altitude:154 meters
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Early/Lower Cretaceous
Stage:Aptian 10 m.y. bin:Cretaceous 3
Key time interval:Aptian
Age range of interval:121.40000 - 113.00000 m.y. ago
Stratigraphy
Formation:Ilek
Stratigraphic resolution:group of beds
Stratigraphy comments: "Ilekskaya Suite"

Recently the fossiliferous level at Shestakovo 1 locality has been dated as Aptian by pollen (Bugdaeva et al., 2017).
Lithology and environment
Primary lithology:concretionary,gray,green,yellow calcareous sandstone
Secondary lithology:planar lamination,brown,gray,green,red siltstone
Includes fossils?Y
Lithology description: "greenish-grey and yellowish-green sands, with calcareous sandstone concretions, interbedded rhythmically with packets of thin-laminated greenish-grey and brown-red siltstones and clay marls." Most bones from sandstone layer; mammal from siltstone just above
Environment:terrestrial indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils,microfossils
Preservation of anatomical detail:good
Articulated whole bodies:some
Associated major elements:some
Disassociated major elements:many
Disassociated minor elements:many
Fragmentation:occasional
Collection methods and comments
Collection methods:bulk,selective quarrying,surface (float),surface (in situ),mechanical,sieve,field collection,observed (not collected)
Reason for describing collection:taxonomic analysis
Museum repositories:PIN
Collectors:A. Mossakovsky, I. Lebedev, E. Mashchenko Collection dates:1953–1960, 1994–1999, 2014–
Collection method comments: discovered 1953 by A. A. Mossakovsky & I. V. Lebedev; work resumed 1994
Metadata
Also known as:LMCCE 004
Database number:39680
Authorizer:M. Carrano, R. Benson, P. Mannion Enterer:M. Carrano, J. Tennant, P. Mannion, M. Oreska, R. Benson
Modifier:M. Carrano Research group:vertebrate
Created:2004-06-03 06:01:58 Last modified:2023-11-15 10:17:10
Access level:the public Released:2004-06-03 06:01:58
Creative Commons license:CC BY
Reference information

Primary reference:
68327. K. B. Yuryev. 1954. Kratkiy obzor nakhodok dinozavrov na territorii SSSR [A brief reivew of dinosaur finds in the USSR]. Uchenyye Zapiski, Leningradskogo Ordena Lenina Gosudarstvennogo Universiteta. Seriya Biologicheskikh Nauk 181(38):183-197 [M. Carrano/M. Carrano]

Secondary references:

32161ETE V. R. Alifanov and Y. L. Bolotsky. 2010. Arkharavia heterocoelica gen. et sp. nov., a new sauropod dinosaur from the Upper Cretaceous of the Far East of Russia. Paleontological Journal 44(1):84-91 [M. Carrano/M. Carrano/M. Carrano]
65771 A. Averianov, S. Ivantsov, P. Skutschas, A. Faingertz, and S. Leshchinskiy. 2018. A new sauropod dinosaur from the Lower Cretaceous Ilek Formation, Western Siberia, Russia. Geobios 51:1-14 [P. Mannion/P. Mannion]
31142 A. O. Averianov. 2007. Mid-Cretaceous ornithocheirids (Pterosauria, Ornithocheiridae) from Russia and Uzbekistan. Paleontological Journal 41:79-86 [R. Butler/R. Butler/R. Butler]
71017 A. O. Averianov, S. V. Ivantsov, and P. P. Skutschas. 2019. Theropod teeth from the Lower Cretaceous Ilek Formation of Western Siberia, Russia. Proceedings of the Zoological Institute, Russian Academy of Sciences 323(2):65-84 [M. Carrano/M. Carrano]
14367ETE A. O. Averianov, S. Leshchinskiy, P. Skutschas, A. Fayngertz, and A. Rezvyi. 2004. Dinosaurs from the Early Cretaceous Ilek Formation in West Siberia, Russia. Second European Association of Vertebrate Palaeontologists Meeting, Moravian Museum, Brno. Abstracts of Papers 6 [M. Carrano/M. Carrano/M. Carrano]
39704 A. O. Averianov and A. Lopatin. 2008. “Protocone” in a pretribosphenic mammal and upper dentition Of tinodontid “symmetrodontans. Journal of Vertebrate Paleontology 28(2):548-552 [R. Benson/R. Benson]
86461 A. O. Averianov and A. V. Lopatin. 2023. Dinosaurs of Russia: a review of the localities. Herald of the Russian Academy of Sciences 93(4) [M. Carrano/M. Carrano/M. Carrano]
85489 A. O. Averianov and A. V. Lopatin. 2023. A long-necked theropod from the Lower Cretaceous of western Siberia, Russia. Journal of Vertebrate Paleontology e2216761:1-4 [M. Carrano/M. Carrano]
71292 A. O. Averianov, P. P. Skutschas, R. Schellhorn, A. V. Lopatin, P. N. Kolosov, V. V. Kolchanov, D. D. Vitenko, D. V. Grigoriev, and T. Martin. 2019. The northernmost sauropod record in the Northern Hemisphere. Lethaia 53:362-368 [P. Mannion/P. Mannion/M. Carrano]
23626ETE A. O. Averianov and H.-D. Sues. 2007. A new troodontid (Dinosauria: Theropoda) from the Cenomanian of Uzbekistan, with a review of troodontid records from the territories of the former Soviet Union. Journal of Vertebrate Paleontology 27(1):87-98 [M. Carrano/M. Carrano]
14369ETE A. O. Averianov and A. V. Voronkevich. 2002. A new crown-group salamander from the Early Cretaceous of western Siberia. Russian Journal of Herpetology 9(3):209-214 [M. Carrano/M. Carrano/M. Carrano]
8702 A. O. Averianov, A. V. Voronkevich, E. N. Maschenko, S. V. Leshchinskiy, and A. V. Fayngertz. 2002. A sauropod foot from the Early Cretaceous of western Siberia, Russia. Acta Palaeontologica Polonica 47(1):117-124 [M. Carrano/M. Carrano/M. Carrano]
19092ETE A. O. Averianov, A. V. Voronkevich, S. V. Lechchinskiy and A. Fayngertz. 2006. A ceratopsian dinosaur Psittacosaurus sibiricus from the Early Cretaceous of west Siberia, Russia and its phylogenetic relationships. Journal of Systematic Palaeontology 4(4):359-395 [M. Carrano/M. Carrano]
86463 E. V. Bugdaeva, V. S. Markevich, and E. B. Volynets. 2017. Palaeoenvironmental and palaeoclimatic reconstruction of the Early Cretaceous psittacosaur localities, Asia. In D. Cheong, Y.-I. Lee, & D. Kim (eds.), Cretaceous Ecosystems and Their Responses to Paleoenvironmental Changes in Asia and the Western Pacific. Proceedings and Field Guidebook for the Fifth International Symposium of International Geoscience Programme IGCP Project 608 31-34 [M. Carrano/M. Carrano]
64010 Z.-W. Cheng. 1982. Páxíng lèi {Reptilia}. Mesozoic Stratigraphic Paleontology in the Guyang Coal-Bearing Basin, Inner Mongolia 123-136 [M. Carrano/M. Carrano]
86243 N. V. Demidenko and E. N. Maschenko. 2016. Results of paleontological excavations 2014–2015 held by the Kemerovo Regional Museum at the Early Cretaceous Shestakovo vertebrate localities. In O. S. Dzyuba, E. B. Pestchevitskaya, & B. N. Shurygin (eds.), Cretaceous Ecosystems and Their Responses to Paleoenvironmental Changes in Asia and the Western Pacific. Short papers for the Fourth International Symposium of International Geoscience Programme, IGCP Project 608 114-116 [M. Carrano/M. Carrano]
14468ETE M. B. Efimov and S. V. Leshchinskiy. 2000. [First discovery of a fossil crocodile skull in Siberia]. In A.V. Komarov (ed.), Materialy regionalanoj konferencii geologov Sibiri, Dalshchnego Vostoka i Severoâ ’Vostoka Rossii. Gala Press, Tomsk 2:361-363 [M. Carrano/M. Carrano/M. Carrano]
34817ETE P. P. Gambaryan, A. O. Averianov, E. N. Maschenko and S. V. Leshchinskiy. 2004. Mammalian humerus from the Early Cretaceous of West Siberia. Russian Journal of Theriology 3(2):51-58 [M. Carrano/M. Oreska]
86245 S. V. Ivantsov, A. V. Fayngerts, S.V. Leshchinskiy, A.S. Rezviy, P.P. Skutschas and A.O. Averianov. 2016. The Lower Cretaceous continental vertebrate fauna from the Shestakovo locality (West Siberia): Results of 20-year research. In O. S. Dzyuba, E. B. Pestchevitskaya, & B. N. Shurygin (eds.), Cretaceous Ecosystems and Their Responses to Paleoenvironmental Changes in Asia and the Western Pacific. Short papers for the Fourth International Symposium of International Geoscience Programme, IGCP Project 608 121-123 [M. Carrano/M. Carrano]
86246 A. E. Kostyunin. 2016. The history of discovery and research of the Shestakovo locality of Early Cretaceous vertebrates. In O. S. Dzyuba, E. B. Pestchevitskaya, & B. N. Shurygin (eds.), Cretaceous Ecosystems and Their Responses to Paleoenvironmental Changes in Asia and the Western Pacific. Short papers for the Fourth International Symposium of International Geoscience Programme, IGCP Project 608 124-126 [M. Carrano/M. Carrano]
38196 E. N. Kurochkin, N. V. Zelenkov, A. O. Averianov and S. V. Leshchinskiy. 2011. A new taxon of birds (Aves) from the Early Cretaceous of Western Siberia, Russia. Journal of Systematic Palaeontology 9(1):109-117 [P. Mannion/P. Mannion]
40059 S. V. Leshchinskiy, A. V. Faingerts, A. V. Voronkevich, E. N. Mashchensko, and A. O. Averianov. 2000. Predvaritel'nye rezul'taty izucheniy mestonakhozhdenii Shestakovskogo kompleksa rannemelovykh pozvonochnykh [Preliminary results of the investigation of the Shestakovo localities of Early Cretaceous vertebrates]. In A.V. Komarov (ed.), Materialy Regional'noy Konferentsii Geologov Sibiri Dal'nego Vostoka i Severo-Vostoka Rossii. Gala Press, Tomsk 2:363-366 [M. Carrano/M. Carrano/M. Carrano]
17882ETE A. V. Lopatin and A. O. Averianov. 2006. Mesozoic mammals of Russia. 9th International Symposium on Mesozoic Terrestrial Ecosystems and Biota, Abstracts and Proceedings Volume 67-70 [M. Carrano/M. Carrano/M. Carrano]
45970 A. V. Lopatin, A. O. Averianov, E. N. Maschenko and S. V. Leschinskiy. 2009. Early Cretaceous mammals of Western Siberia: 2. Tegotheriidae. Paleontological Journal 43(4):92-100 [P. Mannion/J. Tennant]
39702 A. V. Lopatin, E. N. Maschenko, A. O. Averianov, A. S. Rezvyi, P. P. Skutschas and S. V. Leschinskiy. 2005. Early Cretaceous Mammals from Western Siberia: 1. Tinodontidae. Paleontological Journal 39(5):523-534 [R. Benson/R. Benson]
39755 A. V. Lopatin, E. N. Maschenko, and A. O. Averianov. 2010. A new genus of triconodont mammals from the Early Cretaceous of western Siberia. Doklady biological Sciences 433:282-285 [R. Benson/R. Benson/R. Benson]
59918 A. V. Lopatin, E. N. Mashchenko, K. K. Tarasenko, A. V. Podlesnov, N. V. Demidenko and E. A. Kuzmina. 2015. A unique burial site of Early Cretaceous vertebrates in western Siberia (the Shestakovo 3 locality, Kemerovo Province, Russia). Doklady Biological Sciences 462:148-151 [M. Carrano/M. Carrano]
36988 E. N. Maschenko and A. V. Lopatin. 1998. First record of an Early Cretaceous triconodont mammal from Siberia. Bulletin de l'Institute Royal des Sciences Naturelles de Belgique, Sciences de la Terre 68:233-236 [M. Carrano/M. Carrano]
45980 E. N. Maschenko, A. V. Lopatin, and A. V. Voronkevich. 2002. A new genus of the tegotheriid docodonts (Docodonta, Tegotheriidae) from the Early Cretaceous of West Siberia. Russian Journal of Theriology 1(2):75-81 [P. Mannion/J. Tennant/J. Tennant]
39703 E. N. Maschenko and A. V. Lopatin. 2002. A new Early Cretaceous mammal from Western Siberia. Doklady Biological Sciences 386:475-477 [R. Benson/R. Benson/R. Benson]
16510ETE L. A. Nessov. 1995. Dinozavri severnoi Yevrazii: Novye dannye o sostave kompleksov, ekologii i paleobiogeografii [Dinosaurs of northern Eurasia: new data about assemblages, ecology, and paleobiogeography]. Institute for Scientific Research on the Earth's Crust, St. Petersburg State University, St. Petersburg 1-156 [M. Carrano/K. Maguire/M. Carrano]
14209ETE V. M. Podobina, G. M. Tatyanin, S. V. Leschchinskiy, A. V. Voronkevich, and A. V. Faingerts. 2002. A review of the Early Cretaceous fossil vertebrates of the Sehstakovo assemblage (south-east western Siberia, Russia). M. Wagreich (ed.), Aspects of Cretaceous Stratigraphy and Palaeobiogeography, Proceedings of the 6th International Cretaceous Symposium. Osterreichische Akademie der Wissenschaften Schriftenreihe der Erdwissenschaftlichen Kommissionen 15:239-247 [M. Carrano/M. Carrano/M. Carrano]
62664 A. K. Rozhdestvensky. 1960. Mestonakhozhdeniye nizhnemelovykh dinozavrov v Kuzbasse [The location of Lower Cretaceous dinosaurs in Kuzbas]. Paleontologicheskii Zhurnal 3(2):165 [M. Carrano/M. Carrano]
14374ETE A. K. Rozhdestvensky. 1973. The study of Cretaceous reptiles in Russia. Paleontological Journal 1973(2):90-99 [M. Carrano/M. Carrano/M. Carrano]
85555 A. K. Rozhdestvensky and L. I. Khozatsky. 1967. [Late Mesozoic terrestrial vertebrates from the Asiatic part of the USSR]. In G. G. Martinson (ed.), [Stratigraphy and Paleontology of the Mesozoic and Paleogene-Neogene continental deposits of the Asiatic part of the USSR] 82-92 [M. Carrano/M. Carrano]
17903ETE P. Skutschas. 2006. Mesozoic amphibians from Siberia, Russia. 9th International Symposium on Mesozoic Terrestrial Ecosystems and Biota, Abstracts and Proceedings Volume 123-126 [M. Carrano/M. Carrano]
54019 P. P. Skutschas. 2014. Kiyatriton leshchinskiyi Averianov et Voronkevich, 2001, a crown-group salamander from the Lower Cretaceous of Western Siberia, Russia. Cretaceous Research 51:88-94 [P. Mannion/J. Tennant/J. Tennant]
67961 T. A. Tumanova and V. R. Alifanov. 2018. First record of stegosaur (Ornithischia, Dinosauria) from the Aptian–Albian of Mongolia. Paleontological Journal 52(14):1771-1779 [P. Holroyd/P. Holroyd/M. Carrano]