Shestakovo-1: Aptian, Russian Federation
collected by A. Mossakovsky, I. Lebedev, E. Mashchenko 1953–1960, 1994–1999, 2014–
List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia
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Choristodera indet.
(Cope 1876)
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Ivantsov et al. 2016 | 10 specimens | ||||||||
fragmentary dentaries LMCCE 2-3/4, cervical vertebra LMCCE 4/4, dorsal vertebrae LMCCE 5-8/4, proximal rib fragments LMCCE 9-10/4, dorsal vertebra LMCCE 11/4 | ||||||||||
= ? Neochoristodera indet.
Evans and Hecht 1993
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Skutschas and Vitenko 2017 | |||||||||
"Shestakovo choristodere" | ||||||||||
Tagarosuchus kulemzini n. gen., n. sp.
Alifanov et al. 1999
|
Averianov et al. 2002 | |||||||||
Reptilia
- Pterosauria
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Pterodactyloidea indet.
(Plieninger 1901)
|
Ivantsov et al. 2016 | |||||||||
Reptilia
- Pterosauria
- Ornithocheiridae
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Ornithocheiridae indet.
Seeley 1870
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Skutschas 2006 | 3 elements | ||||||||
collection of PM TGU, three teeth | ||||||||||
Reptilia
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Stegosauria indet.
Marsh 1877
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Averianov et al. 2004 | |||||||||
Reptilia
- Hypsilophodontidae
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Hypsilophodontidae informal indet. 1
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Averianov et al. 2004 | |||||||||
"sp. 1" | ||||||||||
Hypsilophodontidae informal indet. 2
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Averianov et al. 2004 | |||||||||
Reptilia
|
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Psittacosaurus sp.
Osborn 1923
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Rozhdestvensky 1960 | |||||||||
(7 measurements) | ||||||||||
= Psittacosaurus mongoliensis
Osborn 1923
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Rozhdestvensky 1973 | |||||||||
= Psittacosaurus sibiricus
Voronkevich and Averianov 2000
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Averianov et al. 2006 | |||||||||
? Sauropoda indet.
Marsh 1878
|
Averianov et al. 2002 | |||||||||
now lost | ||||||||||
Titanosauriformes indet.
Salgado et al. 1997
|
Averianov et al. 2002 | 1 individual | ||||||||
partial pes | ||||||||||
= Sibirotitan astrosacralis n. gen., n. sp.
Averianov et al. 2018
|
Averianov et al. 2018 | |||||||||
PM TGU 120/10-Sh1-22 -holotype (middle dorsal vertebra); PM TGU 16/0 - 81-88 (associated pedal elements) | ||||||||||
Brachiosauridae indet.
Riggs 1904
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Averianov et al. 2002 | 2 specimens | ||||||||
"Pleurocoelus-like" teeth | ||||||||||
= Sibirotitan astrosacralis
Averianov et al. 2018
|
Averianov et al. 2018 | |||||||||
LMCCE 105/4 and 108/4-115/4 (9 teeth) | ||||||||||
Titanosauridae indet.
Lydekker 1885
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Averianov et al. 2002 | 1 specimen | ||||||||
peg-like tooth | ||||||||||
= Sibirotitan astrosacralis
Averianov et al. 2018
|
Averianov et al. 2018 | |||||||||
LMCCE 105/4 and 108/4-115/4 (9 teeth) | ||||||||||
Coelurosauria informal indet. A
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Averianov et al. 2019 | 31 specimens | ||||||||
LMCCE 004/5, 53, 86, 125 (Dromaeosauridae or Tyrannosauroidea) | ||||||||||
Coelurosauria informal indet. B
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Averianov et al. 2019 | 10 specimens | ||||||||
LMCCE 004/74, 83 (Dromaeosauridae or Tyrannosauroidea) | ||||||||||
Coelurosauria informal indet. D
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Averianov et al. 2019 | 7 specimens | ||||||||
LMCCE 004/13, 26, 51, 52, 116 (Dromaeosauridae or Tyrannosauroidea) | ||||||||||
Therizinosauroidea indet.
Russell and Dong 1994
|
Ivantsov et al. 2016 | |||||||||
Aves indet.
Linnaeus 1758
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Skutschas 2006 | |||||||||
Reptilia
- Mystiornithiformes
- Mystiornithidae
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Mystiornis cyrili n. gen., n. sp.
Kurochkin et al. 2011
|
Kurochkin et al. 2011 | 1 specimen | ||||||||
PM TSU 16/5–45 - holotype (left metatarsus) | ||||||||||
Reptilia
|
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Deinonychosauria informal indet. C
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Averianov et al. 2019 | 14 specimens | ||||||||
LMCCE 004/32, 44, 61, 62 (Microraptorinae or Troodontidae) | ||||||||||
Reptilia
- Troodontidae
|
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Troodontidae indet.
Gilmore 1924
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Averianov and Sues 2007 | 3 specimens | ||||||||
PM TGU 16/5-124 | ||||||||||
cf. Urbacodon sp.
Averianov and Sues 2007
|
Averianov et al. 2019 | 5 specimens | ||||||||
LMCCE 004/1, 29, 78 (morphotype E) | ||||||||||
Reptilia
- Dromaeosauridae
|
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Dromaeosauridae indet.
Colbert and Russell 1969
|
Averianov et al. 2002 | |||||||||
cf. Dromaeosauridae indet.
Colbert and Russell 1969
|
Averianov et al. 2004 | |||||||||
Reptilia
|
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cf. Prodeinodon sp.
Osborn 1924
|
Averianov et al. 2004 | |||||||||
Reptilia
- Noasauridae
|
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Coelurosauria indet.
(Huene 1914)
|
Averianov and Lopatin 2023 | 1 specimen | ||||||||
PIN 329/16, incomplete anterior or middle cervical vertebra | ||||||||||
= Kiyacursor longipes
Averianov et al. 2024
|
Averianov et al. 2024 | |||||||||
Kiyacursor longipes n. gen., n. sp.
Averianov et al. 2024
|
Averianov et al. 2024 | 1 specimen | ||||||||
KOKM 5542 (holotype), partial skeleton | ||||||||||
Reptilia
- Testudines
|
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Kirgizemys sp.
Nessov and Khozatskiy 1973
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Leshchinskiy et al. 2000 | |||||||||
Reptilia
|
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Gekkota indet.
Cuvier 1817
|
Skutschas 2006 | |||||||||
"gen. et sp. nov." | ||||||||||
Reptilia
- Paramacellodidae
|
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Paramacellodidae indet.
Estes 1983
|
Podobina et al. 2002 | |||||||||
"gen. et sp. nov." | ||||||||||
= Paramacellodus sp.
Hoffstetter 1967
|
Ivantsov et al. 2016 | |||||||||
Reptilia
- Xenosauridae
|
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Xenosauridae indet.
Cope 1866
|
Podobina et al. 2002 | |||||||||
"gen. et spec. nov." | ||||||||||
Reptilia
|
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Scincomorpha indet.
Camp 1923
|
Podobina et al. 2002 | |||||||||
Docodonta
- Tegotheriidae
|
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Sibirotherium rossicus
Maschenko et al. 2002
|
Lopatin and Averianov 2006 | |||||||||
= Sibirotherium rossicum n. gen., n. sp.
Maschenko et al. 2002
|
Lopatin et al. 2009 | |||||||||
PM TGU 16/5-22, left dentary fragment | ||||||||||
Mammalia
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Mammalia indet.
Linnaeus 1758
|
Averianov et al. 2002 | |||||||||
Mammalia
- Eutriconodonta
- Gobiconodontidae
|
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Gobiconodon sp.
Trofimov 1978
|
Leshchinskiy et al. 2000 | |||||||||
"sp. nov" | ||||||||||
Gobiconodon hoburensis
(Trofimov 1978)
|
Leshchinskiy et al. 2000 | |||||||||
Gobiconodon borissiaki
Trofimov 1978
|
Maschenko and Lopatin 1998 | 1 specimen | ||||||||
PIN N 4463/1 | ||||||||||
Mammalia
- Tinodontidae
|
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? Tinodontidae indet.
Marsh 1887
|
Gambaryan et al. 2004 | 1 specimen | ||||||||
PM TGU 16/5-12, distal left humerus fragment | ||||||||||
Yermakia domitor n. gen., n. sp.
|
Lopatin et al. 2005 | 9 specimens | ||||||||
PM TGU, no. 16/6-16, holotype right dentary fragment; PM TGU, no. 16/7-22, PM TGU, no. 16/4-8, PM TGU, no. 120/9-25, PM TGU, no. 16/6-18, PM TGU, no. 16/9-28, PM TGU, no. 16/6-6, PM TGU, no. 16/7-6; PM TGU 120/5-Sh1-1, left upper molar (possibly M2; reported by Averianov & Lopatin 2008) | ||||||||||
Mammalia
- Zhangheotheriidae
|
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Kiyatherium cardiodens
Maschenko and Lopatin 2002
|
Maschenko and Lopatin 2002 | 1 specimen | ||||||||
P5 | ||||||||||
Mammalia
- Amphilestidae
|
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Amphilestidae indet.
Osborn 1888
|
Averianov et al. 2002 | |||||||||
= Amphilestinae indet.
Osborn 1888
|
Lopatin and Averianov 2006 | |||||||||
Mammalia
- Symmetrodonta
- Amphidontidae
|
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Acinacodus tagaricus n. gen., n. sp.
Lopatin et al. 2010
|
Lopatin et al. 2010 | 1 specimen | ||||||||
PIN, no. 4463/5, holotype partial right dentary | ||||||||||
Tritylodontidae
|
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Tritylodontidae indet.
Cope 1884
|
Podobina et al. 2002 | |||||||||
Xenocretosuchus sibiricus n. gen., n. sp.
Tatarinov and Matschenko 1999
|
Leshchinskiy et al. 2000 | |||||||||
Amphibia
- Temnospondyli
|
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Caudata indet.
(Scopoli 1777)
|
Ivantsov et al. 2016 | |||||||||
Amphibia
- Temnospondyli
- Urodela
|
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Kiyatriton leshchinskiyi n. gen., n. sp.
Averianov and Voronkevich 2002
|
Averianov and Voronkevich 2002 | 6 specimens | ||||||||
Sinamiidae
|
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Sinamiidae indet.
Berg 1940
|
Leshchinskiy et al. 2000 | |||||||||
Palaeonisciformes
|
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Palaeonisciformes indet.
Hay 1902
|
Leshchinskiy et al. 2000 | |||||||||
Pteridopsida
|
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Osmundacidites wellmanii
Couper 1953
|
Bugdaeva et al. 2017 | |||||||||
Polypodiopsida
- Gleicheniaceae
|
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Gleicheniidites senonicus
Ross 1949
|
Bugdaeva et al. 2017 | |||||||||
unclassified
|
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Pilosisporites setiferus
|
Bugdaeva et al. 2017 | |||||||||
Ginkgocycadophytus sp.
Samoilovitch 1953
|
Bugdaeva et al. 2017 | |||||||||
Pinopsida
- Pinales
- Pinaceae
|
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Pinaceae indet.
Rudolphi 1830
|
Bugdaeva et al. 2017 | |||||||||
unclassified
|
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Taurocusporites reduncus
Stover 1962
|
Bugdaeva et al. 2017 | |||||||||
see common names |
Geography
Country: | Russian Federation | State/province: | Kemerovo | County: | Chebulinsky |
Coordinates: | 55.9° North, 88.0° East (view map) | ||||
Paleocoordinates: | 54.2° North, 83.9° East | ||||
Basis of coordinate: | stated in text | ||||
Altitude: | 154 meters | ||||
Geographic resolution: | outcrop |
Time
Period: | Cretaceous | Epoch: | Early Cretaceous |
Stage: | Aptian | 10 m.y. bin: | Cretaceous 3 |
Key time interval: | Aptian | ||
Age range of interval: | 121.4 - 113 m.y. ago |
Stratigraphy
Formation: | Ilek | ||||
Stratigraphic resolution: | group of beds | ||||
Stratigraphy comments: "Ilekskaya Suite"
Recently the fossiliferous level at Shestakovo 1 locality has been dated as Aptian by pollen (Bugdaeva et al., 2017). |
Lithology and environment
Primary lithology: | concretionary,gray,green,yellow calcareous sandstone |
Secondary lithology: | planar lamination,brown,gray,green,red siltstone |
Includes fossils? | Y |
Lithology description: "greenish-grey and yellowish-green sands, with calcareous sandstone concretions, interbedded rhythmically with packets of thin-laminated greenish-grey and brown-red siltstones and clay marls." Most bones from sandstone layer; mammal from siltstone just above | |
Environment: | terrestrial indet. |
Taphonomy
Modes of preservation: | body |
Size of fossils: | macrofossils,mesofossils,microfossils |
Preservation of anatomical detail: | good |
Articulated whole bodies: | some |
Associated major elements: | some |
Disassociated major elements: | many |
Disassociated minor elements: | many |
Fragmentation: | occasional |
Collection methods and comments
Collection methods: | bulk,selective quarrying,surface (float),surface (in situ),mechanical,sieve,field collection,observed (not collected) | ||
Reason for describing collection: | taxonomic analysis | ||
Museum repositories: | PIN | ||
Collectors: | A. Mossakovsky, I. Lebedev, E. Mashchenko | Collection dates: | 1953–1960, 1994–1999, 2014– |
Collection method comments: discovered 1953 by A. A. Mossakovsky & I. V. Lebedev; work resumed 1994 |
Metadata
Also known as: | LMCCE 004 | ||
Database number: | 39680 | ||
Authorizer: | M. Carrano, R. Benson, P. Mannion, E. Vlachos | Enterer: | M. Carrano, R. Benson, M. Oreska, F. Aspromonte, P. Mannion, J. Tennant |
Modifier: | M. Carrano | Research group: | vertebrate |
Created: | 2004-06-03 06:01:58 | Last modified: | 2023-11-15 10:17:10 |
Access level: | the public | Released: | 2004-06-03 06:01:58 |
Creative Commons license: | CC BY |
Reference information
Primary reference:
68327. | K. B. Yuryev. 1954. Kratkiy obzor nakhodok dinozavrov na territorii SSSR [A brief reivew of dinosaur finds in the USSR]. Uchenyye Zapiski, Leningradskogo Ordena Lenina Gosudarstvennogo Universiteta. Seriya Biologicheskikh Nauk 181(38):183-197 [M. Carrano/M. Carrano] |
Secondary references:
32161 | ETE | V. R. Alifanov and Y. L. Bolotsky. 2010. Arkharavia heterocoelica gen. et sp. nov., a new sauropod dinosaur from the Upper Cretaceous of the Far East of Russia. Paleontological Journal 44(1):84-91 [M. Carrano/M. Carrano/M. Carrano] |
65771 | A. Averianov, S. Ivantsov, P. Skutschas, A. Faingertz, and S. Leshchinskiy. 2018. A new sauropod dinosaur from the Lower Cretaceous Ilek Formation, Western Siberia, Russia. Geobios 51:1-14 [P. Mannion/P. Mannion] | |
31142 | A. O. Averianov. 2007. Mid-Cretaceous ornithocheirids (Pterosauria, Ornithocheiridae) from Russia and Uzbekistan. Paleontological Journal 41:79-86 [R. Butler/R. Butler/R. Butler] | |
71017 | A. O. Averianov, S. V. Ivantsov, and P. P. Skutschas. 2019. Theropod teeth from the Lower Cretaceous Ilek Formation of Western Siberia, Russia. Proceedings of the Zoological Institute, Russian Academy of Sciences 323(2):65-84 [M. Carrano/M. Carrano] | |
14367 | ETE | A. O. Averianov, S. Leshchinskiy, P. Skutschas, A. Fayngertz, and A. Rezvyi. 2004. Dinosaurs from the Early Cretaceous Ilek Formation in West Siberia, Russia. Second European Association of Vertebrate Palaeontologists Meeting, Moravian Museum, Brno. Abstracts of Papers 6 [M. Carrano/M. Carrano/M. Carrano] |
39704 | A. O. Averianov and A. Lopatin. 2008. “Protocone” in a pretribosphenic mammal and upper dentition Of tinodontid “symmetrodontans. Journal of Vertebrate Paleontology 28(2):548-552 [R. Benson/R. Benson] | |
86461 | A. O. Averianov and A. V. Lopatin. 2023. Dinosaurs of Russia: a review of the localities. Herald of the Russian Academy of Sciences 93(4) [M. Carrano/M. Carrano/M. Carrano] | |
85489 | A. O. Averianov and A. V. Lopatin. 2023. A long-necked theropod from the Lower Cretaceous of western Siberia, Russia. Journal of Vertebrate Paleontology e2216761:1-4 [M. Carrano/M. Carrano] | |
71292 | A. O. Averianov, P. P. Skutschas, R. Schellhorn, A. V. Lopatin, P. N. Kolosov, V. V. Kolchanov, D. D. Vitenko, D. V. Grigoriev, and T. Martin. 2019. The northernmost sauropod record in the Northern Hemisphere. Lethaia 53:362-368 [P. Mannion/P. Mannion/M. Carrano] | |
88468 | A. O. Averianov, P. P. Skutschas, A. A. Atuchin, D. A. Slobodin, O. A. Feofanova and O. N. Vladimirova. 2024. The last ceratosaur of Asia: a new noasaurid from the Early Cretaceous Great Siberian Refugium. Proceedings of the Royal Society B: Biological Sciences 291(2023):20240537:1-9 [E. Vlachos/F. Aspromonte/M. Carrano] | |
23626 | ETE | A. O. Averianov and H.-D. Sues. 2007. A new troodontid (Dinosauria: Theropoda) from the Cenomanian of Uzbekistan, with a review of troodontid records from the territories of the former Soviet Union. Journal of Vertebrate Paleontology 27(1):87-98 [M. Carrano/M. Carrano] |
8702 | A. O. Averianov, A. V. Voronkevich, E. N. Maschenko, S. V. Leshchinskiy, and A. V. Fayngertz. 2002. A sauropod foot from the Early Cretaceous of western Siberia, Russia. Acta Palaeontologica Polonica 47(1):117-124 [M. Carrano/M. Carrano/M. Carrano] | |
14369 | ETE | A. O. Averianov and A. V. Voronkevich. 2002. A new crown-group salamander from the Early Cretaceous of western Siberia. Russian Journal of Herpetology 9(3):209-214 [M. Carrano/M. Carrano/M. Carrano] |
19092 | ETE | A. O. Averianov, A. V. Voronkevich, S. V. Lechchinskiy and A. Fayngertz. 2006. A ceratopsian dinosaur Psittacosaurus sibiricus from the Early Cretaceous of west Siberia, Russia and its phylogenetic relationships. Journal of Systematic Palaeontology 4(4):359-395 [M. Carrano/M. Carrano] |
86463 | E. V. Bugdaeva, V. S. Markevich, and E. B. Volynets. 2017. Palaeoenvironmental and palaeoclimatic reconstruction of the Early Cretaceous psittacosaur localities, Asia. In D. Cheong, Y.-I. Lee, & D. Kim (eds.), Cretaceous Ecosystems and Their Responses to Paleoenvironmental Changes in Asia and the Western Pacific. Proceedings and Field Guidebook for the Fifth International Symposium of International Geoscience Programme IGCP Project 608 31-34 [M. Carrano/M. Carrano] | |
64010 | Z.-W. Cheng. 1982. Páxíng lèi {Reptilia}. Mesozoic Stratigraphic Paleontology in the Guyang Coal-Bearing Basin, Inner Mongolia 123-136 [M. Carrano/M. Carrano] | |
86243 | N. V. Demidenko and E. N. Maschenko. 2016. Results of paleontological excavations 2014–2015 held by the Kemerovo Regional Museum at the Early Cretaceous Shestakovo vertebrate localities. In O. S. Dzyuba, E. B. Pestchevitskaya, & B. N. Shurygin (eds.), Cretaceous Ecosystems and Their Responses to Paleoenvironmental Changes in Asia and the Western Pacific. Short papers for the Fourth International Symposium of International Geoscience Programme, IGCP Project 608 114-116 [M. Carrano/M. Carrano] | |
14468 | ETE | M. B. Efimov and S. V. Leshchinskiy. 2000. [First discovery of a fossil crocodile skull in Siberia]. In A.V. Komarov (ed.), Materialy regionalanoj konferencii geologov Sibiri, Dalshchnego Vostoka i Severoâ ’Vostoka Rossii. Gala Press, Tomsk 2:361-363 [M. Carrano/M. Carrano/M. Carrano] |
34817 | ETE | P. P. Gambaryan, A. O. Averianov, E. N. Maschenko and S. V. Leshchinskiy. 2004. Mammalian humerus from the Early Cretaceous of West Siberia. Russian Journal of Theriology 3(2):51-58 [M. Carrano/M. Oreska] |
86245 | S. V. Ivantsov, A. V. Fayngerts, S.V. Leshchinskiy, A.S. Rezviy, P.P. Skutschas and A.O. Averianov. 2016. The Lower Cretaceous continental vertebrate fauna from the Shestakovo locality (West Siberia): Results of 20-year research. In O. S. Dzyuba, E. B. Pestchevitskaya, & B. N. Shurygin (eds.), Cretaceous Ecosystems and Their Responses to Paleoenvironmental Changes in Asia and the Western Pacific. Short papers for the Fourth International Symposium of International Geoscience Programme, IGCP Project 608 121-123 [M. Carrano/M. Carrano] | |
86246 | A. E. Kostyunin. 2016. The history of discovery and research of the Shestakovo locality of Early Cretaceous vertebrates. In O. S. Dzyuba, E. B. Pestchevitskaya, & B. N. Shurygin (eds.), Cretaceous Ecosystems and Their Responses to Paleoenvironmental Changes in Asia and the Western Pacific. Short papers for the Fourth International Symposium of International Geoscience Programme, IGCP Project 608 124-126 [M. Carrano/M. Carrano] | |
38196 | E. N. Kurochkin, N. V. Zelenkov, A. O. Averianov and S. V. Leshchinskiy. 2011. A new taxon of birds (Aves) from the Early Cretaceous of Western Siberia, Russia. Journal of Systematic Palaeontology 9(1):109-117 [P. Mannion/P. Mannion] | |
40059 | S. V. Leshchinskiy, A. V. Faingerts, A. V. Voronkevich, E. N. Mashchensko, and A. O. Averianov. 2000. Predvaritel'nye rezul'taty izucheniy mestonakhozhdenii Shestakovskogo kompleksa rannemelovykh pozvonochnykh [Preliminary results of the investigation of the Shestakovo localities of Early Cretaceous vertebrates]. In A.V. Komarov (ed.), Materialy Regional'noy Konferentsii Geologov Sibiri Dal'nego Vostoka i Severo-Vostoka Rossii. Gala Press, Tomsk 2:363-366 [M. Carrano/M. Carrano/M. Carrano] | |
17882 | ETE | A. V. Lopatin and A. O. Averianov. 2006. Mesozoic mammals of Russia. 9th International Symposium on Mesozoic Terrestrial Ecosystems and Biota, Abstracts and Proceedings Volume 67-70 [M. Carrano/M. Carrano/M. Carrano] |
45970 | A. V. Lopatin, A. O. Averianov, E. N. Maschenko and S. V. Leschinskiy. 2009. Early Cretaceous mammals of Western Siberia: 2. Tegotheriidae. Paleontological Journal 43(4):92-100 [P. Mannion/J. Tennant] | |
39702 | A. V. Lopatin, E. N. Maschenko, A. O. Averianov, A. S. Rezvyi, P. P. Skutschas and S. V. Leschinskiy. 2005. Early Cretaceous Mammals from Western Siberia: 1. Tinodontidae. Paleontological Journal 39(5):523-534 [R. Benson/R. Benson] | |
39755 | A. V. Lopatin, E. N. Maschenko, and A. O. Averianov. 2010. A new genus of triconodont mammals from the Early Cretaceous of western Siberia. Doklady biological Sciences 433:282-285 [R. Benson/R. Benson/R. Benson] | |
59918 | A. V. Lopatin, E. N. Mashchenko, K. K. Tarasenko, A. V. Podlesnov, N. V. Demidenko and E. A. Kuzmina. 2015. A unique burial site of Early Cretaceous vertebrates in western Siberia (the Shestakovo 3 locality, Kemerovo Province, Russia). Doklady Biological Sciences 462:148-151 [M. Carrano/M. Carrano] | |
36988 | E. N. Maschenko and A. V. Lopatin. 1998. First record of an Early Cretaceous triconodont mammal from Siberia. Bulletin de l'Institute Royal des Sciences Naturelles de Belgique, Sciences de la Terre 68:233-236 [M. Carrano/M. Carrano] | |
45980 | E. N. Maschenko, A. V. Lopatin, and A. V. Voronkevich. 2002. A new genus of the tegotheriid docodonts (Docodonta, Tegotheriidae) from the Early Cretaceous of West Siberia. Russian Journal of Theriology 1(2):75-81 [P. Mannion/J. Tennant/J. Tennant] | |
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