Blasi 2, Arén: Late Maastrichtian, Spain

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Charales - Feistiellaceae
Feistiella sp. Schudack 1986
López-Martínez et al. 2001
Charales - Characea
López-Martínez et al. 2001
A. c. concava, A. c. mucronata, A. c. ssp.
López-Martínez et al. 2001
López-Martínez et al. 2001
Actinopteri - Pycnodontiformes
López-Martínez et al. 2001
Actinopteri - Lepisosteiformes - Lepisosteidae
López-Martínez et al. 2001
Actinopteri
Teleostei indet. Müller 1846
López-Martínez et al. 2001
Reptilia
Murelaga and Canudo 2005
Joyce 2017
Reptilia - Testudines - Bothremydidae
Murelaga and Canudo 2005
Reptilia - Testudines
López-Martínez et al. 2001
Reptilia - Crocodyliformes
López-Martínez et al. 2001
Puértolas-Pascual et al. 2016
Reptilia - Krokolithidae
López-Martínez 2000
dinosauroid tubospherulithic
    = Krokolithes sp. Hirsch 1985
Pérez-Pueyo et al. 2021
Reptilia
López-Martínez et al. 2001 2 specimens
    = Eusuchia indet. Huxley 1875
Puértolas-Pascual et al. 2016
Crocodylia indet. (Owen 1842)
López-Martínez et al. 2001
Puértolas et al. 2011
MPZ 2010/948-951; 4 teeth
López-Martínez et al. 2001 1 specimen
MPZ-2017/1137, tooth
Pérez-Pueyo et al. 2021
López-Martínez et al. 2001
1 specimen
MPZ2004/5, tooth
Canudo et al. 2005
Torices et al. 2015
Reptilia - Dromaeosauridae
López-Martínez et al. 2001 1 specimen
"Dromaeosauridae type C"; MPZ 2004/6
Torices et al. 2004
Torices et al. 2015
Torices et al. 2004 1 specimen
MPZ 98/75
Canudo et al. 2005
Torices et al. 2004 2 specimens
MPZ 98/72, 74, teeth
Canudo et al. 2005
Torices et al. 2015
Torices et al. 2004 2 specimens
MPZ 98/73, 2004/7, teeth
Canudo et al. 2005
Torices et al. 2015
Reptilia
López-Martínez et al. 2001 3 specimens
MPZ 98/76, 77, 78, teeth
Torices et al. 2015
López-Martínez et al. 2001 50 specimens
Canudo et al. 2005
MPZ 98/79, 80, 81, 82; 50 isolated teeth
Reptilia - Prismatoolithidae
López-Martínez 2000
Reptilia - Hadrosauridae
López-Martínez et al. 2001 4 specimens
BLA 2b/2
Reptilia - Spheroolithidae
Pérez-Pueyo et al. 2021
Reptilia - Squamata
Iguanidae indet. Oppel 1811
López-Martínez et al. 2001 3 specimens
Blain et al. 2010
Indeterminate lizard 1 (probably an iguanid, teiid or lacertid) - 1 partial unidentified bone with 3 teeth [MPZ 2009/323]
Reptilia
Blain et al. 2010 1 specimen
posterior caudal vertebra [MPZ 2009/331]
López-Martínez et al. 2001 2 specimens
Blain et al. 2010
Indeterminate lizard 2 - 1 left dentary [MPZ 2009/324] and 1 incomplete unidentified bone with 2 teeth [MPZ 2009/325]
Reptilia - Anguidae
Anguidae indet. Gray 1825
Blain et al. 2010 5 specimens
5 trunk vertebrae [MPZ 2009/326-330]
Amphibia
Amphibia - Temnospondyli - Palaeobatrachidae
Blain et al. 2010 36 specimens
3 incomplete maxillae [MPZ 2009/300-302], 14 ilia (1 left [MPZ 2009/303], 2 right [MPZ 2009/304-305] and 11 indet. [MPZ 2009/306-316]), 1 right scapula [MPZ 2009/317], 1 vertebra [MPZ 2009/318] and 4 radioulnae [MPZ 2009/319-322]
Amphibia - Temnospondyli
Blain et al. 2010 82 specimens
1 mandible [MPZ 2009/229], 13 ilia (2 left [MPZ 2009/ 230-231], 7 right [MPZ 2009/232-238] and 4 indet. fragments [MPZ 2009/239-242]), 9 humeri [MPZ 2009/243-251], 1 urostyle [MPZ 2009/252], 4 vertebrae [MPZ 2009/253-256], 2 sacral vertebrae [MPZ 2009/257-258], 4 radioulnae [MPZ 2009/259-262], 1 femur [MPZ 2009/263] and 36 tibiofibulae [MPZ 2009/264-299]
Amphibia - Temnospondyli - Albanerpetontidae
Blain et al. 2010 52 specimens
3 premaxillae [MPZ 2009/179-181], 4 maxillae (2 left [MPZ 2009/182-183] and 2 right [MPZ 2009/184-185]); 25 dentaries (14 left [MPZ 2009/186-199], 9 right [MPZ 2009/200-208] and 2 indet. [MPZ 2009/207-208]), 8 indeterminate partial jaws [MPZ 2009/211-218], 1 frontal [MPZ 2009/219], 5 humeri [MPZ 2009/221-225] and 2 trunk vertebrae [MPZ 2009/226-227]: misspelled "nexuosum"
see common names

Geography
Country:Spain State/province:Aragón County:Huesca
Coordinates: 42.3° North, 0.7° East (view map)
Paleocoordinates:30.3° North, 10.6° East (Wright 2013)
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period: Cretaceous Epoch: Late Cretaceous
Stage: Maastrichtian 10 m.y. bin: Cretaceous 8
Key time interval: Late Maastrichtian Foram zone:  Abathomphalus mayaroensis
Age range of interval: 72.2 - 66 m.y. ago
Stratigraphy
Geological group:Tremp Formation:La Posa
Local section:Areny Local bed:8 m
Local order:bottom to top
Stratigraphic resolution:bed
Stratigraphy comments: lower part; 6.5 m interval; upper part of chron c30n, 65.6-68.7 mya
The Conques Formation is equivalent to the "lower Red Garumen" facies. The Conques Formation was previously assigned a late maastrichtian age (Sellés and Villa 2015; Villa et al. 2015) Biostratigraphy and Sr stable isotope geochemistry suggest that the La Posa and Conques Formation are entirely early Maastrichtian, lying within chron C31r (see Caus et al. 2016; Oms et al. 2016; Fondevilla et al. 2016; Fondevilla et al. 2019).
Actually the "Grey Garumnian" = La Posa Fm.
Lithology and environment
Primary lithology:lenticular,gray marl
Includes fossils?Y
Lithology description: "grey marls", 2 m thick
Environment:lagoonal/restricted shallow subtidal
Geology comments: "lagoonal/marsh marls"
Taphonomy
Modes of preservation:body
Degree of concentration:concentrated
Size of fossils:macrofossils,mesofossils
Preservation of anatomical detail:medium
Disassociated major elements:many
Disassociated minor elements:many
Fragmentation:frequent
Collection methods and comments
Collection methods:bulk,sieve,field collection
Rock censused:5000 kg
Reason for describing collection:biostratigraphic analysis
Collection method comments: includes seven types of eggshell
Metadata
Also known as:Blasi 2a, 2b
Database number:45437
Authorizer:M. Carrano, P. Mannion, P. Holroyd Enterer:M. Carrano, P. Holroyd, J. Tennant, P. Mannion
Modifier:M. Carrano Research group:vertebrate
Created:2004-10-22 13:35:56 Last modified:2025-02-22 15:12:02
Access level:the public Released:2004-10-22 13:35:56
Creative Commons license:CC0
Reference information

Primary reference:
17660.ETE J. I. Canudo, L. Ardévol, G. Cuenca-Bescós, N. López-Martínez, X. Murelaga, X. Pereda-Suberbiola, J. I. Ruiz-Omeñaca and X. Orue-Etxebarria. 1999. New dinosaur localities near the Cretaceous/Tertiary boundary from Arén (Huesca, Spain). In J. I. Canudo & G. Cuenca-Bescós (ed.), IV European Workshop on Vertebrate Paleontology. Programme and Abstracts. Field Guide 40-41 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

37367 H.-A. Blain, J.-I. Canudo, G. Cuenca-Bescós and N. López-Martínez. 2010. Amphibians and squamate reptiles from the latest Maastrichtian (Upper Cretaceous) of Blasi 2 (Huesca, Spain). Cretaceous Research 31(4):433-446 [P. Mannion/P. Mannion]
14882ETE J. I. Canudo. 2004. El caso del dinosaurio hadrosaurio de Arén (Huesca): muerte accidental o asesinato? [The case of the hadrosaur dinosaur of Arén (Huesca): accidental death or assassination?]. Naturaleza Aragonesa 13:4-14 [M. Carrano/K. Maguire/M. Carrano]
53564 J. I. Canudo, J. L. Barco, R. Royo-Torres and J. I. Ruiz-Omeñaca. 2001. Los saurópodos (Dinosauria) del Tithónico (Jurásico Superior) y del Cretácico de Aragón [The sauropoda (Dinosauria) from the Tithonian (Upper Jurassic) and Cretaceous of Aragón]. I Jornadas Internacionales sobre Paleontología de Dinosaurios y Su Entorno Actas:309-318 [M. Carrano/M. Carrano/M. Carrano]
17600ETE J. I. Canudo, J. L. Barco, P. Cruzado-Caballero, G. Cuenca-Bescós, J. I. Ruiz-Omeñaca and R. Royo-Torres. 2005. Evidencias de predación de dinosaurios terópodos en el Maastrichtiense superior, Cretácico superior de Arén (Huesca) [Evidence of theropod dinosaur predation in the upper Maastrichtian, Upper Cretaceous of Arén (Huesca)]. Lucas Mallada 12:29-58 [M. Carrano/M. Carrano]
28738ETE J. I. Canudo, G. Cuenca-Bescós, and J. I. Ruiz-Omeñaca. 2004. Los últimos dinosaurios europeos: debates recientes sobre la extinción de los dinosaurios [The last European dinosaurs: recent debates on the extinction of the dinosaurs]. Historia Natural 7:42-51 [M. Carrano/M. Carrano]
17647ETE J. I. Canudo and J. I. Ruiz-Omeñaca. 2003. Los restos directos de dinosaurios teropódos (excluyendo Aves) en España [Direct remains of theropod dinosaurs (excluding Aves) in Spain]. Ciencias de la Tierra 26:347-373 [M. Carrano/M. Carrano]
81303 J. Company, P. Cruzado-Caballero, and J. I. Canudo. 2013. A diminutive hadrosaur from the Late Maastrichtian of the south-central Pyrenees (Spain). In F. Torcida Fernández-Baldor & P. Huerta (ed.), VI Jornadas Internacionales sobre Paleontología de Dinosaurios y su Entorno. Libro de Resúmenes 53-55 [M. Carrano/M. Carrano]
70541 J. Company, P. Cruzado-Caballero, and J. I. Canudo. 2015. Presence of diminutive hadrosaurids (Dinosauria: Ornithopoda) in the Maastrichtian of the south-central Pyrenees (Spain). Journal of Iberian Geology 41(1):71-81 [M. Carrano/M. Carrano]
42686 J. Company and Z. Szentesi. 2012. Amphibians from the Late Cretaceous Sierra Perenchiza Formation of the Chera Basin, Valencia Province, Spain. Cretaceous Research 37:240-245 [P. Mannion/P. Mannion/M. Carrano]
17658ETE P. Cruzado-Caballero and J. I. Canudo. 2006. Nuevas evidencias de ornitópodos "enanos" en el Maastrichtiense superior Europeo (Blasi 5, Arén, Huesca) [New evidences of "dwarf" ornithopods in the European upper Maastrichtian (Blasi 5, Arén, Huesca)]. IV Encuentro de Jóvenes Investigadores en Paleontología, Salamanca, Spain 75-76 [M. Carrano/M. Carrano]
28219ETE P. Cruzado-Caballero, J. I. Canudo, and J. I. Ruiz-Omeñaca. 2007. Los dinosaurios hadrosaurios (Ornithischia: Ornithopoda) del Maastrichtiense superior (Cretácico Superior) de Arén (Huesca, España) [The hadrosaurian dinosaurs (Ornithischia: Ornithopoda) from the upper Maastrichtian (Upper Cretaceous) of Arén (Huesca, Spain)]. IV Jornadas Internacionales sobre Paleontología de Dinosaurios y su Entorno, Salas de los Infantes, Spain 55-56 [M. Carrano/M. Carrano/M. Carrano]
19147ETE F. M. Dalla Vecchia. 2006. Telmatosaurus and the other hadrosaurids of the Cretaceous European Archipelago. An overview. Natura Nascosta 32:1-55 [M. Carrano/M. Carrano]
41120 F. M. Dalla Vecchia. 2009. Telmatosaurus and the other hadrosauroids of the Cretaceous European Archipelago. An update. Natura Nascosta 39:1-18 [M. Carrano/M. Carrano]
56054 F. M. Dalla Vecchia. 2014. An overview of the latest Cretaceous hadrosauroid record in Europe. In D. A. Eberth & D. C. Evans (ed.), Hadrosaurs 268-297 [M. Carrano/M. Carrano]
55204 A. Galobart, A. Santos-Cubedo, M. Suñer and J. I. Canudo. 2011. Paleontological discoveries and studies. In À. Galobart, M. Suñer, & B. Poza (eds.), Dinosaurs of Eastern Iberia 1-23 [M. Carrano/M. Carrano/M. Carrano]
80599 E. Isasmendi, A. Torices, J. I. Canudo, P. J. Currie, and X. Pereda-Suberbiola. 2022. Upper Cretaceous European theropod palaeobiodiversity, palaeobiogeography and the intra‐Maastrichtian faunal turnover: new contributions from the Iberian fossil site of Laño. Papers in Palaeontology 8(1):e1419:1-38 [M. Carrano/M. Carrano]
62124 W. G. Joyce. 2017. A Review of the Fossil Record of Basal Mesozoic Turtles. Bulletin of the Peabody Museum of Natural History 58(1):65-113 [P. Holroyd/P. Holroyd/P. Mannion]
56237 Y. Laurent. 2003. Les faunes de vertébrés continentaux du Maastrichtien supérieur d'Europe: systématique et biodiversité [The continental vertebrate faunas of the upper Maastrichtian of Europe: systematics and biodiversity]. Strata, Série 2 41:1-81 [M. Carrano/M. Carrano/M. Carrano]
61140 N. López-Martínez. 2000. Eggshell sites from the Cretaceous-Tertiary transition in south-central Pyrenees (Spain). In A. M. Bravo & T. Reyes (ed.), First International Symposium on Dinosaur Eggs and Babies, Extended Abstracts 95-115 [M. Carrano/M. Carrano]
17699ETE N. López-Martínez, J. I. Canudo, and G. Cuenca-Bescós. 1999. Latest Cretaceous eggshells from Arén (southern Pyrénées, Spain). First International Symposium on Dinosaur Eggs and Babies, Isona i Conda Dellà, Spain 35-36 [M. Carrano/M. Carrano]
11838ETE N. López-Martínez, J. I. Canudo, L. Ardèvol, X. Pereda Suberbiola, X. Orue-Etxebarria, G. Cuenca-Bescós, J. I. Ruiz-Omeñaca, X. Murelaga, and M. Feist. 2001. New dinosaur sites correlated with Upper Maastrichtian pelagic deposits in the Spanish Pyrenees: implications for the dinosaur extinction pattern in Europe. Cretaceous Research 22(1):41-61 [M. Carrano/M. Carrano/M. Carrano]
83615 M. Moreno-Azanza, L. Ezquerro, M. Pérez-Pueyo and J. M. Gasca. 2021. Huevos de dinosaurio en la Sierra Exteriores de Huesca [Dinosaur eggs in the Sierras Exteriores of Huesca]. Lucas Mallada 23:61-81 [M. Carrano/M. Carrano]
59760 X. Murelaga and J. I. Canudo. 2005. Descripción de los restos de quelonios del Maastrichtiense superior de Aren y Serraduy (Huesca) [Description of chelonian remains from the Upper Maastrichtian of Aren and Seraduy (Huesca)]. Geogaceta 38:51-54 [P. Holroyd/P. Holroyd/M. Uhen]
17698ETE O. Oms and J. I. Canudo. 2004. Datación magnetoestratigrafica de los dinosaurios del Cretácico terminal (Maastrichtiense superior) de Arén (Huesca, Unidad Surpirenaica Central) [Magnetostratigraphic dating of the terminal Cretaceous dinosaurs (upper Maastrichtian) of Arén (Huesca, central Pyrénées)]. Geo-Temas 6(5):51-54 [M. Carrano/M. Carrano]
81262 M. Pérez-Pueyo, P. Cruzado-Caballero, M. Moreno-Azanza, B. Vila, D. Castanera, J. M. Gasca, E. Puértolas-Pascual, B. Bádenas, and J. I. Canudo. 2021. The tetrapod fossil record from the uppermost Maastrichtian of the Ibero-Armorican Island: an integrative review based on the outcrops of the western Tremp Syncline (Aragón, Huesca Province, NE Spain). Geosciences 11:162:1-38 [M. Carrano/M. Carrano]
47749 A. Prieto-Marquez, F. M. D. Vecchia, R. Gaete and A. Galobart. 2013. Diversity, relationships, and biogeography of the lambeosaurine dinosaurs from the European archipelago, with description of the new aralosaurin Canardia garonnensis. PLoS One 8(7):e69835:1-44 [P. Mannion/J. Tennant/M. Carrano]
36564 E. Puértolas, J. I. Canudo, and P. Cruzado-Caballero. 2011. A new crocodylian from the late Maastrichtian of Spain: Implications for the initial radiation of crocodyloids. PLoS One 6(6):e20011:1-12 [P. Mannion/P. Mannion/M. Carrano]
76284 E. Puértolas-Pascual, A. Blanco, C. A. Brochu and J. I. Canudo. 2016. Review of the Late Cretaceous-early Paleogene crocodylomorphs of Europe: Extinction patterns across the K-PG boundary. Cretaceous Research 57:565-590 [P. Mannion/P. Mannion]
56091 A. Torices, P. J. Currie, J. I. Canudo and X. Pereda-Suberbiola. 2015. Theropod dinosaurs from the Upper Cretaceous of the South Pyrenees Basin of Spain. Acta Palaeontologica Polonica 60(3):611-626 [P. Mannion/J. Tennant/M. Carrano]
17618ETE A. Torices, J. I. Ruiz-Omeñaca, J. I. Canudo and N. López-Martínez. 2004. Nuevos datos sobre los dinosaurios terópodos (Saurischia: Theropoda) del Cretácico superior de los Pirineos Sur-Centrales (Huesca y Lleida) [New data on the theropod dinosaurs (Saurischia: Theropoda) from the Upper Cretaceous of the south-central Pyrenées (Huesca and Lleida)]. Geo-Temas 6(5):71-74 [M. Carrano/M. Carrano]
55315 B. Vila, M. Suñer, A. Santos-Cubedo, J. I. Canudo, B. Poza and A. Galobart. 2011. Saurischians through time. In A. Galobart, M. Suñer, & B. Poza (eds.), Dinosaurs of Eastern Iberia 130-168 [M. Carrano/M. Carrano]