Aguada de Lomas: Tortonian, Peru

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Chondrichthyes - Myliobatiformes - Myliobatidae
Myliobatis sp. Cuvier 1816
Chondrichthyes - Lamniformes - Otodontidae
Chondrichthyes - Lamniformes - Lamnidae
Marocco and Muizon 1988
Isurus cf. oxyrhynchus (Rafinesque 1810)
Reptilia - Odontopterygiformes - Pelagornithidae
Pelagornis sp. Lartet 1857
Chavez et al. 2007 1 specimen
Reptilia - Spheniscidae
Spheniscidae indet. Bonaparte 1831
form 2
Stucchi 2002 1 specimen
Reptilia - Suliformes - Sulidae
cf. Sula sp. Brisson 1760
Sula aff. variegata (Tschudi 1843)
Stucchi 2003 1 specimen
Reptilia - Suliformes - Phalacrocoracidae
Reptilia
Crocodylia indet. (Owen 1842)
Salas-Gismondi et al. 2022
MUSM 4010 (AGL817), MUSM 2606 (AGL765), MUSM 4011, MUSM n/n (AGL953), MUSM 4012, MUSM 4013 (AGL954)
Reptilia - Gavialidae
Eusuchia indet. Huxley 1875
Salas-Gismondi et al. 2022
MUSM 163
Salas-Gismondi et al. 2022
MUSM 4008 (AGL816), MUSM 4009 (AGL952) and MUSM 2528
Mammalia - Sirenia - Dugongidae
Domning and Aguilera 2008
Mammalia - Cetacea
Marocco and Muizon 1988
Bouetel and Muizon 2006 1 specimen
Bisconti 2012 1 specimen
Mammalia - Cetacea - Balaenopteridae
Marocco and Muizon 1988
sp. 2
Pilleri 1989 1 specimen
(1 measurement)
Marx and Kohno 2016 2 specimens
Kimura and Hasegawa 2024 1 specimen
GMNH-PV-159
Bisconti et al. 2022 1 specimen
Mammalia - Cetacea - Kogiidae
Muizon 1988
Collareta et al. 2017 1 specimen
Mammalia - Cetacea - Phocoenidae
Marocco and Muizon 1988
Muizon 1986
Mammalia - Cetacea - Pontoporiidae
Pilleri and Siber 1989 1 specimen
Lambert and Muizon 2013 5 specimens
Mammalia - Cetacea - Eurhinodelphinidae
Pilleri 1989
Mammalia - Carnivora
terrestrial
Mammalia - Carnivora - Phocidae
form 2 & form 4
n. sp.
Valenzuela-Toro et al. 2016 2 specimens
Mammalia - Nothrotheriidae
Muizon et al. 2003 1 specimen
Mammalia - Megatheriidae
Marocco and Muizon 1988
Actinopteri - Clupeiformes - Clupeoidei
Alosinae indet. (Svetovidov 1952)
Actinopteri - Siluriformes - Ariidae
Ariidae indet. Berg 1958
Actinopteri - Istiophoriformes - Sphyraenidae
see common names

Geography
Country:Peru State/province:Arequipa
Coordinates: 15.5° South, 74.8° West (view map)
Paleocoordinates:14.9° South, 72.6° West (Wright 2013)
Basis of coordinate:stated in text
Geographic resolution:local area
Time
Period: Neogene Epoch: Miocene
Stage: Tortonian 10 m.y. bin: Cenozoic 6
Key time interval: Tortonian
Age range of interval: 11.63 - 7.246 m.y. ago
Age estimate: 8.85 to 7.3 Ma (other)
Stratigraphy
Formation:Pisco
Local section:Pisco Local bed:AGL
Local order:no particular order
Stratigraphic resolution:bed
Stratigraphy comments: 7-8.8 Ma (Stucchi & Urbina, 2004), Huayquerian, ca. 8 Ma (Muizon & DeVries, 1985), between 7.3 and 7.46 (Ehret et al 2012).
The Pisco Formation at Agua de Lomas has been dated to 8.0–7.0 Ma based on K-Ar dating [74, 75]. (Marx & Fordyce, 2015).
~8.8 and 7.9 Ma for rocks associated to the AGL vertebrate-level (Ochoa et al., 2022)
Lithology and environment
Primary lithology:diatomaceous siltstone
Includes fossils?Y
Environment:foreshore
Geology comments: AGL horizon, shore environment containing protected beaches and reefs. The protected beaches and reefs are exposed to marine currents. There is a high sedimentation rate, with a shallow-to-deep shelf and upwelling.
Taphonomy
Modes of preservation:body,original phosphate
Size of fossils:macrofossils
Collection methods and comments
Collection methods:field collection
Reason for describing collection:taxonomic analysis
Metadata
Database number:45485
Authorizer:M. Uhen, P. Mannion Enterer:N. Pyenson, P. Mannion, M. Uhen, C. Peredo
Modifier:P. Mannion Research group:vertebrate
Created:2004-10-25 11:12:40 Last modified:2025-02-22 15:12:02
Access level:the public Released:2004-10-25 11:12:40
Creative Commons license:CC0
Reference information

Primary reference:
16797. C. Muizon and T. J. DeVries. 1985. Geology and paleontology of late Cenozoic marine deposits in the Sacaco area (Peru). Geologische Rundschau 74(3):547-563 [M. Uhen/M. Uhen/M. Uhen]

Secondary references:

44129 M. Bisconti. 2012. Comparative osteology and phylogenetic relationships of Miocaperea pulchra, the first fossil pygmy right whale genus and species (Cetacea, Mysticeti, Neobalaenidae). Zoological Journal of the Linnean Society 166:876-911 [M. Uhen/M. Uhen]
80424 M. Bisconti, D. Ochoa, M. Urbina and R. Salas-Gismondi. 2022. Archaebalaenoptera eusebioi, a new rorqual from the late Miocene of Peru (Cetacea, Mysticeti, Balaenopteridae) and its impact in reconstructing body size evolution, ecomorphology and palaeobiogeography of Balaenopteridae. Journal of Systematic Palaeontology [M. Uhen/M. Uhen]
18255 V. Bouetel and C. Muizon. 2006. The anatomy and relationships of Piscobalaena nanna (Cetacea, Mysticeti), a Cetotheriidae s.s. from the early Pliocene of Peru. Geodiversitas 28(2):319-395 [M. Uhen/M. Uhen]
39156 L. Brand, M. Urbina, A. Chadwick, T. J. DeVries, and R. Esperante. 2011. A high resolution stratigraphic framework for the remarkable fossil cetacean assemblage of the Miocene/Pliocene Pisco Formation, Peru. Geobios 31:414-425 [M. Uhen/M. Uhen/M. Uhen]
34370 M. Chavez, M. Stucchi, and M. Urbina. 2007. El registro de Pelagornithidae (Aves: Pelecaniformes) y la avifauna neogena del Pacifico sudeste. Bulletin de l'Institut Francais d'Etudes Andines 36(2):175-197 [M. Uhen/M. Uhen]
64039 A. Collareta, O. Lambert, C. Muizon, M. Urbina, and G. Bianucci. 2017. Koristocetus pescei gen. et sp. nov., a diminutive sperm whale (Cetacea: Odontoceti: Kogiidae) from the late Miocene of Peru. Fossil Record 20:259-278 [M. Uhen/M. Uhen]
27454 D. P. Domning and O. A. Aguilera. 2008. Fossil Sirenia of the West Atlantic and Caribbeaan Region. VIII. Nanosiren garciae, gen. et sp. nov. and Nanosiren sanchezi, sp. nov. Journal of Vertebrate Paleontology 28(2):479-500 [M. Uhen/M. Uhen]
46232 D. J. Ehret, B. J. MacFadden, D. S. Jones, T. J. DeVries, D. A. Foster and R. Salas-Gismondi. 2012. Origin of the white shark Carcharodon (Lamniformes: Lamnidae) based on recalibration of the upper Neogene Pisco Formation of Peru. Palaeontology 55(6):1139-1153 [M. Uhen/M. Uhen]
88186 T. Kimura and Y. Hasegawa. 2024. A new species of Late Miocene balaenopterid, Incakujira fordycei, from Sacaco, Peru. Bulletin of the Gunma Museum of Natural History 28:1-14 [M. Uhen/M. Uhen]
46791 O. Lambert and C. Muizon. 2013. A new long-snouted species of the Miocene pontoporiid dolphin Brachydelphis and a review of the Mio-Pliocene marine mammal levels in the Sacaco Basin, Peru. Journal of Vertebrate Paleontology 33(3):709-721 [M. Uhen/M. Uhen]
12182 R. Marocco and C. d.e. Muizon. 1988. Los vertebrados del Neogeno de La Costa Sur del Perú: Ambiente sedimentario y condiciones de fosilización. Bulletin de l'Institut Frances d'Etudes Andines 17(2):105-117 [M. Uhen/M. Uhen/M. Uhen]
55050 F. G. Marx and R. E. Fordyce. 2015. Baleen boom and bust: a synthesis of mysticete phylogeny, diversity and disparity. Royal Society Open Science 2:140434 [M. Uhen/M. Uhen]
60487 F. G. Marx and N. Kohno. 2016. A new Miocene baleen whale from the Peruvian desert. Royal Society Open Science 3(10):160542 [M. Uhen/C. Peredo]
12756 C. Muizon. 1986. Un nouveau Phocoenidae (Odontoceti, Mammalia) du Miocene superieur de la Formation Pisco (Perou). Comptes - rendus hebdomadaires des seances de l'Academie des Sciences Serie II 303(16):1509-1512 [M. Uhen/M. Uhen/M. Uhen]
11895 C. Muizon. 1988. Les vertebres fossiles de la Formation Pisco (Perou). Troisieme partie: Les Odontocetes (Cetacea, Mammalia) du Miocene. Editions Recherche sur les Civilisations (78)1-244 [M. Uhen/N. Pyenson/M. Uhen]
16787 C. Muizon, H. G. McDonald, R. Salas and M. Urbina. 2003. A new early species of the aquatic sloth Thalassocnus (Mammalia, Xenarthra) from the late Miocene of Peru. Journal of Vertebrate Paleontology 23(4):886-894 [M. Uhen/M. Uhen/M. Uhen]
81296 D. Ochoa, T. J. DeVries, K. Quispe, A. Barbosa-Espitia, R. Salas-Gismondi, D. A. Foster, R. Gonzales, S. Revillon, R. Berrospi, L. Pairazamán, J. Cardich, A. Perez, P. Romero, M. Urbina, and M. Carré. 2022. Age and provenance of the Mio-Pleistocene sediments from the Sacaco area, Peruvian continental margin. Journal of South American Earth Sciences 116:103799 [M. Uhen/M. Uhen/M. Uhen]
15412 G. Pilleri. 1989. Balaenoptera siberi, ein neuer spatmiozaner bartenwal aus der Pisco-formation Perus. Beitrage zur Palaontologie der Cetaceen Perus 65-84 [M. Uhen/M. Uhen/M. Uhen]
15413 G. Pilleri. 1989. Vorkommen von Eurhinodelphidae (Cetacea: Odontoceti) in der Pisco-Formation Perus?. Beitrage zur Paleontologie der Cetaceen Perus 221-223 [M. Uhen/M. Uhen]
26379 G. Pilleri. 1990. Paratypus von Balaenoptera siberi (Cetacea: Mysticeti) aus der Pisco Formation Perus. Beiträge zur Paläontologie der Cetaceen und Pinnipedier der Pisco Formation Perus II:205-236 [M. Uhen/M. Uhen/J. Alroy]
12843 G. Pilleri and H. J. Siber. 1989. Piscorhynchus aenigmaticus, ein neuer Miozaner Zahnwal aus der Pisco-Formation Perus. Beitrage zur Palaontologie der cetaceen Perus 195-203 [M. Uhen/M. Uhen/M. Uhen]
81650 R. Salas-Gismondi, D. Ochoa, S. Jouve, P. E. Romero, J. Cardich, A. Perez, T. DeVries, P. Baby, M. Urbina and M. Carré. 2022. Miocene fossils from the southeastern Pacific shed light on the last radiation of marine crocodylians. Proceedings of the Royal Society B 289:20220380:1-10 [P. Mannion/P. Mannion]
34468 M. Stucchi. 2002. Una nueva especie de Spheniscus (Aves: Spheniscidae) de la Formacion Pisco, Peru. Boletim de la Sociedad Geologica del Peru 94:17-24 [M. Uhen/M. Uhen/M. Uhen]
54566 M. Stucchi. 2003. Los Piqueros (Aves: Sulidae) de la Formacion Pisco, Peru. Boletin de la Sociedad Geologica de Peru 95:75-91 [M. Uhen/M. Uhen]
18990 M. Stucchi and M. Urbina. 2004. Ramphastosula (aves, Sulidae): a new genus from the early Pliocene of the Pisco Formation, Peru. Journal of Vertebrate Paleontology 24(4):974-978 [M. Uhen/M. Uhen]
57210 A. Valenzuela-Toro, N. D. Pyenson, C. S. Gutstein and M. E. Suarez. 2016. A new dwarf seal from the Late Neogene of South America and the evolution of pinnipeds in the southern hemisphere. Papers in Palaeontology 2(1):101-115 [M. Uhen/M. Uhen/M. Uhen]