Coten End Quarry: Anisian, United Kingdom

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Dipnoi - Ceratodontidae
Benton and Spencer 1995 1 specimen
Amphibia - Temnospondyli - Cyclotosauridae
Paton 1974 4 specimens
WARMS Gz 13, 14, 26, 36
Labyrinthodon leptognathus n. sp. Owen 1841
Owen 1842 4 specimens
WARMS Gz 6, 11, 35, 38
Amphibia - Temnospondyli
Benton et al. 1994
Benton and Spencer 1995
Benton et al. 1994
Benton and Spencer 1995
Reptilia
? Sauria indet. Gauthier 1984
1 specimen
tooth, vertebra
Galton 2012 6 specimens
CAMSM G.353A; WARMS G.1036, G.981/982
Galton 2012 1 specimen
BGS GSLC 4873
Galton 2012 2 specimens
BGS GSLC 90742, 90743
Galton 2012 1 specimen
WARMS G.957
Galton 2012 1 specimen
WARMS G.956.1
Rauisuchia indet. (Huene 1942)
Galton 2012 6 specimens
WARMS G.4713, G.4; G.128; BIRUG 2473; CAMSM G.344a-f, G.357
Galton 2012 1 specimen
WARMS G.954
Galton 2012 1 specimen
WARMS G.969
Galton 2012 2 specimens
CAMSM G.352; BGS GSLC 59744
Allen 1909 1 specimen
Benton et al. 1994
BMNH R2628, cervical vertebra
Galton and Walker 1996
Galton 2012
Benton et al. 1994 5 specimens
WARMS Gz 3a,b, right ilium (holotype). WARMS G.1/2, sacral vertebrae; WARMS 121, dorsal centrum; WARMS G.5, WARMS G.970
4 specimens
    = Cladyodon lloydii n. gen., n. sp. Owen 1841
Owen 1842
WARMS Gz 7, 8, 954, 957, teeth
Newton 1905
Ezcurra et al. 2016 14 specimens
WARMS Gz17, dorsal vertebra; Gz19, left ischium; Gz34, interclavicle; Gz950/1, 2, right dentary and partial splenial; Gz955, small right maxilla; Gz959, posterior right dentary; Gz960, right maxilla; Gz4712, left dentary; Gz4715, partial snout; Gz6097, partial skull; BMNH R2623, partial dentary; R8494, right maxilla with portions of palatine, ectopterygoid & jugal; BGS/GSM 59745, 90493, left maxillae
Allen 1909 1 specimen
BMNH R8495, mandible and ventral parts of skull, fits to WARMS Gz6097 (holotype)
Benton 1990
Previously referred specimens were considered indeterminate rhynchosaurids by Ezcurra et al. (2016)
Benton and Walker 1996 2 specimens
WARMS Gz 21, proximal left femur; Gz4714, left ilium
see common names

Geography
Country:United Kingdom State/province:England County:Warwickshire
Coordinates: 52.3° North, 1.6° West (view map)
Paleocoordinates:29.6° North, 20.3° East (Wright 2013)
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period: Triassic Epoch: Middle Triassic
Stage: Anisian 10 m.y. bin: Triassic 2
Key time interval: Anisian
Age range of interval: 246.7 - 241.464 m.y. ago
Stratigraphy
Geological group:Sherwood Sandstone Formation:Bromsgrove Sandstone
Stratigraphic resolution:group of beds
Stratigraphy comments: equate with the former 'Waterstones' and 'Building Stones'. Reptile specimens were found mostly in the 'Building Stones' and 'Dirt-bed' horizons while most amphibians were found in the 'Waterstones'
Palynological data indicates an Anisian age (Benton et al. 1994)
Lithology and environment
Primary lithology:"cross stratification",fine,brown,red poorly lithified sandstone
Secondary lithology: marl
Includes fossils?Y
Lithology description: "10-11 m of channeled and cross-bedded water-laid buff and red sandstone in units 1-3 m thick, with occasional impersistent marl and clay bands 0.1-0.5 m thick."
Environment:"channel"
Geology comments: "overbank pools subsequently broken up by flood waters"
Taphonomy
Modes of preservation:body,trace
Size of fossils:macrofossils,mesofossils
Preservation of anatomical detail:good
Articulated whole bodies:none
Associated major elements:none
Disassociated major elements:many
Disassociated minor elements:many
Fragmentation:occasional
Temporal resolution:time-averaged
Spatial resolution:parautochthonous
Collection methods and comments
Collection methods:selective quarrying,mechanical,field collection,survey of museum collection
Reason for describing collection:taxonomic analysis
Museum repositories:BMNH
Collection method comments: quarried for building stone, abandoned by 1890
Metadata
Also known as:Coton End Quarry
Database number:57944
Authorizer:M. Carrano, R. Butler Enterer:K. Maguire, M. Carrano, R. Butler
Modifier:M. Carrano Research group:vertebrate
Created:2006-01-11 11:02:56 Last modified:2025-02-22 15:12:02
Access level:the public Released:2006-01-11 11:02:56
Creative Commons license:CC0
Reference information

Primary reference:
51968. R. I. Murchison and H. E. Strickland. 1837. On the upper formations of the New Red System in Gloucestershire, Worcestershire, and Warwickshire; showing that the Red or Saliferous, including a peculiar band of sandstone, represent the "Keuper" or "Marnes irisées;" with some account of the underlying sandstone of Ombersley, Bromsgrove, and Warwick, priving that it is the "Bunter Sandstein" or "Grès bigarré" of foreign geologists. Transactions of the Geological Society of London, series 2 5(26):331-348 [M. Carrano/M. Carrano]

Secondary references:

55053 H. A. Allen. 1909. List of British Triassic fossils in the Warwick Museum. Report of the British Association for the Advancement of Science 78:274-277 [M. Carrano/M. Carrano/M. Carrano]
28945 M. J. Benton. 1990. The species of Rhynchosaurus, a rhynchosaur (Reptilia, Diapsida) from the Middle Triassic of England. Philosophical Transactions of the Royal Society of London, Series B 328:213-306 [J. Mueller/T. Liebrecht/R. Butler]
29836 M. J. Benton and D. J. Gower. 1997. Richard Owen's giant Triassic frogs: archosaurs from the Middle Triassic of England. Journal of Vertebrate Paleontology 17(1):74-88 [R. Butler/R. Butler]
15587ETE M. J. Benton and P. S. Spencer. 1995. Fossil Reptiles of Great Britain. Chapman & Hall, London 1-386 [M. Carrano/K. Maguire/J. Tennant]
33374 M. J. Benton and A. D. Walker. 1996. Rhombopholis, a prolacertiform reptile from the Middle Triassic of England. Palaeontology 39(3):763-782 [R. Butler/R. Butler]
24929ETE M. J. Benton, G. Warrington, A. J. Newell and P. S. Spencer. 1994. A review of the British Middle Triassic tetrapod assemblages. In N. C. Fraser & H.-D. Sues (eds.), In the Shadow of the Dinosaurs: Early Mesozoic Tetrapods. Cambridge University Press, Cambridge 131-160 [M. Carrano/M. Carrano]
60747 M. D. Ezcurra, F. Montefeltro, and R. J. Butler. 2016. The early evolution of rhynchosaurs. Frontiers in Ecology and Evolution 3:142 [R. Butler/R. Butler/M. Carrano]
57297 P. M. Galton. 2012. Notes on bones and teeth of Bromsgroveia and other archosaurian reptiles from the lower Middle Triassic (Anisian) of the English Midlands. Revue de Paleobiologie, Genève 31(1):171-204 [R. Butler/R. Butler/M. Carrano]
33375 P. M. Galton and A. D. Walker. 1996. Supposed prosauropod dinosaurs from Middle Triassic of England referred to Parasuchia and Dinosauriformes. Neues Jahrbuch für Geologie und Paläontologie, Monatshefte 1996:727-738 [R. Butler/R. Butler]
55054 A. R. Horwood. 1909. Bibliographical notes on the flora and fauna of the Trias, 1826–76. Report of the British Association for the Advancement of Science 78:277-282 [M. Carrano/M. Carrano]
16877ETE F. v. Huene. 1908. Eine Zusammenstellung über die englische Trias und das Alter ihrer Fossilien [A compilation on the English Trias and the age of its fossils]. Centralblatt für Mineralogie, Geologie und Paläontologie 1908:9-17 [M. Carrano/M. Carrano]
63762 G. A. Mantell. 1854. The Medals of Creation; or, First Lessions in Geology, and the Study of Organic Remains 2:447-930 [M. Carrano/M. Carrano/M. Carrano]
51970 E. T. Newton. 1905. Notes on the Triassic fossils (excluding Rhaetic) in the Museum of the Geological Survey at Jermyn Street, London. Report of the British Association for the Advancement of Science 1904:282-285 [M. Carrano/M. Carrano]
14071ETE R. Owen. 1842. Report on British fossil reptiles, part II. Report of the British Association for the Advancement of Science 11:60-204 [M. Carrano/M. Carrano/M. Carrano]
90505% 37320R. L. Paton. 1974. Capitosauroid labyrinthodonts from the Trias of England. Palaeontology 17(2):253-289 [J. Head/J. Head/J. Head]
82858 W. A. S. Sarjeant. 1974. A history and bibliography of the study of fossil vertebrate footprints in the British Isles. Palaeogeography, Palaeoclimatology, Palaeoecology 16:265-378 [M. Carrano/M. Carrano]
30971ETE A. S. Woodward and C. D. Sherborn. 1890. A Catalogue of British Fossil Vertebrata. Dulao & Company, London 1-396 [M. Carrano/M. Carrano/M. Carrano]