Stark Quarry, Schamhaupten: Late Kimmeridgian, Germany
collected by H. Balzer, G. Viohl, U. Gerhard, M. Röper 1988–1998

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Phaeophyceae
Viohl and Zapp 2007
Equisetopsida - Araucariaceae
Renesto and Viohl 1997
Araucarites sp. Presl 1833
Viohl and Zapp 2007
Gymnospermopsida - Corystospermales - Corystospermaceae
Viohl and Zapp 2007
Sphenopteridae
Sphenopteris sp. Sternberg 1825
Viohl and Zapp 2007
unclassified
Muensteria vermicularis Sternberg 1833
Viohl and Zapp 2007
Anthozoa - Scleractinia
Renesto and Viohl 1997
Echinoidea
Renesto and Viohl 1997
Echinoidea - Pygasteridae
Viohl and Zapp 2007
Crinoidea - Roveacrinida - Saccocomidae
Saccocoma tenella (Goldfuss 1831)
Viohl and Zapp 2007
Crinoidea - Millericrinida - Millericrinidae
Millericrinus sp. d'Orbigny 1840
Viohl and Zapp 2007
Chondrichthyes - Heterodontiformes - Heterodontidae
Kriwet 2008 1 individual
JM Scha 728
Coelacanthimorpha - Coelacanthiformes - Latimeriidae
Viohl and Zapp 2007
Reptilia - Eurysternidae
Joyce and Zelenitsky 2002 1 individual
JM SCHA 101, shell with eggs
Reptilia
Karl et al. 2012 2 individuals
JME Scha 97, 70
Joyce 2000 1 individual
JME Scha 70
Reptilia - Pterosauria - Rhamphorhynchidae
Viohl and Zapp 2007 2 specimens
Reptilia
Göhlich and Chiappe 2006 1 individual
JME SCh 200 (8 measurements)
Reptilia - Metriorhynchidae
Dakosaurus maximus (Plieninger 1846)
Andrade et al. 2010
Reptilia - Ichthyosauria
Ichthyosauria indet. de Blainville 1835
Viohl and Zapp 2007 1 specimen
sclerotic ring
Reptilia - Rhynchocephalia - Sphenodontidae
Renesto and Viohl 1997 1 individual
JME SCHA 40
Actinopteri - Lepisosteiformes
Viohl and Zapp 2007
JME Scha 80
López-Arbarello and Sfercoa 2009
Actinopteri - Macrosemiiformes
Arratia and Schultze 2012 1 individual
JM-E SCH 102 (found on same slab as Juravenator starki holotype)
Actinopteri - Ichthyodectiformes
1 individual
JM SCh 30a, b
Allothrissops sp. Nybelin 1964
1 individual
BM.f.7286
Actinopteri - Leptolepidiformes - Leptolepididae
Tharsis sp. Giebel 1848
Viohl and Zapp 2007
unclassified
Viohl and Zapp 2007
Renesto and Viohl 1997
Viohl and Zapp 2007
Actinopteri - Elopiformes - Elopidae
1 individual
JM SCh 85
= "Elops-like sp. 1"
Actinopteri - Pycnodontiformes
Viohl and Zapp 2007
Ostracoda
Renesto and Viohl 1997
Malacostraca - Decapoda - Axiidae
Etallonia hoellorum n. sp. Polz 1999
Viohl and Zapp 2007
Malacostraca - Decapoda - Glypheidae
Glyphea sp. Von Meyer 1835
Viohl and Zapp 2007
Malacostraca - Decapoda
Palinuroidea indet. Latreille 1802
Viohl and Zapp 2007
Malacostraca - Decapoda - Coleiidae
Coleia longipes (Fraas 1855)
Viohl and Zapp 2007
JME Scha 1531
Malacostraca - Isopoda
Isopoda indet. Latreille 1817
Viohl and Zapp 2007
on skeleton of Juravenator
Insecta - Orthoptera
Renesto and Viohl 1997
Viohl and Zapp 2007
Bivalvia
Renesto and Viohl 1997
Bivalvia - Ostreida - Gryphaeidae
Viohl and Zapp 2007
Bivalvia - Pectinida - Propeamussiidae
Viohl and Zapp 2007
Gastropoda - Trochidae
Buckmanina sp. Cossmann 1920
Viohl and Zapp 2007
Gastropoda - Pleurotomariidae
Viohl and Zapp 2007
Cephalopoda - Belemnitida - Mesohibolitidae
Viohl and Zapp 2007
Cephalopoda - Belemnitida - Belemnotheutidae
Acanthoteuthis sp. Wagner in Munster 1839
Viohl and Zapp 2007
Cephalopoda - Vampyromorpha
Viohl and Zapp 2007
Cephalopoda - Octopoda - Muensterellidae
Fuchs et al. 2019 1 specimen
JME SOS Scha 709
Cephalopoda - Ammonoidea
Renesto and Viohl 1997
35 spp.
Cephalopoda - Ammonitida - Oppeliidae
Oppeliidae indet. Bonarelli 1894
Viohl and Zapp 2007
Brachiopoda
Brachiopoda indet. (Cuvier 1805)
Renesto and Viohl 1997
Rhynchonellata - Rhynchonellida - Basiliolidae
Lacunosella sp. Wisniewska 1932
Viohl and Zapp 2007
Rhynchonellata - Rhynchonellida - Cyclothyrididae
Viohl and Zapp 2007
Rhynchonellata - Terebratulida - Zeilleriidae
Viohl and Zapp 2007
Rhynchonellata - Terebratulida - Terebratulidae
Viohl and Zapp 2007
Stenolaemata - Tubuliporata - Diastoporidae
"Berenicea" sp. Lamouroux 1821
Viohl and Zapp 2007
unclassified
Viohl and Zapp 2007
Goniolina sp. de Saporta 1891
Viohl and Zapp 2007
see common names

Geography
Country:Germany State/province:Bayern County:Eichstätt
Coordinates: 48.9° North, 11.6° East (view map)
Paleocoordinates:32.3° North, 24.2° East (Wright 2013)
Basis of coordinate:estimated from map
Geographic resolution:outcrop
Time
Period: Jurassic Epoch: Late Jurassic
Stage: Kimmeridgian 10 m.y. bin: Jurassic 5
Key time interval: Late Kimmeridgian
Age range of interval: 152.21 - 149.2 m.y. ago
Stratigraphy
Geological group:Weißjura Formation:Painten
Stratigraphic resolution:group of beds
Stratigraphy comments: Malm Epsilon 2 (setatum subzone), Weissjura epsilon; listed as both Solnhofen Formation, and as Painten Fm./Schamhaupten Subformation, "Profilschicht E3". Beckeri zone, Ulmense subzone.
Lithology and environment
Primary lithology: cherty/siliceous lime mudstone
Secondary lithology: bindstone
Includes fossils?Y
Includes fossils?Y
Lithology description: "silicified, laminated limestone" and bindstones
Environment:lagoonal/restricted shallow subtidal
Taphonomy
Modes of preservation:body,adpression,soft parts
Lagerst�tten type:conservation
Size of fossils:macrofossils
Spatial orientation:life position
Preservation of anatomical detail:excellent
Articulated whole bodies:all
Temporal resolution:snapshot
Spatial resolution:autochthonous
Collection methods and comments
Collection methods:selective quarrying,surface (in situ),mechanical,field collection
Reason for describing collection:taxonomic analysis
Collectors:H. Balzer, G. Viohl, U. Gerhard, M. Röper Collection dates:1988–1998
Collection method comments: collected by Jura Museum and University of Bonn. Taxa not separated into beds, but clearly there are differences in abundances between layers - could be separated into collections if future publications provide more info.
Metadata
Database number:59175
Authorizer:M. Carrano, J. Alroy, P. Mannion, M. Clapham Enterer:K. Maguire, J. Alroy, M. Carrano, J. Tennant, M. Clapham, C. Feng
Modifier:M. Carrano Research group:vertebrate
Created:2006-03-20 10:17:39 Last modified:2025-02-22 15:12:02
Access level:the public Released:2006-03-20 10:17:39
Creative Commons license:CC0
Reference information

Primary reference:
27128.ETE G. Arratia. 2000. Remarkable teleostean fishes from the Late Jurassic of southern Germany and their phylogenetic relationships. Mitteilungen aus dem Museum für Naturkunde in Berlin, Geowissenschaften Reihe 3:137-179 [M. Carrano/M. Carrano]

Secondary references:

43156 M. B. Andrade, M. T. Young, J. B. Desojo and S. L. Brusatte. 2010. The evolution of extreme hypercarnivory in Metriorhynchidae (Mesoeucrocodylia: Thalattosuchia) based on evidence from microscopic denticle morphology. Journal of Vertebrate Paleontology 30(5):1451-1465 [M. Young/M. Young]
71014 G. Arratia and H.-P. Schultze. 2012. The macrosemiiform fish companion of the Late Jurassic theropod Juravenator from Schamhaupten, Bavaria, Germany. Fossil Record 15(5):5-25 [M. Carrano/M. Carrano]
38017 L. M. Chiappe and U. B. Göhlich. 2010. Anatomy of Juravenator starki (Theropoda: Coelurosauria) from the Late Jurassic of Germany. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 258(3):257-296 [M. Carrano/M. Carrano]
69677 D. Fuchs, Y. Iba, A. Heyng, M. Iijima, C. Klug, N. L. Larson, and G. Schweigert. 2019. The Muensterelloidea: phylogeny and character evolution of Mesozoic stem octopods. Papers in Palaeontology 6:31-92 [R. Butler/E. Dunne/M. Clapham]
16916ETE U. B. Göhlich and L. M. Chiappe. 2006. A new carnivorous dinosaur from the Late Jurassic Solnhofen archipelago. Nature 440:329-332 [M. Carrano/K. Maguire/M. Carrano]
26842ETE U. B. Göhlich, H. Tischlinger, and L. M. Chiappe. 2006. Juravenator starki (Reptilia, Theropoda), ein neuer Raubdinosaurier aus dem Oberjura der Südlichen Frankenalb (Süddeutschland): Skelettanatomie und Weichteilbefunde [Juravenator starki (Reptilia, Theropoda), a new predatory dinosaur from the Upper Jurassic of the southern Franconian Alb (southern Germany): skeletal anatomy and soft tissues]. Archaeopteryx 24:1-26 [M. Carrano/M. Carrano]
27130ETE W. Joyce and D. K. Zelenitsky. 2002. Turtle egg pseudomorphs from the Late Jurassic of Schamhaupten, Germany. Archaeopteryx 20:57-62 [M. Carrano/M. Carrano]
44043 W. G. Joyce. 2000. The first complete skeleton of Solnhofia parsoni (Cryptodira, Eurysternidae) from the Upper Jurassic of Germany and its taxonomic implications. Journal of Paleontology 74(4):684-700 [R. Benson/R. Benson]
46818 H.-V. Karl, G. Tichy, and D. Valdisberri. 2012. Sontiochelys cretacea Stache, 1905 and new description of the families Eurysternidae Dollo, 1886 and Thalassemydidae Rutimeyer, 1873 (Testudines: Cryptodira). Studia Geologica Salmanticensia 48(1):45-76 [P. Mannion/J. Tennant/P. Wagner]
30504 J. Kriwet. 2008. Anew species of extinct bullhead sharks, Paracestracion viohli sp. nov. (Neoselachii, Heterodontiformes), from the Upper Jurassic of South Germany. Acta Geologica Polonica 58(2):235-241 [J. Alroy/J. Alroy]
37653 A. López-Arbarello and E. Sfercoa. 2009. New Semionotiform (Actinopterygii, Neopterygii) from the Late Jurassic of Southern Germany. Journal of Systematic Palaeontology 9:197-215 [M. Clapham/C. Feng/M. Clapham]
64581 O. W. M. Rauhut, C. Foth, and H. Tischlinger. 2018. The oldest Archaeopteryx (Theropoda: Avialiae): a new specimen from the Kimmeridgian/Tithonian boundary of Schamhaupten, Bavaria. PeerJ 6:e4191 [M. Uhen/M. Uhen]
78879 A. G. Reisdorf and M. Wuttke. 2012. Re-evaluating Moodie’s opisthotonic-posture hypothesis in fossil vertebrates part I: reptiles—the taphonomy of the bipedal dinosaurs Compsognathus longipes and Juravenator starki from the Solnhofen Archipelago (Jurassic, Germany). Palaeobiodiversity and Palaeoenvironments 92(1):119-168 [M. Carrano/M. Carrano]
27129ETE S. Renesto and G. Viohl. 1997. A sphenodontid (Reptilia, Diapsida) from the late Kimmeridgian of Schamhaupten (Southern Franconian Alb, Bavaria, Germany). Archaeopteryx 15:27-46 [M. Carrano/M. Carrano]
27315ETE G. Viohl and M. Zapp. 2007. Schamhaupten, an outstanding Fossil-Lagerstätte in a silicified Plattenkalk around the Kimmeridgian-Tithonian boundary (Southern Franconian Alb, Bavaria). Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 245(1):127-142 [M. Carrano/M. Carrano]