Verdigris Coulee (UA-MR-6): Late Santonian, Canada

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Chondrichthyes - Hybodontiformes - Lonchidiidae
Lonchidion sp. Estes 1964
Brinkman et al. 2017 1 specimen
Chondrichthyes - Orectolobiformes - Orectolobidae
Orectolobidae indet. Jordan and Fowler 1903
Brinkman et al. 2017 6 specimens
Chondrichthyes - Rajiformes
Pseudomyledaphus madseni Kirkland et al. 2013
Brinkman et al. 2017 76 specimens
Actinopteri - Lepisosteiformes - Lepisosteidae
Lepisosteidae indet. Cuvier 1825
Brinkman et al. 2017 17 specimens
Actinopteri - Amiiformes - Amiidae
Amiinae indet. Bonaparte 1837
Brinkman et al. 2017 25 specimens
Actinopteri - Aspidorhynchiformes - Aspidorhynchidae
Belonostomus longirostris (Lambe 1902)
Brinkman et al. 2017 1 specimen
Actinopteri
Teleostei informal indet. U-4
Brinkman et al. 2017 20 specimens
"type U-4"
Teleostei informal indet. O
Brinkman et al. 2017 10 specimens
"type O"
Actinopteri - Hiodontiformes - Hiodontidae
Hiodontidae indet.
Brinkman et al. 2017 11 specimens
Actinopteri - Osteoglossiformes
Wilsonichthys sp. Murray et al. 2016
Brinkman et al. 2017 41 specimens
Actinopteri
Elopomorpha indet. (Greenwood et al. 1966)
Brinkman et al. 2017 3 specimens
Actinopteri - Elopiformes
? Ostariostoma sp. Schaeffer 1949
Brinkman et al. 2017 3 specimens
Ostariostoma sp. Schaeffer 1949
Brinkman et al. 2017 1 specimen
Actinopteri
Otophysa informal indet. U-3/BvD
Brinkman et al. 2017 64 specimens
"Teleost U-3/BvD"
Actinopteri - Ellimmichthyiformes
Horseshoeichthys sp. Newbrey et al. 2010
Brinkman et al. 2017 26 specimens
Actinopteri - Esociformes - Esocidae
Estesesox foxi Wilson et al. 1992
Brinkman et al. 2017 3 specimens
Actinopteri
Acanthomorpha indet. (Rosen 1973)
Brinkman et al. 2017 11 specimens
Amphibia - Temnospondyli - Albanerpetontidae
Albanerpeton sp. Estes and Hoffstetter 1976
Brinkman et al. 2017 1 specimen
Albanerpeton galaktion n. sp. Fox and Naylor 1982
Fox and Naylor 1982 39 specimens
UA 16203-16235, 16237-16242
Albanerpeton nexuosus Estes 1981
Gardner 2000 40 specimens
Amphibia - Temnospondyli - Urodela
Urodela indet. (Duméril 1805)
Brinkman et al. 2017 104 specimens
Amphibia - Temnospondyli
Anura indet. (Fischer von Waldheim 1813)
Brinkman et al. 2017 2 specimens
Reptilia - Teiidae
Sphenosiagon simplex Gao and Fox 1991
Gao and Fox 1996 2 specimens
UALVP 29836-29837 (partial dentaries)
Reptilia - Anguidae
Anguidae indet. Gray 1825
Gao and Fox 1996 3 specimens
"gen. et sp. indet."; UALVP 33379 (left dentary) plus many unnumbered fragments
Reptilia - Xenosauridae
Xenosauridae indet. Cope 1866
Gao and Fox 1996 5 specimens
"gen. et sp. indet."; UALVP 29914 (jugal), 29915-29918 (maxilla) plus unnumbered specimens
Reptilia - Necrosauridae
Necrosauridae indet. Hoffstetter 1943
Gao and Fox 1996 2 individuals
"gen. et sp. nov."; UALVP 29787, 33383 (right dentaries)
Reptilia - Chamopsiidae
Chamops sp. Marsh 1892
Gao and Fox 1996
partial maxillae/dentaries
Reptilia - Iguanidae
Iguanidae informal indet. A
Gao and Fox 1996 1 specimen
"genus and species new A"; UALVP 29908 (partial left maxilla)
Cnephasaurus locustivorus n. gen., n. sp. Gao and Fox 1996
Gao and Fox 1996 1 specimen
UALVP 29921 (holotype right maxilla), 33386 (partial maxilla)
Reptilia - Scincidae
Aocnodromeus corrugatus n. gen., n. sp. Gao and Fox 1996
Gao and Fox 1996 2 individuals
UALVP 29905 (holotype left dentary), 29906 (left dentary)
Penemabuya antecessor n. gen., n. sp. Gao and Fox 1996
Gao and Fox 1996 6 specimens
UALVP 29789 (holotype dentary), 29788, 29827-29828, and several unnumbered jaw fragments
Reptilia
Coniophis cf. precedens Marsh 1892
Fox 1975
isolated vertebrae
Reptilia - Aniliidae
Aniliidae indet. Fitzinger 1826
Fox 1975
isolated vertebra (UA 12019)
Mammalia
Potamotelses aquilensis n. sp. Fox 1972
Fox 1972
Theria indet. Parker and Haswell 1897
UA 12088
Mammalia - Alphadontidae
Alphadon sp. Simpson 1927
Fox 1971
    = Alphadontidae indet. Marshall et al. 1990
Alroy 2002
said by Fox 1979a to be a new genus
Alphadon creber n. sp. Fox 1971
Fox 1971
Albertatherium primus n. sp. Fox 1971
Fox 1971
Mammalia - Pediomyidae
Iqualadelphis lactea n. sp. Fox 1987
Fox 1987
UAVP 22823-30
Mammalia
Pediomys exiguus Fox 1971
Fox 1971
Mammalia - Aquiladelphidae
Aquiladelphis incus Fox 1971
Fox 1971
Aquiladelphis minor Fox 1971
Fox 1971
Mammalia - Stagodontidae
Eodelphis sp. Matthew 1916
Fox 1971
Mammalia - Picopsidae
cf. Picopsis sp. Fox 1980
Fox 1980
Picopsis pattersoni n. sp. Fox 1980
Fox 1980
Mammalia - Eutriconodonta - Triconodontidae
? Triconodontidae indet. Marsh 1887
UA 12084
Alticonodon lindoei Fox 1969
Mammalia - Multituberculata - Cimolomyidae
? Cimolomys informal sp. A
Fox 1971
? Cimolomys informal sp. B
Fox 1971
Mammalia - Multituberculata
Cimexomys magister n. sp. Fox 1971
Fox 1971
Cimexomys antiquus n. sp. Fox 1971
Fox 1971
Mammalia - Multituberculata - Neoplagiaulacidae
? Mesodma sp. Jepsen 1940
Fox 1971
Mesodma senecta Fox 1971
Fox 1971
Mammalia - Multituberculata - Cimolodontidae
Cimolodon electus Fox 1971
Fox 1971
Cimolodon similis n. sp. Fox 1971
Fox 1971
Mammalia - Spalacotheriidae
Symmetrodontoides canadensis n. sp. Fox 1976
see common names

Geography
Country:Canada State/province:Alberta
Coordinates: 49.1° North, 111.8° West (view map)
Paleocoordinates:54.4° North, 73.7° West
Basis of coordinate:based on nearby landmark
Geographic resolution:small collection
Time
Period: Cretaceous Epoch: Late Cretaceous
Stage: Santonian 10 m.y. bin: Cretaceous 6
*Epoch:Senonian
*International age/stage:Campanian
Key time interval: Late Santonian
Age range of interval: 86.3 - 83.6 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Milk River Member:Deadhorse Coulee
Stratigraphic resolution:bed
Stratigraphy comments: Upper part of formation
Lithology and environment
Primary lithology:brown,gray argillaceous sandstone
Secondary lithology: sandy "shale"
Includes fossils?Y
Includes fossils?Y
Environment:terrestrial indet.
Geology comments: The upper member [of the Milk River Formation] is composed of nonmarine, drab gray, brown, and dark gray argillaceous sandstones and sandy shale with streaks of impure lignite.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection excludes:some genera
Collection methods:bulk,sieve,field collection
Reason for describing collection:taxonomic analysis
Metadata
Also known as:UALVP MR-6
Database number:14425
Authorizer:J. Alroy, R. Benson, P. Holroyd, M. Carrano Enterer:J. Alroy, M. Carrano, E. Leckey, P. Holroyd, R. Benson
Modifier:M. Carrano Research group:vertebrate
Created:1994-04-30 00:00:00 Last modified:2015-08-13 10:03:53
Access level:the public Released:1994-04-30 00:00:00
Creative Commons license:CC BY
Reference information

Primary reference:
1514. R. C. Fox. 1976. Additions to the mammalian local fauna from the Upper Milk River Formation (Upper Cretaceous), Alberta. Canadian Journal of Earth Sciences 13:1105 [J. Alroy/J. Alroy/J. Ju]

Secondary references:

6294 J. Alroy. 2002. Synonymies and reidentifications of North American fossil mammals. [J. Alroy/J. Alroy/M. Kouvari]
64013 D. B. Brinkman, A. G. Neuman, and J. D. Divay. 2017. Non-marine fishes of the late Santonian Milk River Formation of Alberta, Canada – evidence from vertebrate microfossil localities. Vertebrate Anatomy Morphology Paleontology 3:7-46 [P. Holroyd/P. Holroyd/M. Carrano]
1510 R. C. Fox. 1971. Early Campanian multituberculates (Mammalia: Allotheria) from the upper Milk River Formation, Alberta. Canadian Journal of Earth Sciences 8:916-938 [J. Alroy/J. Alroy/M. Carrano]
1511 R. C. Fox. 1971. Marsupial mammals from the early Campanian Milk River Formation, Alberta, Canada. Zoological Journal of the Linnean Society 50(suppl. 1):145-164 [J. Alroy/J. Alroy/M. Carrano]
1512 R. C. Fox. 1972. An Upper Cretaceous Symmetrodont from Alberta, Canada. Nature 239:170-171 [J. Alroy/J. Alroy/J. Ju]
9537 R. C. Fox. 1975. Fossil snakes from the Upper Milk River Formation (Upper Cretaceous), Alberta. Canadian Journal of Earth Sciences 12:1557-1563 [J. Alroy/E. Leckey/P. Mannion]
1521 R. C. Fox. 1980. Picopsis pattersoni, n.gen. and sp., an unusual therian from the Upper Cretaceous of Alberta, and the classification of primitive tribosphenic mammals. Canadian Journal of Earth Sciences 17:1489 [J. Alroy/J. Alroy/J. Ju]
1533 R. C. Fox. 1987. An ancestral marsupial and its implications for early marsupial evolution. Occasional Paper of the Tyrrell Museum of Palaeontology 3:236-240 [J. Alroy/J. Alroy/E. Leckey]
39274 R. C. Fox and B. G. Naylor. 1982. A reconsideration of the relationships of the fossil amphibian Albanerpeton. Canadian Journal of Earth Sciences 19:118-128 [M. Carrano/M. Carrano]
37933 K. -Q. Gao and R. C. Fox. 1996. Taxonomy and evolution of Late Cretaceous lizards (Reptilia: Squamata) from western Canada. Bulletin of the Carnegie Museum of natural History 33:1-107 [R. Benson/R. Benson]
42675 J. D. Gardner. 2000. Albanerpetontid amphibians from the Upper Cretaceous (Campanian and Maastrichtian) of North America. Geodiversitas 22(3):349-388 [M. Carrano/M. Carrano/M. Carrano]