PBDB Collection

Nikiti-2: MN 11, Greece

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Mammalia - Proboscidea - Choerolophodontidae

	Choerolophodon pentelici (Gaudry and Lartet 1856)
Mammalia - Perissodactyla - Equidae

	Hipparion sp. de Christol 1832
	"large-sized"

	Hipparion dietrichi (Wehrli 1941)

	Hipparion macedonicum Koufos 1984
Mammalia - Bovidae

	Palaeoryx cf. pallasi (Wagner 1857)	Kostopoulos 2016
	left maxilla with P2-M3, NIK-1063.

	Tragoportax aff. rugosifrons (Schlosser 1904)
	"Tragoportax aff. rugosiforns"
	= ? Miotragocerus sp. Stromer 1928	Kostopoulos 2016
	Partial skull with associate mandible, NIK- 764; anterior part of skull with palate, NIK-422, 1690; lower dentitions, NIK-1103, 1140, 1680, 1681.

	Tragoportax amalthea (Roth and Wagner 1854)	Kostopoulos 2016
	Skull, NIK-1688; partial skull, NIK-1687; frontlets of young individuals, NIK-1135, 1689; partly preserved hornless skull, NIK-1684; palate, NIK-423, 1685; right P2-M3, NIK- 1141; mandible, NIK-1181; left part of mandible, NIK-1675; lower tooth rows, NIK-425, 433, 1676, 1691.

	Gazella sp. Blaineville 1816
	= Gazella cf. capricornis	Kostopoulos 2016
	Partially preserved skull, NIK-1812; partially preserved skull with mandible, NIK-1142; frontlet, NIK-699; right horn-core, NIK-1819; basal part of left horn-core, NIK-1820; P2-M3, NIK-607, 1845. Possibly ascribed: right mandibular ramus with p4-m3, NIK-1892; left mandibular ramus with p3-m3, NIK- 1889, 1896; right mandibular ramus with p4-m3, NIK-1897.

	Gazella pilgrimi Bohlin 1935	Kostopoulos 2016
	Partially preserved skull, NIK-1811; frontlets, NIK-813, 1813, 1814, 1817; left horn-core, NIK-700, 1815; right horn-core, NIK-1816, 1818; palate, NIK-1899; right P2-M3, NIK-607; part of mandible with i1-m3 sin. and i1, p2-m3 dex., NIK- 778; right mandibular ramii, NIK-589, 1888, 1890, 1898, 1902; left mandibular ramii, NIK-13’, 590, 1891, 1893-1895, 1900, 1901.

	cf. Ouzocerus sp.
	= Palaeoreas lindermayeri Wagner 1848	Kostopoulos 2016

	Nisidorcas planicornis (Pilgrim 1939)
Mammalia - Giraffidae

	Palaeotraginae indet. Pilgrim 1911	Kostopoulos 2016
	Part of first phalanx lacking proximal epiphysis, NIK-211; proximal epiphysis of first phalanx, NIK-264 (young); second phalanges of young individuals, NIK-1916 (posterior?), NIK-1917 (anterior?).

	Palaeotragus rouenii (Gaudry 1861)	Kostopoulos 2016
	Left P2-(part)M1, NIK-1823; distal part of humerus, NIK-1810; radius of young individual, NIK-1809; metacarpal III + IV, NIK-1807; calcaneum, NIK-1912; part of cuboscaphoid, NIK-1914; proximal metatarsal III + IV, NIK-1808; first phalanx, NIK-1913; first, second and third phalanges, NIK- 1915.

	Helladotherium duvernoyi Gaudry and Lartet 1856
	"Helledotherium duvernoyi"
Reptilia - Testudines - Testudinidae

	Testudo cf. marmorum Gaudry 1862	Garcia et al. 2016
	complete shell
	= Testudo marmorum Gaudry 1862	Vlachos et al. 2019
Reptilia - Struthioniformes - Struthionidae

	Struthio cf. karatheodoris Forsyth Major 1888	Koufos et al. 2016
	tarsometatarsus
see common names

Country:	Greece
Coordinates:	40.2° North, 23.7° East (view map)
Paleocoordinates:	38.0° North, 24.5° East (Wright 2013)
Basis of coordinate:	estimated from map
Geographic resolution:	outcrop

Period:	Neogene	Epoch:	Miocene
Stage:	Tortonian	10 m.y. bin:	Cenozoic 6
Key time interval:	MN 11
Age range of interval:	8.7 - 7.75 m.y. ago

Formation:	Nikiti
Stratigraphic resolution:	bed
Stratigraphy comments: about 20 m higher than the Vallesian collection Nikiti 1 (NKT) "early Turolian... beginning of MN11"

Primary lithology:	hematitic siltstone
Lithology description: lens of "red beds"; stratigraphic section indicates fine sandstone mixed with clay, presumably meaning siltstone
Environment:	terrestrial indet.

Modes of preservation:	body
Size of fossils:	macrofossils
Associated major elements:	some
Spatial resolution:	parautochthonous

Collection methods:	field collection
Reason for describing collection:	taxonomic analysis

Also known as:	NIK
Database number:	73869
Authorizer:	J. Alroy, E. Vlachos	Enterer:	J. Alroy, P. Kampouridis, E. Vlachos
Modifier:	J. Alroy	Research group:	vertebrate
Created:	2007-07-09 11:21:58	Last modified:	2025-02-22 15:12:02
Access level:	the public	Released:	2007-07-09 11:21:58
Creative Commons license:	CC0

61581		G. Garcia, D. S. Kostopoulos, X. Valentin and G. D. Koufos. 2016. Testudines. Geobios 49:23-27 [E. Vlachos/E. Vlachos]
71311		D. S. Kostopoulos. 2016. Artiodactyla - Palaeontology of the upper Miocene vertebrate localities of Nikiti (Chalkidiki Peninsula, Macedonia, Greece). Geobios 49:119-234 [E. Vlachos/E. Vlachos/E. Vlachos]
67329		G. D. Koufos, D. S. Kostopoulos, and G. E. Konidaris. 2016. Palaeontology of the upper Miocene vertebrate localities of Nikiti (Chalkidiki Peninsula, Macedonia, Greece) Aves. Geobios 49:29-36 [E. Vlachos/P. Kampouridis]
72005		E. Vlachos, G. L. Georgalis, S. Roussiakis, M. Bohme, and G. Theodorou. 2019. The Pikermian tortoises (Testudines, Testudinidae) from the late Miocene of the South Balkans. Journal of Vertebrate Paleontology 39(5):1-23 [E. Vlachos/E. Vlachos]