Bromacker Quarry (upper level of Tambach sandstone): Artinskian, Germany
collected by Schäfer-Gotha 1887, 1893
List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Amphibia
- Temnospondyli
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Dissorophoidea informal new sp.
(Bolt 1969)
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Martens et al. 2005 | |||||||||
"family indeterminate" | ||||||||||
Amphibia
- Temnospondyli
- Trematopidae
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Trematopidae informal new sp.
(Williston 1910)
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Martens et al. 2005 | |||||||||
= Rotaryus gothae n. gen., n. sp.
Berman et al. 2011
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Berman et al. 2011 | |||||||||
MNG 10182 holotype | ||||||||||
Tambachia trogallas n. gen., n. sp.
Sumida et al. 1998
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Sumida et al. 1998 | 1 individual | ||||||||
MNG 7722 holotype | ||||||||||
Amphibia
- Temnospondyli
- Dissorophidae
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Dissorophidae informal new sp.
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Martens et al. 2005 | |||||||||
Amphibia
- Temnospondyli
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Georgenthalia clavinasica n. gen., n. sp.
Anderson et al. 2008
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Anderson et al. 2008 | 1 individual | ||||||||
MNG 11135 holotype | ||||||||||
Amphibia
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Amphisauropus sp.
Haubold 1970
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Voigt et al. 2007 | |||||||||
unclassified
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Tambachichnium sp.
Müller 1954
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Voigt et al. 2007 | |||||||||
Ichnium brachydactylum
Pabst 1908
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Pabst 1908 | 2 individuals | ||||||||
Gotha n. 1841, 1843 | ||||||||||
Brachystelechidae
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Bromerpeton subcolossus n. gen., n. sp.
MacDougall et al. 2024
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MacDougall et al. 2024 | 1 specimen | ||||||||
Holotype: MNG 16545, a partial skull and mandible with left humerus and largely complete right forelimb | ||||||||||
Sphenacodontidae
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Dimetrodon teutonis n. sp.
Berman et al. 2001
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Berman et al. 2001 | 1 individual | ||||||||
MNG 10598 holotype | ||||||||||
Varanopidae
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Varanopseidae informal new sp.
(Romer and Price 1940)
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Martens et al. 2005 | 2 specimens | ||||||||
= Tambacarnifex unguifalcatus n. gen., n. sp.
Berman et al. 2014
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Berman et al. 2014 | |||||||||
holotype: MNG 10596 (greater part of postcranial skeleton preserved both in bone and as impressions on counterpart blocks); paratype: MNG 15037 (partial left dentary exposed in lateral view) | ||||||||||
Caseidae
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Caseidae informal new sp.
Williston 1911
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Martens et al. 2005 | |||||||||
Martensius bromackerensis n. gen., n. sp.
Berman et al. 2020
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Berman et al. 2020 | 1 specimen | ||||||||
MNG 13814 (Holotype), adult and largest of the specimens, consisting of a nearly complete, well–preserved skeleton preserved in several, closely associated, articulated sections | ||||||||||
Reptilia
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Thuringothyris mahlendorffae n. gen., n. sp.
Boy and Martens 1991
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Müller et al. 2006 | 8 individuals | ||||||||
MNG 7729 holotype; MNG 10183, 10647, 10652, 11191 referred specimens | ||||||||||
Dimetropus sp.
Romer and Price 1940
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Voigt et al. 2007 | |||||||||
Ichnium acrodactylum n. sp.
Pabst 1908
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Pabst 1908 | 12 individuals | ||||||||
incl. subspecies alternans, curvata, dispar, and microdactylum; Gotha no. 1760, 1762, 1765–1766, 1823, 1825, 1827–1828, 1838, 1983, 1985–1986 | ||||||||||
Reptilia
- Bolosauridae
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Eudibamus cursoris n. gen., n. sp.
Berman et al. 2000
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Berman et al. 2000 | 1 individual | ||||||||
MNG 8852 holotype | ||||||||||
Diadectidae
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Ichniotherium cottae
(Pohlig 1885)
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Pabst 1908 | 11 specimens | ||||||||
Orobates pabsti n. gen., n. sp.
Berman et al. 2004
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Berman et al. 2004 | 4 individuals | ||||||||
MNG 10181 holotype; MNG 8760, 8980, 11133, 11134 paratypes | ||||||||||
Diadectes absitus n. sp.
Berman et al. 1998
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4 specimens | |||||||||
MNG 8853 (Holotype: nearly complete articulated skeleton), MNG 8747 (nearly complete skull), MNG 7721 (nearly complete postcranial skeleton), and MNG 8978 (nearly complete but disarticulated postcranial skeleton) | ||||||||||
unclassified
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Varanopus sp.
Moodie 1929
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Voigt et al. 2007 | |||||||||
Ichnium dolichodactylum
Pabst 1908
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Pabst 1908 | 1 individual | ||||||||
Gotha no. 1785 | ||||||||||
Seymouriidae
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Seymouria sanjuanensis
Vaughn 1966
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Berman et al. 2000 | 4 individuals | ||||||||
MNG 7727, 8759, 10553, 10554 | ||||||||||
Myriapoda
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Myriapoda sp.
(Latreille 1802)
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Martens et al. 2005 | |||||||||
"family indeterminate, 1-2 undescribed new species" | ||||||||||
Insecta
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Insecta sp.
Linnaeus 1758
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Martens et al. 2005 | |||||||||
"family indeterminate, 2-3 undescribed new species" | ||||||||||
Insecta
- Blattodea
- Mylacridae
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Opsiomylacris sp.
Haupt 1952
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Martens et al. 2005 | |||||||||
Moravamylacris kukalovae
Schneider 1980
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Martens et al. 2005 | |||||||||
Insecta
- Phyloblattidae
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Kunguroblattina sp.
Martynov 1930
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Martens et al. 2005 | |||||||||
Phylloblatta sp.
(Handlirsch 1906)
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Martens et al. 2005 | |||||||||
cf. Anthracoblattina sp.
Scudder 1879
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Martens et al. 2005 | |||||||||
Branchiopoda
- Lioestheriidae
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Lioestheria monticula
Martens 1983
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Martens et al. 2005 | |||||||||
unclassified
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Ichnium sphaerodactylum n. sp.
Pabst 1908
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Pabst 1908 | 42 specimens | ||||||||
Gotha no. 1351–1352, 1367–1371, 1393–1395, 1401 2, 1747–1749, 1814, 1819, 1840 | ||||||||||
see common names |
Geography
Country: | Germany | State/province: | Thuringia | County: | Gotha |
Coordinates: | 50.8° North, 10.6° East (view map) | ||||
Paleocoordinates: | 8.9° North, 22.6° East | ||||
Basis of coordinate: | based on nearby landmark | ||||
Geographic resolution: | outcrop |
Time
Period: | Permian | Epoch: | Cisuralian |
Stage: | Artinskian | 10 m.y. bin: | Permian 2 |
Key time interval: | Artinskian | ||
Age range of interval: | 290.1 - 283.5 m.y. ago |
Stratigraphy
Geological group: | Rotliegend | Formation: | Tambach | Member: | Tambach-Sandstein |
Stratigraphic resolution: | group of beds | ||||
Stratigraphy comments: "At the Bromacker quarry site are two superimposed stratigraphic successions that can be characterized by their facies associations and are referred to as the Lower Beds and Upper Beds (Eberth et al. 2000). All the vertebrates from the Bromacker quarry come from the Upper Beds [...]. The vertebrates are almost exclusively restricted to two [... horizons ...] which are separated by 50 cm in a stratigraphic interval of 1.2 m." (Berman et al., 2001)
The Tambach Formation is of Artinskian age according to "Stratigraphische Tabelle von Deutschland 2002", however, a Kungurian age is proposed by Lützner (2006) in LITHOLEX stratigraphic online database [http://www.bgr.de/app/LithoLex/gesamt_ausgabe_neu.php5?id=25] |
Lithology and environment
Primary lithology: | very fine,brown,red lithified sandstone | ||
Includes fossils? | Y | ||
Lithology description: "massive, red-brown, very fine-grained sandstone and siltstone" (Berman et al., 2001). For further details see Eberth et al. (2000). | |||
Environment: | fluvial-lacustrine indet. | Tectonic setting: | intramontane basin |
Geology comments: "alluvial paleochannel and sheetflood facies and lacustrine suspension deposits" (Berman et al., 2001).
"During the time that the Lower and Upper beds were deposited, the Tambach Basin was internally-drained and probably characterized by a wet-and-dry tropical or wetter climate that was hot year-round with seasonal-to-subseasonal cycles of rainfall and drying. [...] In the Bromacker area, the floodbasin was never deeply desiccated or heavily vegetated between flooding events." Eberth et al. (2000). |
Taphonomy
Modes of preservation: | body,permineralized |
Size of fossils: | macrofossils |
Preservation of anatomical detail: | excellent |
Articulated whole bodies: | some |
Associated major elements: | some |
Disassociated major elements: | some |
Disassociated minor elements: | some |
Temporal resolution: | snapshot |
Collection methods and comments
Collection excludes: | all macrofossils | ||
Collection methods: | bulk,surface (in situ),field collection | ||
Reason for describing collection: | taxonomic analysis | ||
Collectors: | Schäfer-Gotha | Collection dates: | 1887, 1893 |
Collection method comments: MNG = Museum der Natur, Gotha |
Metadata
Database number: | 80112 | ||
Authorizer: | R. Reisz, E. Dunne, A. Dunhill, J. Mueller, M. Carrano | Enterer: | R. Reisz, M. Carrano, E. Dunne, T. Liebrecht, B. Allen |
Modifier: | M. Carrano | Research group: | vertebrate |
Created: | 2008-04-04 13:49:44 | Last modified: | 2021-12-07 11:20:18 |
Access level: | authorizer only | Released: | 2011-04-04 13:49:44 |
Creative Commons license: | CC BY |
Reference information
Primary reference:
26922. | D.S. Berman, S.S. Sumida, and T. Martens. 1998. Diadectes (Diadectomorpha: Diadectidae) from the Early Permian of Central Germany, with description of a new species. Annals of Carnegie Museum 67(1):53-93 [R. Reisz/D. Mazierski] |
Secondary references:
27263 | ETE | J. S. Anderson, A. C. Henrici, S. S. Sumida, T. Martens, and D. S. Berman. 2008. Georgenthalia clavinasica, a new genus and species of dissorophoid temnospondyl from the Early Permian of Germany, and the relationships of the family Amphibamidae. Journal of Vertebrate Paleontology 28(1):61-75 [M. Carrano/M. Carrano] |
5765 | ETE 1524 | D. S. Berman, A. C. Henrici, S. S. Sumida and T. Martens. 2000. Redescription of Seymouria sanjuanensis (Seymouriomorpha) from the Lower Permian of Germany based on complete, mature specimens with a discussion of paleoecology of Bromacker locality assemblage. Journal of Vertebrate Paleontology 20(2):253-268 [A. Behrensmeyer/A. Behrensmeyer/J. Alroy] |
30738 | D. S. Berman, A. C. Henrici, R. A. Kissel, S. S. Sumida, and T. Martens. 2004. A New Diadectid (Diadectomorpha), Orobates pabsti, from the Early Permian of Central Germany. Bulletin of Carnegie Museum of Natural History 35(1):1-36 [J. Mueller/T. Liebrecht] | |
40304 | D. S. Berman, A. C. Henrici, T. Martens, S. S. Sumida, and J. S. Anderson. 2011. Rotaryus gothae, a New Trematopid (Temnospondyli: Dissorophoidea) from the Lower Permian of Central Germany. Annals of Carnegie Museum 80(1):49-65 [J. Mueller/T. Liebrecht] | |
50796 | D. S. Berman, A. C. Henrici, S. S. Sumida, T. Martens, and V. Pelletier. 2014. First European record of a Varanodontine (Synapsida: Varanopidae): member of a unique Early Permian upland paleoecosystem, Tambach Basin, central Germany. In C. F. Kammerer, K. D. Angielczyk, J. Fröbsich (eds.), Early Evolutionary History of the Synapsida 69-86 [R. Benson/R. Benson] | |
72029 | D. S. Berman, H. C. Maddin, A. C. Henrici, S. S. Sumida, D. Scott and R. R. Reisz. 2020. New primitive caseid (Synapsida, Caseasauria) from the early Permian of Germany. Annals of Carnegie Museum 86(1):43-75 [E. Dunne/E. Dunne] | |
28679 | D. S. Berman, R. R. Reisz, D. Scott, A. C. Henrici, S. S. Sumida and T. Martens. 2000. Early Permian Bipedal Reptile. Science 290:969-972 [J. Mueller/T. Liebrecht] | |
28661 | D. S. Berman, R. R. Reisz, T. Martens and A. C. Henrici. 2001. A new species of Dimetrodon (Synapsida: Sphenacodontidae) from the Lower Permian of Germany records first occurrence of genus outside of North America. Canadian Journal of Earth Sciences 38(5):803-812 [J. Mueller/T. Liebrecht] | |
5766 | ETE 1525 | D. A. Eberth, D. S. Berman, S.S. Sumida and H. Hopf. 2000. Lower Permian Terrestrial Paleoenvironments and Vertebrate Paleoecology of the Tambach Basin (Thuringia, Central Germany): The Upland Holy Grail. Palaios 15(4):293-313 [A. Behrensmeyer/A. Behrensmeyer/T. Liebrecht] |
87733 | M. J. MacDougall, A. Jannel, A. C. Henrici, D. S. Berman, S. S. Sumida, T. Martens, N. B. Fröbisch and J. Fröbisch. 2024. A new recumbirostran ‘microsaur’ from the lower Permian Bromacker locality, Thuringia, Germany, and its fossorial adaptations. Scientific Reports 14(4200) [E. Dunne/E. Dunne] | |
40162 | T. Martens, D. S. Berman, A. C. Henrici and S. S. Sumida. 2005. The Bromacker Quarry - the Most Important Locality of Lower Permian Terrestrial Vertebrate Fossils Outside of North America. The Nonmarine Permian. New Mexico Museum of Natural History and Science Bulletin 30:214-215 [J. Mueller/T. Liebrecht] | |
28678 | J. Müller, D. S. Berman, A. C. Henrici, T. Martens, and S. S. Sumida. 2006. The Basal Reptile Thuringothyris mahlendorffae (Amniota: Eureptilia) from the Lower Permian of Germany. Journal of Paleontology 80(4):726-739 [J. Mueller/T. Liebrecht] | |
78836 | W. Pabst. 1908. Die Tierfährten in dem Rotliegenden “Deutschlands” [The animal tracks in the “German” Rotliegenden]. Nova Acta. Abhandlungen der Kaiserlichen Leopoldinisch-Carolinischen Deutschen Akademie der Naturforscher 89(2):315-480 [M. Carrano/M. Carrano] | |
30596 | S. S. Sumida, D. S. Berman, and T. Martens. 1998. A New Trematopid Amphibian from the Lower Permian of Central Germany. Palaeontology 41(4):605-629 [J. Mueller/T. Liebrecht] | |
67270 | S. Voigt, D. S. Berman, and A. C. Henrici. 2007. First well-established track-trackmaker association of paleozoic tetrapods based on Ichniotherium trackways and diadectid skeletons from the Lower Permian of Germany. Journal of Vertebrate Paleontology 27(3):553-570 [A. Dunhill/B. Allen] |