PBDB Collection

Czatkowice 1 quarry: Olenekian, Poland
collected 1978

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Osteichthyes

	Osteichthyes indet. Huxley 1880
Amphibia - Temnospondyli

	Czatkobatrachus polonicus n. gen., n. sp. Evans and Borsuk-Bialynicka 1998	Evans and Borsuk-Bialynicka 1998	76 elements
Amphibia - Temnospondyli - Tupilakosauridae

	Batrachosuchoides sp. Shishkin 1967	Shishkin and Sulej 2009	8 elements
	A series of incomplete isolated bones, collection ZPAL AbIV: postparietal 101; ectoptery− goids 36, 53; exoccipitals 48, 120, 152; surangular 69; clavicle 61
Amphibia - Temnospondyli

	Amphibia indet. Linnaeus 1758		53 elements
	= Parotosuchus speleus n. sp. Shishkin and Sulej 2009	Shishkin and Sulej 2009
	ZPAL AbIV/105, left vomer (holotype). A series of isolated bones and bone fragments from the type locality, all belonging to collection ZPAL AbIV. The specimens include: postfrontal 43; postorbital 151; vomers 37, 38, 58, 59, 65–67, and 73; palatine 97; ectopterygoids 46, 50, and 107; exoccipitals 33 and 104; premaxillae 62 and 92; maxillae 31,106,116,118, and 153; maxillae or dentaries (poorly identifiable fragments) 47, 51, 52, 57, 60, 64, 74, 91, and 93; dentaries 32, 34, 35, 82, and 119; angular 63; surangular 68; clavicles 75 and 87; interclavicle 102; neural arches 158–160; scapulae (?) 88 and 103; humeri 84 and 86; radius (?) 96; ribs 85 and 99.
Reptilia

	Lepidosauromorpha indet. Benton 1983		84 elements
	= Sophineta cracoviensis n. gen., n. sp. Evans and Borsuk−Białynicka 2009	Evans and Borsuk−Białynicka 2009
	ZPAL RV/175, a nearly complete right maxilla (holotype). ZPAL RV/226–230, 445, 1086 (frontals); ZPAL RV/231, 232, 443 (parietals); ZPAL RV/1056 (prefrontal); ZPAL RV/233–235, 966, 967, 1060 (postfrontals); ZPAL RV/236–238, 1058, 1062 (postorbitals); ZPAL RV/10, 239–244, 965, 1054, 1059 (jugals); ZPAL RV/245–249, 1055, 1057, 1069 (squamosals); ZPAL RV/968–973 (pterygoids); ZPAL RV/13, and 974 (quadrates); ZPAL RV/3, 7, 174–178, 455, 472, 493, 823, 824 (maxillae); ZPAL RV/1061 (premaxilla); ZPAL RV/174, 189, 392, 493, 512, 1101 (dentaries); ZPAL RV/23, 627, 628, 1079, 1080, 1098, 1108, 1110, 1121 (vertebrae); ZPAL RV/948–950, 952, 959, 1053, 1063 (ilia).
Reptilia - Kuehneosauridae

	Diapsida indet. Osborn 1903		114 elements
	= Pamelina polonica n. gen., n. sp. Evans 2009	Evans 2009
	ZPAL RV/1036 (maxilla, holotype). ZPAL RV/378, 381, 383–384, 387, 537, 1087 (frontals); ZPAL RV/157, 975–978, 1028 (parietals); ZPAL RV/ 979, 980, 1001, 1002 (prefrontals); ZPAL RV/148, 1003, 1004, 1006, 1007, 1027 (postfrontals); ZPAL RV/806, 1005, 1072, 1077, 1078 (postorbitals); ZPAL RV/1008–1010 (jugals); ZPAL RV/147, 366, 1011–1026 (squamosals); ZPAL RV/1029–1033, 1083 (quadrates); ZPAL RV/1, 6, 1034–1042, 1081 (maxillae); ZPAL RV/146, 151–155, 184, 201, 451, 1043–1045, 1082 (premaxillae); ZPAL RV/1048–1050 (sphenoids); ZPAL RV/142–144, 149, 162, 185, 186, 1046, 1047 (dentaries); ZPALRV/1066, 67 (possible palatines); ZPAL RV/ 613, 617, 627, 1194–1210 (vertebrae); ZPAL RV/1211–1213 (ribs); ZPAL RV/555, 981 (ilia).
Reptilia

	Archosauriformes indet. Gauthier 1986	Borsuk−Białynicka and Sennikov 2009
Reptilia - Euparkeriidae

	Archosauria indet. Cope 1869		260 specimens
	= Osmolskina czatkowicensis n. gen., n. sp. Borsuk-Bialynicka and Evans 2003	Borsuk-Bialynicka and Evans 2003
	Approximately 100 cranial elements. Catalogued postcranial specimens are: 63 measured vertebrae (Supplement 1); twelve scapulae: ZPAL RV/881–885, 902, 1161, 1261, 1282–1286; eight coracoids: ZPAL RV/890–891, 903, 1166–1169, 1311; eleven humeri ZPAL RV/1164, 1262–1263, 1266–1272, 1277; ten ilia: ZPAL RV/630, 678, 679, 911–913, 918, 919, 924, 925; four ischia: ZPAL RV/887–889, 892; five pubes: 905–907, 909, 910; fifteen femora: ZPAL RV/938–940, 1252, 1254, 1264, 1287–1302, 1304; 18 tibiae: ZPAL RV/1165, 1171, 1176, 1177, 1221, 1222, 1257–1260, 1265, 1303, 1306, 1317–1321, fragmentary radius ZPAL RV/1232 and ulnae ZPAL RV/1230 and 1279; fragmentary fibulae ZPAL RV/1182, 1225, 1247; numerous tarsal bones, of which the astragalus: ZPAL RV/811, a calcaneum ZPAL RV/810, and a fourth distal tarsal ZPAL RV/812 are catalogued and figured.
Reptilia

	Collilongus rarus n. gen., n. sp. Borsuk−Białynicka and Sennikov 2009	Borsuk−Białynicka and Sennikov 2009	20 elements
	Four cervicals: ZPAL RV/579, 580, 581, 596; ?four dorsals: ZPAL RV/584, 585, 588, 694; one sacral ZPAL RV/1369; and twelve caudals: ZPAL RV/583, 594, 661, 662, 663, 1362, 1363, 1364, 1365, 1366, 1367, 1368

	Prolacertiformes indet. Camp 1945
	= Czatkowiella harae n. gen., n. sp. Borsuk−Białynicka and Evans 2009	Borsuk−Białynicka and Evans 2009
	ZPAL R.V/100, right maxilla. Many referred specimens
Reptilia - Procolophonidae

	Procolophonidae informal unnamed genus 1	Borsuk−Białynicka and Lubka 2009
	Maxillae ZPAL RV/734; ZPAL RV/739; dentary: ZPAL RV/733; vertebrae type B2 (Lubka 1999): ZPAL RV/840–842, 849, 851–853, 860, and numerous unnumbered specimens.

	Procolophonidae informal unnamed genus 2	Borsuk−Białynicka and Lubka 2009
	Dentary fragment ZPAL RV/507; Vertebrae type B3 (Lubka 1999): ZPAL RV/767–772; ZPAL RV/846–870, and several unnumbered vertebrae

	Procolophonidae informal unnamed genus 3	Borsuk−Białynicka and Lubka 2009	1 element
	ZPAL RV/716, dentary

	Procolophonida indet.
	= Procolina teresae n. gen., n. sp. Borsuk−Białynicka and Lubka 2009	Borsuk−Białynicka and Lubka 2009
	Many disarticulated premaxillae, maxillae, dentaries, palatal bones and quadrates, vertebrae referred to as type B1 vertebrae by Lubka (1999), and numerous bone fragments
Dipnoi - Gnathorhizidae

	Gnathorhiza sp. Cope 1883	Borsuk-Bialynicka et al. 2003
see common names

Geography

Country:	Poland	State/province:	Malopolskie	County:	Krakowski
Coordinates:	50.1° North, 19.6° East (view map)
Paleocoordinates:	17.8° North, 26.3° East
Basis of coordinate:	based on nearby landmark
Altitude:	318 meters
Geographic resolution:	outcrop

Time

Period:	Triassic	Epoch:	Early Triassic
Stage:	Olenekian	10 m.y. bin:	Triassic 1
Key time interval:	Olenekian
Age range of interval:	251.2 - 247.2 m.y. ago

Stratigraphy

Stratigraphic resolution:	bed
Stratigraphy comments: earliest Late Olenekian (Yarenskian) on the basis of tetrapod (temnospondyl) biostratigraphy (Shishkin and Sulej 2009)

Lithology and environment

Primary lithology:	fine,yellow sandy "limestone"
Secondary lithology:	"limestone"
Includes fossils?	Y
Lithology description: "fine grained sandy yellow limestone with some calcite cement [with] bones, occasional clasts, and concretions. A few samples show evidence of sedimentary bedding and parallel orientation of the bones, but this is not generally the case." bone breccia within a Carboniferous limestone
Environment:	fissure fill
Geology comments: single karst fissure; arid environment with sparse vegetation and occasional pools with aquatic communities

Taphonomy

Modes of preservation:	body
Lagerst�tten type:	concentrate
Degree of concentration:	-multiple events
Size of fossils:	macrofossils,mesofossils
Spatial orientation:	random
Preservation of anatomical detail:	excellent
Articulated whole bodies:	none
Associated major elements:	none
Disassociated major elements:	many
Disassociated minor elements:	many
Size sorting:	medium
Fragmentation:	frequent
Temporal resolution:	time-averaged
Spatial resolution:	allochthonous

Collection methods and comments

Collection methods:	selective quarrying,chemical,mechanical,field collection
Reason for describing collection:	general faunal/floral analysis
Collection dates:	1978
Collection method comments: ZPAL: Institute of Paleobiology, Warsaw

Metadata

Database number:	38528
Authorizer:	M. Carrano, R. Butler	Enterer:	M. Carrano, R. Butler
Modifier:	M. Carrano	Research group:	vertebrate
Created:	2004-04-23 10:17:39	Last modified:	2021-10-19 10:57:09
Access level:	the public	Released:	2004-04-23 10:17:39
Creative Commons license:	CC BY

Reference information

Primary reference:

10334.

M. Borsuk-Bialynicka, E. Cook, S. E. Evans and T. Maryanska. 1999. A microvertebrate assemblage from the Early Triassic of Poland. Acta Palaeontologica Polonica 44(2):167-188 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

38012		M. Borsuk−Białynicka and S. E. Evans. 2009. A long−necked archosauromorph from the Early Triassic of Poland. Palaeontologia Polonica 65:203-234 [R. Butler/R. Butler]
38013		M. Borsuk−Białynicka and S. E. Evans. 2009. Cranial and mandibular osteology of the Early Triassic archosauriform Osmolskina czatkowicensis from Poland. Palaeontologia Polonica 65:235-281 [R. Butler/R. Butler]
38010		M. Borsuk−Białynicka and M. Lubka. 2009. Procolophonids from the Early Triassic of Poland. Palaeontologia Polonica 65:107-144 [R. Butler/R. Butler]
38014		M. Borsuk−Białynicka and A. G. Sennikov. 2009. Archosauriform postcranial remains from the Early Triassic karst deposits of southern Poland. Palaeontologia Polonica 65:283-328 [R. Butler/R. Butler]
10333		M. Borsuk-Bialynicka and S. E. Evans. 2003. A basal archosauriform from the Early Triassic of Poland. Acta Palaeontologica Polonica 48(4):649-652 [M. Carrano/M. Carrano/M. Carrano]
10336		M. Borsuk-Bialynicka, T. Maryanska, and M. A. Shishkin. 2003. New data on the age of the bone breccia from the locality Czatkowice 1 (Cracow Upland, Poland). Acta Palaeontologica Polonica 48(1):153-155 [M. Carrano/M. Carrano/M. Carrano]
38007		E. Cook and C. Trueman. 2009. Taphonomy and geochemistry of a vertebrate microremains assemblage from the Early Triassic karst deposits at Czatkowice 1, southern Poland. Palaeontologia Polonica 65:17-30 [R. Butler/R. Butler]
38011		S. E. Evans. 2009. An early kuehneosaurid reptile (Reptilia: Diapsida) from the Early Triassic of Poland. Palaeontologia Polonica 65:145-178 [R. Butler/R. Butler]
38009		S. E. Evans and M. Borsuk−Białynicka. 2009. The Early Triassic stem−frog Czatkobatrachus from Poland. Palaeontologica Polonica 65:79-105 [R. Butler/R. Butler]
35304		S. E. Evans and M. Borsuk−Białynicka. 2009. A small lepidosauromorph reptile from the Early Triassic of Poland. Palaeontologia Polonica 65:179-202 [R. Butler/R. Butler]
10335		S. E. Evans and M. Borsuk-Bialynicka. 1998. A stem-group frog from the Early Triassic of Poland. Acta Palaeontologica Polonica 43(4):573-580 [M. Carrano/M. Carrano/M. Carrano]
78367		M. Paszkowski. 2009. The Early Triassic karst of Czatkowice 1, southern Poland. Palaeontologia Polonica 65:7-16 [M. Carrano/M. Carrano]
38008		M. A. Shishkin and T. Sulej. 2009. The Early Triassic temnospondyls of the Czatkowice 1 tetrapod assemblage. Palaeontologia Polonica 65:31-77 [R. Butler/R. Butler]