Daohugou (CNU 2012 collection): Callovian - Oxfordian, China

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Insecta - Hymenoptera - Pamphiliidae
Scabolyda orientalis n. gen., n. sp. Wang et al. 2014
Wang et al. 2014 3 specimens
CNU-HYM-NN2012108, 109, 110 (5 measurements)
Insecta - Hymenoptera - Xyelydidae
Ferganolyda insolita n. sp. Wang et al. 2015
Wang et al. 2015 2 specimens
CNU-HYM-NN2012136, 137 (4 measurements)
Ferganolyda eucalla n. sp. Wang et al. 2015
Wang et al. 2015 2 specimens
CNU-HYM-NN2012132, 133 (4 measurements)
Ferganolyda chungkuei Rasnitsyn et al. 2006
Wang et al. 2015 1 specimen
CNU-HYM-NN2012135 (1 measurement)
Ferganolyda scylla Rasnitsyn et al. 2006
Wang et al. 2015 1 specimen
CNU-HYM-NN2012134 (2 measurements)
Brevilyda provecta n. gen., n. sp. Wang and Rasnitsyn 2016
Wang et al. 2016 3 specimens
CNU-HYM-NN-2012151, 149, 150 (2 measurements)
Strenolyda retrorsa n. sp. Wang and Rasnitsyn 2016
Wang et al. 2016 4 specimens
CNU-HYM-NN-2012164, 167, 158, 163 (2 measurements)
Strenolyda marginalis n. gen., n. sp. Wang and Rasnitsyn 2016
Wang et al. 2016 3 specimens
CNU-HYM-NN-2012166, 165, 170 (2 measurements)
Medilyda distorta n. sp. Wang and Rasnitsyn 2016
Wang et al. 2016 1 specimen
CNU-HYM-NN-2012171 (2 measurements)
Medilyda procera n. gen., n. sp. Wang and Rasnitsyn 2016
Wang et al. 2016 2 specimens
CNU-HYM-NN-2012143, 144 (2 measurements)
Insecta - Hymenoptera
Prolyda elegantula n. sp. Wang et al. 2016
Wang et al. 2016 1 specimen
CNU-HYM-NN-2012148 (2 measurements)
Prolyda dimidia n. sp. Wang et al. 2016
Wang et al. 2016 1 specimen
CNU-HYM-NN-2012147 (2 measurements)
Insecta - Hymenoptera - Mirolydidae
Mirolyda hirta n. gen., n. sp. Wang et al. 2017
Wang et al. 2017 2 specimens
CNU-HYM-NN2012103, 171 (2 measurements)
Insecta - Hymenoptera - Megalodontesidae
Brevisiricius partialis n. gen., n. sp. Wang et al. 2015
Wang et al. 2016 1 specimen
CNU-HYM-NN-2012131 (2 measurements)
Limbisiricius complanatus n. sp. Wang et al. 2015
Wang et al. 2016 1 specimen
CNU-HYM-NN-2012130 (2 measurements)
Limbisiricius aequalis n. gen., n. sp. Wang et al. 2015
Wang et al. 2016 1 specimen
CNU-HYM-NN-2012129 (2 measurements)
Insecta - Hymenoptera - Heloridae
Archaeohelorus tensus n. sp. Shi et al. 2014
Shi et al. 2014 1 specimen
CNU-HYM-NN2012056 (2 measurements)
Archaeohelorus polyneurus n. sp. Shi et al. 2014
Shi et al. 2014 1 specimen
CNU-HYM-NN2012052 (1 measurement)
Insecta - Hymenoptera - Praeaulacidae
Archaulacus probus n. gen., n. sp. Li et al. 2014
Li et al. 2014 1 specimen
CNU-HYM-NN2012033 (2 measurements)
Praeaulacus obtutus n. sp. Li et al. 2014
Li and Shih 2015 1 specimen
CNU-HYM-NN2012039 (2 measurements)
Praeaulacus byssinus n. sp. Wang et al. 2015
Wang et al. 2015 1 specimen
CNU-HYM-NN-2012038 (2 measurements)
Praeaulacus tenellus n. sp. Li et al. 2014
Li et al. 2014 1 specimen
CNU-HYM-NN2012035 (2 measurements)
Praeaulacus subrhombeus n. sp. Li et al. 2014
Li et al. 2014 1 specimen
CNU-HYM-NN2012034 (2 measurements)
Nevania retenta Zhang and Rasnitsyn 2007
Li et al. 2014 1 specimen
CNU-HYM-NN2012001 (2 measurements)
Nevania aspectabilis n. sp. Li et al. 2014
Li et al. 2014 3 specimens
CNU-HYM-NN2012005, 012, 013 (4 measurements)
Nevania robusta Zhang and Rasnitsyn 2007
Li et al. 2014 2 specimens
CNU-HYM-NN2012002, 015 (4 measurements)
Nevania perbella n. sp. Li et al. 2014
Li et al. 2014 4 specimens
CNU-HYM-NN2012003, 010, 014, 018 (7 measurements)
Insecta - Hymenoptera - Anomopterellidae
Synaphopterella patula n. gen., n. sp. Li et al. 2013
Li et al. 2013 1 specimen
CNU-HYM-NN2012019 (2 measurements)
Anomopterella sp. Rasnitsyn 1975
Li et al. 2013 1 specimen
CNU-HYM-NN2012029 (2 measurements)
Anomopterella informal sp. 1
Li et al. 2014 1 specimen
CNU-HYM-NN2012029 (listed as A. coalita in Li et al., 2013)
Anomopterella informal sp. 2
Li et al. 2014 1 specimen
CNU-HYM-NN2012046
Anomopterella informal sp. 3
Li et al. 2014 1 specimen
CNU-HYM-NN2012048
Anomopterella informal sp. 4
Li et al. 2014 1 specimen
CNU-HYM-NN2012049
Anomopterella brachystelis n. sp. Li et al. 2013
Li et al. 2013 2 specimens
CNU-HYM-NN2012020, 025 (4 measurements)
Anomopterella pygmea n. sp. Li et al. 2014
Li et al. 2014 1 specimen
CNU-HYM-NN2012045 (2 measurements)
Anomopterella brevis n. sp. Li et al. 2014
Li et al. 2014 1 specimen
CNU-HYM-NN2012043 (2 measurements)
Anomopterella coalita n. sp. Li et al. 2013
Li et al. 2013 3 specimens
CNU-HYM-NN2012030, 023, 028 (not 029; Li et al., 2014) (6 measurements)
Anomopterella ovalis n. sp. Li et al. 2013
Li et al. 2013 3 specimens
CNU-HYM-NN2012021, 022; 044 in Li et al., 2014 (6 measurements)
Anomopterella ampla n. sp. Li et al. 2013
Li et al. 2013 2 specimens
CNU-HYM-NN2012024; 047 in Li et al., 2014 (4 measurements)
Anomopterella divergens n. sp. Li et al. 2013
Li et al. 2013 2 specimens
CNU-HYM-NN2012027, 026 (3 measurements)
Insecta - Hymenoptera - Ephialtitidae
Praeproapocritus flexus n. sp. Li et al. 2013
Li et al. 2013 1 specimen
CNU-HYM-NN2012031 (2 measurements)
Proapocritus bialatus n. sp. Li et al. 2014
Li and Shih 2015 1 specimen
CNU-HYM-NN2012040 (2 measurements)
Proapocritus parallelus n. sp. Li et al. 2013
Li et al. 2013 1 specimen
CNU-HYM-NN2012032 (2 measurements)
Symphytopterus graciler n. sp. Wang et al. 2015
Wang et al. 2015 1 specimen
CNU-HYM-NN-2012036 (2 measurements)
Insecta - Hymenoptera - Blasticotomidae
Aethotoma aninomorpha n. gen., n. sp. Gao et al. 2016
Gao et al. 2016 1 specimen
CNU-HYM-NN2012003 (2 measurements)
Insecta - Hymenoptera - Xyelidae
Cathayxyela extensa n. gen., n. sp. Wang et al. 2014
Wang et al. 2014 1 specimen
CNU-HYM-NN2012106 (2 measurements)
Platyxyela unica n. gen., n. sp. Wang et al. 2012
Wang et al. 2012 2 specimens
CNU-HYM-NN2012100, 101 (4 measurements)
Aequixyela immensa n. gen., n. sp. Wang et al. 2014
Wang et al. 2014 1 specimen
CNU-HYM-NN2012107 (2 measurements)
Daohugoa bella n. sp. Wang et al. 2019
Wang et al. 2019 1 specimen
CNU-HYM-NN-2012173
Daohugoa longa n. sp. Wang et al. 2019
Wang et al. 2019 1 specimen
CNU-HYM-NN-2012174
Insecta - Lepidoptera
Lepidoptera indet. Linnaeus 1758
Zhang et al. 2013 1 specimen
CNU-LEP-NN2012013 (2 measurements)
Insecta - Lepidoptera - Ascololepidopterigidae
Pegolepidopteron latiala n. gen., n. sp. Zhang et al. 2013
Zhang et al. 2013 1 specimen
CNU-LEP-NN2012001 (2 measurements)
Trionolepidopteron admarginis n. gen., n. sp. Zhang et al. 2013
Zhang et al. 2013 1 specimen
CNU-LEP-NN2012012 (2 measurements)
Ascololepidopterix multinerve n. gen., n. sp. Zhang et al. 2013
Zhang et al. 2013 1 specimen
CNU-LEP-NN2012028 (2 measurements)
Insecta - Lepidoptera - Mesokristenseniidae
Kladolepidopteron parva n. sp. Zhang et al. 2013
Zhang et al. 2013 1 specimen
CNU-LEP-NN2012011 (2 measurements)
Kladolepidopteron subaequalis n. sp. Zhang et al. 2013
Zhang et al. 2013 2 specimens
CNU-LEP-NN2012020, 022 (1 measurement)
Mesokristensenia trichophora n. sp. Zhang et al. 2013
Zhang et al. 2013 1 specimen
CNU-LEP-NN2012032 (2 measurements)
Insecta - Lepidoptera - Eolepidopterigidae
Akainalepidopteron elachipteron n. gen., n. sp. Zhang et al. 2013
Zhang et al. 2013 3 specimens
CNU-LEP-NN2012024, 023, 026 (5 measurements)
Grammikolepidopteron extensus n. gen., n. sp. Zhang et al. 2013
Zhang et al. 2013 1 specimen
CNU-LEP-NN2012006 (2 measurements)
Quadruplecivena celsa n. gen., n. sp. Zhang et al. 2013
Zhang et al. 2013 1 specimen
CNU-LEP-NN2012027 (2 measurements)
Dynamilepidopteron aspinosus n. gen., n. sp. Zhang et al. 2013
Zhang et al. 2013 1 specimen
CNU-LEP-NN2012014 (2 measurements)
Petilicorpus cristatus n. gen., n. sp. Zhang et al. 2013
Zhang et al. 2013 1 specimen
CNU-LEP-NN2012007 (2 measurements)
Longcapitalis excelsus n. gen., n. sp. Zhang et al. 2013
Zhang et al. 2013 1 specimen
CNU-LEP-NN2012025 (2 measurements)
Insecta - Mecoptera
Miriholcorpa forcipata n. gen., n. sp. Wang et al. 2013
Wang et al. 2013 1 specimen
CNU-MEC-NN-2012001 (2 measurements)
Fortiholcorpa paradoxa n. gen., n. sp. Wang et al. 2013
Wang et al. 2013 1 specimen
CNU-MEC-NN-2012002 (2 measurements)
Insecta - Hemiptera - Tettigarctidae
Hirtaprosbole erromera n. gen., n. sp. Liu et al. 2016
Liu et al. 2016 3 specimens
CNU-HEM-NN2012399, 161, 227 (1 measurement)
Insecta - Hemiptera - Procercopidae
Anthoscytina perpetua n. sp. Li et al. 2013
Li et al. 2013 2 specimens
CNU-HEM-NN2012002, 003 (2 measurements)
Insecta - Hemiptera - Progonocimicidae
Cicadocoris anisomeridis n. sp. Dong et al. 2014
Dong et al. 2014 4 specimens
CNU-PRO-NN2012240, 238, 239, 250 (2 measurements)
Insecta - Odonata - Euthemistidae
Sinoeuthemis daohugouensis n. gen., n. sp. Li et al. 2013
1 specimen
CNU-ODO-NN2012004 (2 measurements)
Insecta - Diptera - Axymyiidae
Raraxymyia parallela n. gen., n. sp. Shi et al. 2013
Shi et al. 2013 2 specimens
CNU-DIP-NN2012874, 844 (2 measurements)
Raraxymyia proxima n. sp. Shi et al. 2013
Shi et al. 2013 2 specimens
CNU-DIP-NN2012191, 107 (2 measurements)
see common names

Geography
Country:China State/province:Nei Mongol
Coordinates: 41.3° North, 119.2° East (view map)
Paleocoordinates:42.9° North, 122.8° East
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period: Jurassic Epoch: Middle Jurassic - Late Jurassic
Stage: Callovian - Oxfordian 10 m.y. bin: Jurassic 5
Key time interval: Callovian - Oxfordian
Age range of interval: 165.3 - 154.8 m.y. ago
Stratigraphy
Formation:Daohugou
Stratigraphic resolution:group of beds
Stratigraphy comments: The age of the Daohugou fauna has been controversial. Liu et al. (2012) obtained U-Pb shrimp ages of 160.5 and 161.0 Myr on ash beds associated with fossils of the Daohugou fauna, suggesting a Callovian-Oxfordian age.
Lithology and environment
Primary lithology:tuffaceous lithified "shale"
Includes fossils?Y
Lithology description: "tuffaceous shale"; "freshwater shales interbedded with tuffs"
Environment:lacustrine - large Tectonic setting:rift
Geology comments: Basins characterized by flexure and early rifting (Li et al., 2004)
Taphonomy
Modes of preservation:mold/impression,adpression
Lagerst�tten type:conservation
Size of fossils:macrofossils,mesofossils
Preservation of anatomical detail:excellent
Collection methods and comments
Collection methods:bulk,selective quarrying,mechanical
Reason for describing collection:taxonomic analysis
Collection method comments: Repository: Capital Normal University, Beijing, China.
Metadata
Database number:138748
Authorizer:M. Clapham Enterer:M. Clapham
Modifier:M. Clapham Research group:paleoentomology
Created:2013-01-26 09:26:01 Last modified:2023-05-27 10:47:40
Access level:the public Released:2013-01-26 09:26:01
Creative Commons license:CC BY
Reference information

Primary reference:
44452. Y. J. Li, A. Nel, C. K. Shih, D. Ren, and H. Pang. 2013. The first euthemistid damsel-dragonfly from the Middle Jurassic of China (Odonata, Epiproctophora, Isophlebioptera). ZooKeys 261:41-50 [M. Clapham/M. Clapham]

Secondary references:

52743 Q. P. Dong, Y. Z. Yao, and D. Ren. 2014. New fossil Progonocimicidae (Hemiptera: Coleorrhyncha: Progonocimicoidea) from the Upper Mesozoic of northeastern China, with a phylogeny of Coleorrhyncha. Systematic Entomology 39:773-782 [M. Clapham/M. Clapham]
60113 T. P. Gao, C. K. Shih, M. S. Engel and D. Ren. 2016. A new xyelotomid (Hymenoptera) from the Middle Jurassic of China displaying enigmatic venational asymmetry. BMC Evolutionary Biology 16(155):1-7 [M. Clapham/M. Clapham]
49470 L. F. Li, A. P. Rasnitsyn, C. K. Shih and D. Ren. 2013. Anomopterellidae restored, with two new genera and its phylogeny in Evanioidea (Hymenoptera). PLoS One 8(12):e82587 [M. Clapham/M. Clapham]
49483 L. F. Li, C. K. Shih, and D. Ren. 2013. Two new wasps (Hymenoptera: Stephanoidea: Ephialtitidae) from the Middle Jurassic of China. Acta Geologica Sinica 87:1486-1494 [M. Clapham/M. Clapham]
49900 L. F. Li, C. K. Shih, and D. Ren. 2014. Two new species of Nevania (Hymenoptera: Evanioidea: Praeaulacidae: Nevaniinae) from the Middle Jurassic of China. Alcheringa 39:140-147 [M. Clapham/M. Clapham]
53548 L. F. Li, C. K. Shih, and D. Ren. 2014. Revision of Anomopterella Rasnitsyn, 1975 (Insecta, Hymenoptera, Anomopterellidae) with two new Middle Jurassic species from northeastern China. Geologica Carpathica 65:365-374 [M. Clapham/M. Clapham]
51749 L. F. Li, C. K. Shih, and D. Ren. 2014. New fossil Praeaulacinae wasps (Insecta: Hymenoptera: Evanioidea: Praeaulacidae) from the Middle Jurassic of China. Zootaxa 3814:432-442 [M. Clapham/M. Clapham]
54461 L. F. Li and C. K. Shih. 2015. Two new fossil wasps (Insecta: Hymenoptera: Apocrita) from northeastern China. Journal of Natural History 49:829-840 [M. Clapham/M. Clapham]
48796 S. Li, C. K. Shih, C. Wang, H. Pang, and D. Ren. 2013. Forever love: the hitherto earliest record of copulating insects from the Middle Jurassic of China. PLoS One 8(11):e78188 [M. Clapham/M. Clapham]
60278 X. H. Liu, Y. Li, Y. Z. Yao and D. Ren. 2016. A hairy-bodied tettigarctid (Hemiptera: Cicadoidea) from the latest Middle Jurassic of northeast China. Alcheringa 40:383-389 [M. Clapham/M. Clapham]
48499 G. F. Shi, Y. Zhu, C. K. Shih and D. Ren. 2013. A new axymyiid genus with two new species from the Middle Jurassic of China (Diptera: Nematocera: Axymyiidae). Acta Geologica Sinica 87:1228-1234 [M. Clapham/M. Clapham/M. Clapham]
49819 X. Q. Shi, Y. Y. Zhao, C. K. Shih and D. Ren. 2014. Two new species of Archaeohelorus (Hymenoptera, Proctotrupoidea, Heloridae) from the Middle Jurassic of China. ZooKeys 369:49-59 [M. Clapham/M. Clapham]
58300 C. Wang, C. K. Shih, A. P. Rasnitsyn and M. Wang. 2016. Two new species of Prolyda from the Middle Jurassic of China (Hymenoptera, Pamphilioidea). ZooKeys 569:71-80 [M. Clapham/M. Clapham]
52502 M. Wang, A. P. Rasnitsyn, and D. Ren. 2014. Two new fossil sawflies (Hymenoptera, Xyelidae, Xyelinae) from the Middle Jurassic of China. Acta Geologica Sinica 88:1027-1033 [M. Clapham/M. Clapham]
51882 M. Wang, A. P. Rasnitsyn, C. K. Shih and D. Ren. 2014. A new fossil genus in Pamphiliidae (Hymenoptera) from China. Alcheringa 38:391-397 [M. Clapham/M. Clapham]
54287 M. Wang, A. P. Rasnitsyn, C. K. Shih and D. Ren. 2015. New fossil records of bizarre Ferganolyda (Hymenoptera: Xyelydidae) from the Middle Jurassic of China. Alcheringa 39:99-108 [M. Clapham/M. Clapham]
57528 M. Wang, A. P. Rasnitsyn, C. K. Shih, M. J. Sharkey, and D. Ren. 2016. New fossils from China elucidating the phylogeny of Praesiricidae (Insecta: Hymenoptera). Systematic Entomology 41:41-55 [M. Clapham/M. Clapham]
59079 M. Wang, A. P. Rasnitsyn, H. Li, C. K. Shih, M. J. Sharkey and D. Ren. 2016. Phylogenetic analyses elucidate the inter-relationships of Pamphilioidea (Hymenoptera, Symphyta). Cladistics 32:239-260 [M. Clapham/M. Clapham]
61854 M. Wang, A. P. Rasnitsyn, Z. Q. Yang, C. K. Shih, H. B. Wang and D. Ren. 2017. Mirolydidae, a new family of Jurassic pamphilioid sawfly (Hymenoptera) highlighting mosaic evolution of lower Hymenoptera. Scientific Reports 7(43944):1-9 [M. Clapham/M. Clapham]
69145 M. Wang, A. P. Rasnitsyn, H. C. Zhang, C. K. Shih, and D. Ren. 2019. Revising the systematic position of the extinct family Daohugoidae (basal Hymenoptera). Journal of Systematic Palaeontology 17:1025-1035 [M. Clapham/M. Clapham/M. Clapham]
52332 M. Wang, C. K. Shih, and D. Ren. 2012. Platyxyela gen. nov. (Hymenoptera, Xyelidae, Macroxyelinae) from the Middle Jurassic of China. Zootaxa 3456:82-88 [M. Clapham/M. Clapham]
57581 M. L. Wang, L. F. Li, and C. K. Shih. 2015. New fossil wasps (Hymenoptera, Apocrita) from the Middle Jurassic of China. Insect Systematics & Evolution 46:471-484 [M. Clapham/M. Clapham]
47955 Q. Wang, C. K. Shih, and D. Ren. 2013. The earliest case of extreme sexual display with exaggerated male organs by two Middle Jurassic mecopterans. PLoS One 8(8):e71378 [M. Clapham/M. Clapham/M. Clapham]
49149 W. T. Zhang, C. K. Shih, C. C. Labandeira, J. C. Sohn, D. R. Davis, J. A. Santiago-Blay, O. Flint and D. Ren. 2013. New fossil Lepidoptera (Insecta: Amphiesmenoptera) from the Middle Jurassic Jiulongshan Formation of northeastern China. PLoS One 8(11):e79500 [M. Clapham/M. Clapham]