Biesjiespoort, Victoria West: Wuchiapingian, South Africa

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Ictiodon agilis
Gorgonopidae
Gorgonopsidae indet. Lydekker 1890
Sigogneau-Russell 1989 1 individual
BMNH R 5746, isolated braincase
Lycaenops ornatus n. gen., n. sp. Broom 1925
Sigogneau-Russell 1989 1 individual
AMNH 2240, compressed skull with lower jaw, and most of the postcranial skeleton
Cynarioides tenuis n. gen., n. sp. Broom 1925
Broom 1925 1 individual
MMK 2046, crushed skull with lower jaw
Gorgonops torvus Owen 1876
Sigogneau-Russell 1989 1 individual
BP/1/386, crushed skull with right lower jaw
unclassified
Cynariops sp. Broom 1925
Bendel et al. 2018
Cynariops robustus n. gen., n. sp. Broom 1925
Sigogneau-Russell 1989 1 individual
BMNH R 5743
Cynarioides gracilis n. sp. Broom 1930
1 individual
Sectioned skull (location of type specimen currently unknown)
Anomodontia - Cryptodontidae
Bainia peavoti n. sp. Broom 1921
Broom 1932
A complete skull
Dicynodon andrewsi n. sp. Broom 1921
Broom 1921 1 individual
MMK 4169, a strongle laterallly sheared skull missing the left squamosal and portions of both temporal arches
Dicynodon wilmanae n. sp. Broom 1928
Broom 1928 1 individual
MMK 4167, a well-preserved skull missing the right zygomatic arch
Dicynodon curtus n. sp. Broom 1921
Broom 1921 1 individual
SAM-PK-7850, a distorted skull and disarticulated lower jaw
Anomodontia
Dicynodon sollasi n. sp. Broom 1921
Broom 1921 1 individual
SAM-PK-7420, a dorsoventrally crushed complete skull and lower jaws
Dicynodon psittacops Broom 1912
Cistecephalus angusticeps n. sp. Broom 1932
Broom 1932
BMNH R 5763, a skull
Palemydops platysoma n. gen., n. sp. Broom 1921
Broom 1921
BMNH R 4955, a skull
Emydopsis longus n. sp. Broom 1921
3 individuals
Three skulls, including the holotype BMHN R. 4956-9
Storthyggognathus haughtoni n. gen., n. sp. Janensch 1952
Janensch 1952 1 individual
BMNK, a skull
Anomodontia - Pristerodontidae
Emydorhynchus formosus n. sp. Broom 1935
Broom 1935 1 individual
TM 241, a crushed skull
Emyduranus platyops n. gen., n. sp. Broom 1921
Broom 1921
BMNH R4959, a skull
Therocephalia
Hofmeyria atavus Broom 1935
Broom 1936
unclassified
Lycaenodon longiceps n. gen., n. sp. Broom 1925
Sidor 2003 1 individual
BMNH R5700, anterior two-thirds of a skull lacking lower jaws
see common names

Geography
Country:South Africa State/province:Northern Cape County:Victoria West
Coordinates: 31.4° South, 23.1° East (view map)
Paleocoordinates:61.3° South, 28.7° West
Basis of coordinate:based on nearby landmark
Geographic resolution:local area
Time
Period: Permian Epoch: Lopingian
Stage: Wuchiapingian 10 m.y. bin: Permian 4
Key time interval: Wuchiapingian Zone:  Cistecephalus Assemblage Zone
Age range of interval: 259.51 - 254.14 m.y. ago
Age estimate: 256.25 to 255.2 Ma (U/Pb)
Stratigraphy
Geological group:Beaufort
Stratigraphic resolution:group of beds
Stratigraphy comments: Direct dates for Cistecephalus Assemblage Zone from Rubidge et al., 2013
Lithology and environment
Primary lithology: not reported
Environment:terrestrial indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Reason for describing collection:taxonomic analysis
Metadata
Also known as:Biesiespoort, Nobelsfontein
Database number:174666
Authorizer:R. Butler, M. Uhen, A. Dunhill Enterer:J. Benito Moreno, B. Allen, A. Morgan, R. Butler
Modifier:R. Butler Research group:vertebrate
Created:2015-11-23 09:18:19 Last modified:2016-05-16 06:56:48
Access level:database members Released:2017-11-23 09:18:19
Creative Commons license:CC BY
Reference information

Primary reference:

27216. J. W. Kitching. 1977. The distribution of the Karroo vertebrate fauna. Memoirs of the Bernard Price Institute for Palaeontological Research 1:1-131 [H. Sims/J. Camp]

Secondary references:

78808 E.-M. Bendel, C.F. Kammerer, N. Kardjilov, V. Fernandez, and J. Fröbisch. 2018. Cranial anatomy of the gorgonopsian Cynariops robustus based on CT-reconstruction. PLoS ONE 13(11):e0207367 [M. Uhen/A. Morgan/M. Uhen]
37442 R. Broom. 1921. On some new genera and species of anomodont reptiles from the Karroo beds of South Africa. Proceedings of the Zoological Society of London 647-674 [J. Fröbisch/M. Walther/M. Uhen]
59059 R. Broom. 1925. On some carnivorous therapsids. Records of the Albany Museum 3:309-326 [R. Butler/R. Butler]
58726 R. Broom. 1928. On three new species of Dicynodon. Annals of the South African Museum 22:421-425 [R. Butler/J. Benito Moreno]
27212 R. Broom. 1932. The mammal-like reptiles of South Africa and the Origin of Mammals [H. Sims/J. Camp]
37750 R. Broom. 1935. A new genus and some new species of mammal-like reptiles. Annals of the Transvaal Museum 18(1):1-12 [J. Fröbisch/M. Walther]
67672 R. Broom. 1936. On the structure of the skull in the mammal-like reptiles of the suborder Therocephalia. Philosophical Transactions of the Royal Society of London, Series B, Biological Sciences Sciences 226(529):1-42 [A. Dunhill/B. Allen/M. Carrano]
43467 W. Janensch. 1952. Zwei neue Anomodontier mit Molaren und Crista masseterica aus den Karroo-Schichten Südafrikas [Two new anomodonts with molars and a masseteric crest from the Karroo Beds of South Africa]. Palaeontologische Zeitschrift 26(3/4):218-228 [M. Carrano/M. Carrano]
57953 C. A. Sidor. 2003. The naris and palate of Lycaenodon longiceps (Therapsida: Biarmosuchia), with comments on their early evolution in the Therapsida. Journal of Paleontology 77(5):977-984 [R. Butler/R. Butler]
57886 D. S. Sigogneau-Russell. 1989. Theriodontia I: Phthinosuchia, Biarmosuchia, Eotitanosuchia, Gorgonopsia. Encyclopedia of Paleoherpetology [R. Butler/R. Butler/R. Butler]