PBDB Collection

Berezovsk Quarry: Bathonian, Russian Federation
collected by S. Krasnolutskii et al. 2000-2005, 2010–2012

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Chondrichthyes - Hybodontiformes - Hybodontidae

	Hybodus sp. Agassiz 1834	Averianov et al. 2016
Sinamiidae

	Sinamiidae indet. Berg 1940	Averianov et al. 2016
Acipenseriformes

	Acipenseriformes indet. Berg 1940	Averianov et al. 2016
Palaeonisciformes

	Palaeonisciformes indet. Hay 1902	Skutschas 2006
Amphibia - Temnospondyli - Discoglossidae

	? Eodiscoglossus sp. Villalta 1957	Skutschas et al. 2016	1 specimen
	ZIN PH 27/144
Amphibia - Temnospondyli

	Caudata informal indet. A	Averianov et al. 2016
	Berezovsk salamander A
Amphibia - Temnospondyli - Urodela

	Kiyatriton krasnolutskii n. sp. Skutchas 2016	Averianov et al. 2016
Amphibia - Temnospondyli - Karauridae

	cf. Karauridae indet. Ivachnenko 1978	Skutschas 2006	1 specimen
Amphibia - Temnospondyli

	Caudata indet. Scopoli 1777	Lopatin and Averianov 2006	1 specimen
	= Urupia monstrosa Skutschas and Krasnolutskii 2011	Averianov et al. 2016

	Urupia monstrosa n. gen., n. sp. Skutschas and Krasnolutskii 2011	Skutschas and Krasnolutskii 2011	6 specimens
	ZIN PH 1/144 (holotype atlas), 3/144, 4/144 (dorsal vertebrae); PM TGU 200.10, ZIN PH 2/144 (anterior dentary fragments); PM TGU 200/11 (femur)

	Egoria malashichevi n. gen., n. sp. Skutschas et al. 2020	Skutschas et al. 2020	6 specimens
Reptilia - Testudines

	Testudines informal indet. 1	Averianov et al. 2016	1 specimen
	ZIN PH, no. 1/186, left costal VIII.

	Testudines informal indet. 2	Danilov et al. 2018	2 specimens
	ZIN PH, no. 2/186, left xiphiplastron fragment; and specimen ZIN PH, no. 3/186, shell fragment.

	Testudines informal indet. 3	Danilov et al. 2018	1 specimen
	ZIN PH, no. 4/186, distal part of a right humerus.
Reptilia - Testudines - Xinjiangchelyidae

	Xinjiangchelys sp. Yeh 1986	Skutschas 2006
	= Annemys sp. Sukhanov and Narmandakh 2006	Averianov et al. 2016

	Annemys sp. Sukhanov and Narmandakh 2006	Danilov et al. 2018	8 specimens
	ZIN PH, no. 431/50, costal VIII; ZIN PH, nos. 1/207 and 2/207, medial parts of costal VIII; ZIN PH, nos. 14/187 and 17/187, xiphiplastra; ZIN PH, no. 3/207, xiphiplastron fragment; ZIN PH, no. 6/185, right humerus; and ZIN PH, no. 4/207, distal part of the left humerus.
	= Annemys variabilis n. sp. Obraztsova et al. 2022	Obraztsova et al. 2022
	Holotype. ZIN PH 26/184, braincase (Fig. 3E–H). Referred material. Thousands of isolated bones and several fragmentary specimens in the following collections: ZIN PH 50, more than 2000 isolated shell bones; ZIN PH 178, more than 350 basisphenoids; ZIN PH 183, vertebrae and cervical ribs; ZIN PH 184, three braincases, and isolated skull bones (in total more than 6500 specimens); ZIN PH 185, bones of girdles and limbs; ZIN PH 187, parts of the skeleton, big shell fragments and complete plates; SRM n/n, carapace and plastron restored based on fragments of several individuals (Krasnolutskii 2006, fig. 19; Skutschas et al. 2009, foto 1, fig. 4)
Reptilia - Choristodera

	? Choristodera indet. Cope 1876	Lopatin and Averianov 2006
Reptilia - Goniopholididae

	cf. Sunosuchus sp. Young 1948	Skutschas 2006	6 specimens
	= Goniopholididae indet. Cope 1875	Kuzmin et al. 2013
	ZIN PH 1/176 (maxillary fragment); ZIN PH 2/176 (fragment of left nasal); ZIN PH 3/176 (anterior fragment of right splenial); ZIN PH 4-8/176, (teeth and 17 unnumbered teeth); ZIN PH 9/176 (fragment of the dorsal osteoderm); ZIN PH 10/176 (fragmentary ventral osteoderm)
Reptilia - Mamenchisauridae

	Titanosauriformes indet. Salgado et al. 1997	Skutschas 2006
	= Mamenchisauridae indet. Chao 1965	Averianov et al. 2016
	PM TGU 200/5-BR-308-322; SM 4557; ZIN PH 41-62/117 (teeth); SM 4551-4554, 4556; ZIN PH 39, 40/117 (caudal vertebrae)
Reptilia - Heterodontosauridae

	Heterodontosauridae indet. Romer 1966	Skutschas 2006
Reptilia

	Stegosauria indet. Marsh 1877	Skutschas 2006	2 individuals
	ZIN PH 1–2/117; SHRM various
Reptilia - Theropoda - Proceratosauridae

	cf. Dromaeosauridae indet. Colbert and Russell 1969	Skutschas 2006
	= Kileskus aristotocus Averianov et al. 2010	Averianov et al. 2016

	Kileskus aristotocus n. gen., n. sp. Averianov et al. 2010	Averianov et al. 2010
Reptilia - Pterosauria - Rhamphorhynchidae

	Pterodactyloidea indet. Plieninger 1901	Skutschas 2006
	= Rhamphorhynchidae indet. Seeley 1870	Averianov et al. 2016
Reptilia

	cf. Paramacellodidae indet. Estes 1983	Skutschas 2006
	= Scincomorpha indet. Camp 1923	Averianov et al. 2016
Docodonta - Tegotheriidae

	Hutegotherium yaomingi n. gen., n. sp. Averianov et al. 2010	Averianov et al. 2010	3 specimens
	PM TGU 200/5-BR-2; PIN 5087/6, 5087/8
Docodonta - Simpsonodontidae

	Docodonta indet. Kretzoi 1946	Lopatin and Averianov 2006	2 specimens
	PIN 5087/1, posterior fragment of a right denta− ry with alveoli for the ultimate molar, posterior alveolus for the penultimate molar and partially preserved coronoid and pseudangular processes. PM TGU 200/3−BR−1, posterior fragment of a right dentary with alveoli for the two last mo− lars and partially preserved coronoid process
	= Simpsonodon sibiricus n. sp. Averianov et al. 2010	Averianov et al. 2010
	PIN 5087/5
Docodonta

	Itatodon tatarinovi n. gen., n. sp. Lopatin and Averianov 2005	Lopatin and Averianov 2006	4 specimens
	PIN 5087/2 (holotype, isolated right lower molar), 5087/7, PM TGU 200/3-BR-7
Mammalia

	Mammalia indet. Linnaeus 1758
Mammalia - Amphitheriidae

	Amphibetulimus krasnolutskii n. gen., n. sp. Lopatin and Averianov 2007	Lopatin and Averianov 2007	7 specimens
	PIN, no. 5087/3, right dentary fragment with M3 and alveoli and roots of M1, M2, M4, and M5; PIN 5087/34, right upper molar. PIN 5087/39, left dentary fragment with p1, alveoli or roots for i1–4, double rooted canine, and p2. PIN 5087/44, right dentary fragment with alveoli for two middle molars. PIN 5087/43, right dentary fragment with alveoli for ultimate and penultimate molars. PIN 5087/40, right dentary fragment with ultimate molar and roots of the penultimate molar. PIN 5087/45, right dentary fragment with alveolus for ultimate molar
Mammalia - Dryolestidae

	Dryolestidae indet. Marsh 1879	Lopatin and Averianov 2006	11 specimens
	= Anthracolestes sergeii n. gen., n. sp. Averianov et al. 2014	Averianov et al. 2014
	Holotype—PIN 5087/22, left lower molar. Referred Specimens—PIN 5087/23, damaged right lower molar; PIN 5087/36, right lower molar missing talonid; PIN 5087/27, left dentary fragment with alveoli for i1–4, double-rooted canine, and p1–3; PIN 5087/29, right dentary fragment with alveoli for i1–4, double-rooted canine, and p1–3; PIN 5087/32, left dentary with alveoli for double-rooted canine and p1–4; PM TGU 200/3- BR-3, left dentary fragment with alveoli for four last molars; PIN 5087/24, 5087/30, and 5087/31, left dentary fragments with alveoli for three last molars; PIN 5087/33, left dentary fragment with alveoli for two middle molars.
Mammalia - Eutriconodonta

	Eutriconodonta indet. Kermack et al. 1973	Averianov et al. 2008	1 specimen
	PIN 5087/4, left upper molariform tooth
Mammalia

	Sineleutherus issedonicus n. sp. Averianov et al. 2011	Averianov et al. 2011	10 specimens
	PIN 5087/9 to 5087/18
Mammalia - Multituberculata

	Tagaria antiqua n. gen., n. sp. Averianov et al. 2020	Averianov et al. 2020
	Holotype: PIN 5087/108, right P5; LMCCE 004/10, left M2; PIN 5087/50, right M2

	Tashtykia primaeva n. gen., n. sp. Averianov et al. 2020	Averianov et al. 2020
	Holotype: PIN 5087/52, left p4; PIN 5087/25, right I2; LMCCE 010/2, right upper deciduous premolar; PIN 5087/46, right upper anterior premolar; PIN 5087/21, left P5
Tritylodontidae

	Tritylodontidae indet. Cope 1884	Lopatin and Averianov 2006	18 specimens
	= Stereognathus sp. Charlesworth 1854	Averianov et al. 2017
	ZIN PH 19/117–38/117
Dipnoi

	Dipnoi indet. Müller 1884	Lopatin and Averianov 2006
	= Ferganoceratodus sp. Nessov and Kaznyshkin 1985	Averianov et al. 2016
see common names

Geography

Country:	Russian Federation	State/province:	Krasnoyarsk	County:	Sharypovo
Coordinates:	55.7° North, 89.2° East (view map)
Paleocoordinates:	58.2° North, 86.7° East
Basis of coordinate:	stated in text
Geographic resolution:	small collection

Time

Period:	Jurassic	Epoch:	Middle Jurassic
Stage:	Bathonian	10 m.y. bin:	Jurassic 4
Key time interval:	Bathonian
Age range of interval:	168.2 - 165.3 m.y. ago

Stratigraphy

Formation:	Itat	Member:	Upper
Stratigraphic resolution:	group of beds
Stratigraphy comments: upper part of formation The age of the Upper Mem- ber of the Itat Formation is Bathonian according to the pollen complex, which consists of Cyathidites minor, Piceapollenites sp., Eboracia torosa, Quadraeculina limbata, and Classopollis sp. (Raevskaya et al. 1993)

Lithology and environment

Primary lithology:	gray claystone
Includes fossils?	Y
Lithology description: "grey clays" Fossil rich layer tops a thick coal seam
Environment:	fluvial indet.
Geology comments: By mid-Jurassic times, a low-profile alluvial plain had developed in this area that was characterized by anas- tomosing rivers and well-defined floodplains (Le Heron et al. 2008). Due to the fluvial origin of the sediments, vertebrate remains are mostly disarticulated and often water worn. However, the predominantly good preserva- tion of vertebrate remains demonstrates that transport occurred over short distances.

Taphonomy

Modes of preservation:	body
Lagerst�tten type:	concentrate
Size of fossils:	macrofossils,mesofossils,microfossils
Preservation of anatomical detail:	good
Disassociated major elements:	many
Disassociated minor elements:	many
Fragmentation:	occasional

Collection methods and comments

Collection excludes:	some genera,species names
Collection methods:	bulk,selective quarrying,surface (in situ),mechanical,sieve,field collection
Reason for describing collection:	general faunal/floral analysis
Museum repositories:	PIN
Collectors:	S. Krasnolutskii et al.	Collection dates:	2000-2005, 2010–2012
Collection method comments: Itatodon tatarinovi material housed in the Paleontological Museum of Tomsk State University, Tomsk, Russia (PM TGU), and in the Paleontological Institute of the Russian Academy of Sciences, Moscow, Russia (PIN) 2010-2012 collection by a Joint Russian-German expedition: "The newly collected mammal specimens were obtained during screen-washing of ~12.5 tons of matrix. The jaw fragments were found in the field by picking the coarse fraction of fossiliferous concentrate. The teeth were recovered during sorting of the fine fraction in the laboratory. The specimens described here are housed in the Borissiak Paleontological Institute of the Russian Academy of Sciences, Moscow (PIN). "...other specimens were collected in 2010–2012 during a joint Russian-German expedition to Berezovsk Quarry. The edentulous jaw fragments were found in the field while picking the coarse fraction of fossiliferous concentrate obtained from screenwashing of ∼12.5 tons of matrix" (Avaerianov et al. 2014)"

Metadata

Also known as:	Stegosaur Quarry; Berezovsk coal mine
Database number:	61891
Authorizer:	M. Carrano, O. Rauhut, P. Mannion, E. Vlachos, R. Benson	Enterer:	M. Carrano, J. Tennant, O. Rauhut, R. Benson, G. Varnham, P. Mannion, E. Vlachos
Modifier:	G. Varnham	Research group:	vertebrate
Created:	2006-07-06 14:56:38	Last modified:	2022-03-30 07:45:38
Access level:	the public	Released:	2006-07-06 14:56:38
Creative Commons license:	CC BY

Reference information

Primary reference:

44653.

A. O. Averianov, A. V. Lopatin, P. P. Skutschas, N. V. Matynovich, S. V. Leschinkskiy, A. S. Rezvyi, S. A. Krasnolutskii and A. V. Fayngertz. 2005. Discovery of Middle Jurassic mammals from Siberia. Acta Palaeontologica Polonica 50(4):789-797 [P. Mannion/J. Tennant]

Secondary references:

68147		A. Averianov, S. Krasnolutskii, S. Ivantsov, P. Skutschas, R. Schellhorn, J. Schultz, and T. Martin. 2019. Sauropod remains from the Middle Jurassic Itat Formation of West Siberia, Russia. Paläontologische Zeitschrift [P. Mannion/P. Mannion]
55679		A. Averianov, T. Martin, A. Lopatin and S. Krasnolutskii. 2015. Stem therian mammal Amphibetulimus from the Middle Jurassic of Siberia. Palaeontologische Zeitschrift 89:197-206 [P. Mannion/J. Tennant]
61822		A. Averianov, T. Martin, P. Skutschas, I. Danilov, J. Schultz, R. Schellhorn, E. Obraztsova, A. Lopatin, E. Sttchevskaya, I. Kuzmin, S. Krasnotutskii and S. Ivantsov. 2016. Middle Jurassic vertebrate assemblage of Berezovsk coal mine in western Siberia (Russia). Global Geology 19(4):187-204 [E. Vlachos/E. Vlachos/M. Carrano]
34709	ETE	A. O. Averianov and S. A. Krasnolutskii. 2009. Stegosaur remains from the Middle Jurassic of west Siberia. Proceedings of the Zoological Institute RAS 313(2):153-167 [M. Carrano/M. Carrano]
36119		A. O. Averianov, S. A. Krasnolutskii, and S. V. Ivantsov. 2010. A new basal coelurosaur (Dinosauria: Theropoda) from the Middle Jurassic of Siberia. Proceedings of the Zoological Institute, Russian Academy of Science 314(1):42-57 [O. Rauhut/O. Rauhut]
44655		A. O. Averianov and A. V. Lopatin. 2006. Itatodon tatarinovi (Tegotheriidae, Mammalia), a docodont from the Middle Jurassic of Western Siberia and phylogenetic analysis of Docodonta. Paleontological Journal 40(6):668-677 [P. Mannion/J. Tennant]
44847		A. O. Averianov, A. V. Lopatin, and S. A. Krasnolutskii. 2008. An amphilestid-grade eutriconodontan from the Middle Jurassic of Russia. Russian Journal of Theriology 7(1):1-4 [P. Mannion/J. Tennant/J. Tennant]
34572	ETE	A. O. Averianov, A. V. Lopatin, S. A. Krasnolutskii and S. V. Ivantsov. 2010. New docodontans from the Middle Jurassic of Siberia and reanalysis of Docodonta interrelationships. Proceedings of the Zoological Institute RAS 314(2):121-148 [M. Carrano/M. Carrano]
36940	ETE	A. O. Averianov, A. V. Lopatin, and S. A. Krasnolutskii. 2011. The first haramiyid (Mammalia, Allotheria) from the Jurassic of Russia. Doklady Biological Sciences 437:103-106 [M. Carrano/M. Carrano/M. Carrano]
86461		A. O. Averianov and A. V. Lopatin. 2023. Dinosaurs of Russia: a review of the localities. Herald of the Russian Academy of Sciences 93(4) [M. Carrano/M. Carrano/M. Carrano]
51915		A. O. Averianov, T. Martin, and A. Lopatin. 2014. The oldest dryolestid mammal from the Middle Jurassic of Siberia. Journal of Vertebrate Paleontology 34(4):924-931 [R. Benson/R. Benson]
65142		A. O. Averianov, T. Martin, A. V. Lopatin, J. A. Schultz, P. P. Skutschas, R. Schellhorn, and S. A. Krasnolutskii. 2017. A tritylodontid synapsid from the Middle Jurassic of Siberia and the taxonomy of derived tritylodontids. Journal of Vertebrate Paleontology 37(5):e1363767:1-10 [M. Carrano/M. Carrano]
81047		A. O. Averianov, T. Martin, A. V. Lopatin, J. A. Schultz, R. Schellhorn, S. Krasnolutskii, P. Skutschas and S. Ivantsov. 2020. Multituberculate mammals from the Middle Jurassic of western Siberia, Russia, and the origin of Multituberculata . Papers in Palaeontology 7(2):769-787 [P. Mannion/G. Varnham]
71292		A. O. Averianov, P. P. Skutschas, R. Schellhorn, A. V. Lopatin, P. N. Kolosov, V. V. Kolchanov, D. D. Vitenko, D. V. Grigoriev, and T. Martin. 2019. The northernmost sauropod record in the Northern Hemisphere. Lethaia 53:362-368 [P. Mannion/P. Mannion/M. Carrano]
64692		I. G. Danilov, E. M. Obraztsova, E. A. Boitsova and P. P. Skutchas. 2018. Diversity of Middle Jurassic Turtles from the Berezovsk Quarry Locality, Krasnoyarsk Region, Russia: Morphology and Histological Study. Paleontological Journal 52(1):58-70 [E. Vlachos/E. Vlachos]
49777		I. T. Kuzmin, P. P. Skutschas, O. I. Grigorieva and S. A. Krasnolutskii. 2013. Goniopholidid crocodylomorph from the Middle Jurassic Berezovsk Quarry Locality (Western Siberia, Russia). Proceedings of the Zoological Institute of the Russian Academy of Sciences 317(4):452-458 [P. Mannion/P. Mannion/P. Mannion]
45969		A. V. Lopatin and A. O. Averianov. 2005. A new docodont (Docodonta, Mammalia) from the Middle Jurassic of Siberia. General Biology 405(2):434-436 [P. Mannion/J. Tennant/J. Tennant]
17882	ETE	A. V. Lopatin and A. O. Averianov. 2006. Mesozoic mammals of Russia. 9th International Symposium on Mesozoic Terrestrial Ecosystems and Biota, Abstracts and Proceedings Volume 67-70 [M. Carrano/M. Carrano/M. Carrano]
44659		A. V. Lopatin and A. O. Averianov. 2007. The earliest Asiatic pretribosphenic mammal (Cladotheria, Amphitheriidae) from the Middle Jurassic of Siberia. Doklady Biological Sciences 417:432-434 [P. Mannion/J. Tennant]
82558		E. M. Obraztsova, S. A. Krasnolutskii, V. B. Sukhanov and I. G. Danilov. 2022. Xinjiangchelyid turtles from the Middle Jurassic of the Berezovsk coal mine (Krasnoyarsk Territory, Russia): systematics, skeletal morphology, variation, relationships and palaeobiogeographic implications. Journal of Systematic Palaeontology 20(1):1-61 [E. Vlachos/E. Vlachos/E. Vlachos]
17903	ETE	P. Skutschas. 2006. Mesozoic amphibians from Siberia, Russia. 9th International Symposium on Mesozoic Terrestrial Ecosystems and Biota, Abstracts and Proceedings Volume 123-126 [M. Carrano/M. Carrano]
78277		P. Skutschas, V. Kolchanov, S. Krasnolutskii, A. Averianov, R. Schellhorn, J. Schultz, and T. Martin. 2020. A new small-sized stem salamander from the Middle Jurassic of Western Siberia, Russia. PLoS One 15(2):e0228610 [M. Uhen/K. Eleazer]
38941		P. P. Skutschas and S. A. Krasnolutskii. 2011. A new genus and species of basal salamanders from the Middle Jurassic of western Siberia, Russia. Proceedings of the Zoological Institute of the Russian Academy of Sciences 315(2):167-175 [R. Benson/R. Benson/P. Mannion]
58264		P. P. Skutschas, T. Martin, and S. A. Krasnolutskii. 2016. First Jurassic frog from Siberia, Russia. Journal of Vertebrate Paleontology 36:e1101376:1-3 [P. Mannion/P. Mannion]