Cambay amber, Tadkeshwar lignite mine (AMNH/BSIPL collection): Ypresian, India

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Insecta - Diptera - Ceratopogonidae
Ceratopogonidae indet. Newman 1834
Stebner et al. 2017 2 specimens
AMNH Tad-615, 684
Stilobezzia sp. Kieffer 1911
Stebner et al. 2017 3 specimens
AMNH Tad-515a, 518, 853a
Eohelea indica n. sp. Stebner and Szadziewski 2017
Stebner et al. 2017 1 specimen
AMNH Tad-855 (2 measurements)
Gedanohelea gerdesorum n. sp. Stebner and Szadziewski 2017
Stebner et al. 2017 3 specimens
AMNH Tad-857, 513, 661 (2 measurements)
Serromyia sp. Meigen 1818
Stebner et al. 2017 1 specimen
AMNH Tad-647
Mantohelea sp. Szadziewski 1988
Stebner et al. 2017 1 specimen
AMNH Tad-673
Brachypogon sp. Kieffer 1899
Stebner et al. 2017 3 specimens
AMNH Tad-508, 851, 854
Camptopterohelea odora n. sp. Stebner et al. 2017
Stebner et al. 2016 1 specimen
AMNH Tad-859a (2 measurements)
Meunierohelea borkenti n. sp. Stebner and Szadziewski 2017
Stebner et al. 2017 1 specimen
AMNH Tad-516 (2 measurements)
Meunierohelea orientalis n. sp. Stebner and Szadziewski 2017
Stebner et al. 2017 1 specimen
AMNH Tad-858a (2 measurements)
Meunierohelea cambayana n. sp. Stebner and Szadziewski 2017
Stebner et al. 2017 1 specimen
AMNH Tad-507 (2 measurements)
Meunierohelea aff. cambayana Stebner and Szadziewski 2017
Stebner et al. 2017 1 specimen
AMNH Tad-519
Forcipomyia sp. Meigen 1818
Stebner et al. 2017 10 specimens
AMNH Tad-163, 511a, 511b, 565, 602, 616a, 643, 856, 860, 862a
Leptoconops sp. Skuse 1889
Stebner et al. 2017 2 specimens
AMNH Tad-506, 852a
Insecta - Diptera - Psychodidae
Sycorax longistyla n. sp. Wagner et al. 2022
Wagner et al. 2022 7 specimens
AMNH Tad 877, 64, 876, 769, 356A, 878, 880 (7 measurements)
Insecta - Diptera - Dolichopodidae
Dolichopodidae indet. Latreille 1809
Bickel et al. 2022 3 specimens
AMNH Tad-41b, 165, 321b
Medeterinae indet. Lioy 1864
Bickel et al. 2022 8 specimens
AMNH Tad-66, 271, 295, 401, 406, 447, 468, 498
Rajpardia grimaldii n. gen., n. sp. Bickel 2022
Bickel et al. 2022 1 specimen
AMNH Tad-372 (1 measurement)
Sympycninae indet. Aldrich 1905
Bickel et al. 2022 2 specimens
AMNH Tad-359, 486
Diaphorinae indet. Schiner 1864
Bickel et al. 2022 5 specimens
AMNH Tad-65, 176, 177, 378, 481
Gujaratmyia rotunda n. gen., n. sp. Bickel 2022
Bickel et al. 2022 1 specimen
AMNH Tad-67 (2 measurements)
Palaeomedeterus sp. (Meunier 1895)
Bickel et al. 2022 1 specimen
AMNH Tad-475
Palaeomedeterus cambayensis n. sp. Bickel 2022
Bickel et al. 2022 7 specimens
AMNH Tad-476B, 124, 165, 448, 476A, 483; KU 173A (2 measurements)
Insecta - Diptera - Campichoetidae
Pareuthychaeta eoindica n. sp. Grimaldi 2012
Grimaldi and Singh 2012 1 specimen
AMNH Tad-227 (2 measurements)
Insecta - Diptera - Keroplatidae
Indorrhina sahnii n. gen., n. sp. Stebner and Grimaldi 2017
Stebner et al. 2017 1 specimen
BSIP Tad-418 (2 measurements)
Palaeognoriste orientale n. sp. Stebner and Grimaldi 2017
Stebner et al. 2017 4 specimens
BSIP Tad-887, AMNH Tad-587, Tad-37A, Tad-324 (8 measurements)
Lygistorrhina indica n. sp. Stebner and Grimaldi 2017
Stebner et al. 2017 3 specimens
BSIP Tad-442, AMNH Tad-492, Tad-888 (4 measurements)
Insecta - Diptera - Chironomidae
Stempellina stebneri n. sp. Giłka and Zakrzewska 2020
Zakrzewska et al. 2020 1 specimen
BSIP Tad-882 (2 measurements)
Stempellinella pollex n. sp. Giłka and Zakrzewska 2020
Zakrzewska et al. 2020 2 specimens
BSIP Tad-571a, 546 (4 measurements)
Gujaratomyia miripes n. gen., n. sp. Giłka and Zakrzewska 2018
Zakrzewska et al. 2018 3 specimens
BSIP Tad-607a, b; Tad610 in Zakrzewska et al., 2020 (2 measurements)
Tanytarsus spp. van der Wulp 1874
Zakrzewska et al. 2020 3 specimens
BSIP Tad-533, 883a, 884
Tanytarsus forfex n. sp. Giłka and Zakrzewska 2020
Zakrzewska et al. 2020 1 specimen
BSIP Tad-568a (1 measurement)
Tanytarsus ramus n. sp. Giłka and Zakrzewska 2020
Zakrzewska et al. 2020 1 specimen
BSIP Tad-521 (2 measurements)
Insecta - Blattodea - Termitidae
Nanotermes isaacae n. gen., n. sp. Engel and Grimaldi 2011
Engel et al. 2011 1 specimen
BSIPL Tad-262 (2 measurements)
Insecta - Blattodea - Stylotermitidae
Prostylotermes kamboja n. gen., n. sp. Engel and Grimaldi 2011
Engel et al. 2011 2 specimens
BSIPL Tad-321C (2 specimens) (2 measurements)
Parastylotermes krishnai n. sp. Engel and Grimaldi 2011
Engel et al. 2011 2 specimens
BSIPL Tad-277, AMNH Tad-96 (2 measurements)
Insecta - Blattodea - Rhinotermitidae
Rhinotermitidae indet. Froggatt 1897
Engel et al. 2011 1 specimen
AMNH Tad-155
? Zophotermes sp. Engel 2011
Engel et al. 2011 4 specimens
AMNH Tad-95, Tad-278, SEMC-F000157, AMNH Tad-304 (2 measurements)
Zophotermes ashoki n. gen., n. sp. Engel and Singh 2011
Engel et al. 2011 2 specimens
BSIPL Tad-42, AMNH Tad-97 (2 measurements)
Insecta - Embioptera - Scelembiidae
Kumarembia hurleyi n. gen., n. sp. Engel and Grimaldi 2011
2 specimens
AMNH Tad-261-A, Tad-253 (2 measurements)
Insecta - Hymenoptera - Braconidae
Trichelyon tadkeshwarense n. gen., n. sp. Ortega-Blanco and Engel 2012
Ortega-Blanco et al. 2012 2 specimens
BSIPL Tad-420
Insecta - Hymenoptera - Apidae
? Protobombus sp. Cockerell 1908
Engel et al. 2013 1 specimen
AMNH Tad-41-A (2 measurements)
Melikertini indet. Engel 2001
Engel et al. 2013 1 specimen
AMNH Tad-179
Melikertes (Melikertes) kamboja n. sp. Engel and Ortega-Blanco 2013
Engel et al. 2013 2 specimens
AMNH Tad-272-A, STB-092-T10 (2 measurements)
Melikertes (Paramelikertes) gujaratensis n. sp. Engel and Ortega-Blanco 2013
Engel et al. 2013 1 specimen
AMNH Tad-41-A (2 measurements)
Insecta - Neuroptera - Coniopterygidae
Spiloconis eominuta n. sp. Grimaldi and Engel 2013
Grimaldi et al. 2013 2 specimens
BSIP Tad-316D, AMNH Tad-316B (1 measurement)
Insecta - Coleoptera - Chrysomelidae
Protorthaltica setosella n. gen., n. sp. Nadein 2019
Nadein and Perkovsky 2019 1 specimen
BSIP Tad-89 (1 measurement)
Davidaltica cambayensis n. gen., n. sp. Nadein 2019
Nadein and Perkovsky 2019 3 specimens
BSIP Tad-342, 258, 87 (1 measurement)
Cambaltica paleoindica n. gen., n. sp. Nadein 2019
Nadein and Perkovsky 2019 1 specimen
BSIP Tad-265 (1 measurement)
Insecta - Coleoptera - Staphylinidae
Paleosorius cambayensis n. gen., n. sp. Ortega-Blanco et al. 2013
Ortega-Blanco et al. 2013 1 specimen
SEMC-F000134
Protoclaviger trichodens n. gen., n. sp. Parker and Grimaldi 2014
Parker and Grimaldi 2014 1 specimen
AMNH Tad-490 (2 measurements)
Yprezethinus grimaldii n. gen., n. sp. Parker 2022
Parker 2022 1 specimen
AMNH Tad-130 (2 measurements)
Insecta - Hemiptera - Diaspididae
Normarkicoccus cambayae n. gen., n. sp. Vea and Grimaldi 2015
Vea and Grimaldi 2015 1 specimen
BSIPL Tad-135 (2 measurements)
Insecta - Hemiptera - Margarodidae
Heteromargarodes hukamsinghi n. sp. Vea and Grimaldi 2015
Vea and Grimaldi 2015 1 specimen
BSIPL Tad-139 (2 measurements)
Insecta - Hemiptera - Leptopodidae
Archaesalepta schuhi n. gen., n. sp. Grimaldi and Engel 2013
Grimaldi et al. 2013 1 specimen
BSIP Tad-15 (1 measurement)
Arachnida - Amblypygi - Paracharontidae
Paracharonopsis cambayensis n. gen., n. sp. Engel and Grimaldi 2014
Engel and Grimaldi 2014 1 specimen
AMNH Tad 456
Arachnida - Araneae - Archaeidae
Myrmecarchaea sp. Wunderlich 2004
Wood et al. 2021 1 specimen
BSIP 41985
see common names

Geography
Country:India State/province:Gujarat
Coordinates: 21.4° North, 73.1° East (view map)
Paleocoordinates:4.3° South, 63.1° East
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period: Paleogene Epoch: Eocene
Stage: Ypresian 10 m.y. bin: Cenozoic 2
Key time interval: Ypresian
Age range of interval: 56 - 47.8 m.y. ago
Stratigraphy
Stratigraphy comments: Age 50-52 Ma based on microfossils
Lithology and environment
Primary lithology: amber
Environment:terrestrial indet.
Taphonomy
Modes of preservation:body,soft parts,original chitin,amber
Size of fossils:mesofossils
Collection methods and comments
Reason for describing collection:taxonomic analysis
Museum repositories:AMNH
Collection method comments: Material collected in January 2009, 2010, and 2011 and split among BSIPL, Birbal Sahni Institute of Palaeobotany, Lucknow, India; AMNH, American Museum of Natural History, New York, and SEMC, Snow Entomological Collections, Division of Entomology, University of Kansas Natural History Museum, Lawrence. Type material is presently in the American Museum of Natural History, New York, USA (AMNH) and the Steinmann Institut für Geologie, Mineralogie, und Paläontologie, Bonn, Germany (SIPB), but primary types will eventually be transferred to the amber collection of the Birbal Sahni Institute of Palaeobotany, Lucknow, India
Metadata
Database number:124463
Authorizer:M. Clapham Enterer:M. Clapham
Modifier:M. Clapham Research group:paleoentomology
Created:2012-02-16 04:53:40 Last modified:2022-12-30 18:18:04
Access level:the public Released:2012-02-16 04:53:40
Creative Commons license:CC BY
Reference information

Primary reference:
40175. M. S. Engel, D. A. Grimaldi, H. Singh and P. C. Nascimbene. 2011. Webspinners in early Eocene amber from western India (Insecta: Embiodea). ZooKeys 148:197-208 [M. Clapham/M. Clapham]

Secondary references:

83515 D. J. Bickel, J. Martin, P. Agnihotri and H. Singh. 2022. Dolichopodidae (Diptera) from the Eocene amber deposits of Cambay and Kutch basins, India. Palaeoentomology 5:475-486 [M. Clapham/M. Clapham]
40183 M. S. Engel, D. A. Grimaldi, P. C. Nascimbene and H. Singh. 2011. The termites of Early Eocene Cambay amber, with the earliest record of the Termitidae (Isoptera). ZooKeys 148:105-123 [M. Clapham/M. Clapham]
52166 M. S. Engel and D. A. Grimaldi. 2014. Whipspiders (Arachnida: Amblypygi) in amber from the early Eocene and mid-Cretaceous, including maternal care. Novitates Paleoentomologicae 9:1-17 [M. Clapham/M. Clapham/M. Clapham]
49519 M. S. Engel, J. Ortega-Blanco, P. C. Nascimbene and H. Singh. 2013. The bees of early Eocene Cambay amber (Hymenoptera: Apidae). Journal of Melittology 25:1-12 [M. Clapham/M. Clapham]
45310 D. Grimaldi, M. S. Engel, P. C. Nascimbene and H. Singh. 2013. Coniopterygidae (Neuroptera: Aleuropteryginae) in amber from the Eocene of India and the Miocene of Hispaniola. American Museum Novitates 3770:1-20 [M. Clapham/M. Clapham]
41206 D. Grimaldi and H. Singh. 2012. The extinct genus Pareuthychaeta in Eocene ambers (Diptera: Schizophora: Ephydroidea). The Canadian Entomologist 144:17-28 [M. Clapham/M. Clapham]
48247 D. A. Grimaldi, M. S. Engel, and H. Singh. 2013. Bugs in the biogeography: Leptosaldinae (Heteroptera: Leptopodidae) in amber from the Miocene of Hispaniola and Eocene of India. Journal of the Kansas Entomological Society 86:226-243 [M. Clapham/M. Clapham]
71108 K. S. Nadein and E. E. Perkovsky. 2019. Small and common: the oldest tropical Chrysomelidae (Insecta: Coleoptera) from the lower Eocene Cambay amber of India. Alcheringa 43:597-611 [M. Clapham/M. Clapham/M. Clapham]
48264 J. Ortega-Blanco, S. Chatzimanolis, H. Singh and M. S. Engel. 2013. The oldest fossil of the subfamily Osoriinae (Coleoptera: Staphylinidae), from Eocene Cambay amber (India). The Coleopterists Bulletin 67:304-308 [M. Clapham/M. Clapham]
44208 J. Ortega-Blanco, H. Singh, and M. S. Engel. 2012. First amber fossil Rhysipolini (Hymenopera: Braconidae): a new genus and species in early Eocene Cambay amber. Acta Entomologica Musei Nationalis Pragae 52:585-594 [M. Clapham/M. Clapham]
83505 J. Parker. 2022. Transitional morphology and Afrotropical affinity of a bythinoplectine rove beetle from the early Eocene of India (Coleoptera: Staphylinidae: Pselaphinae). Palaeoentomology 5:452-460 [M. Clapham/M. Clapham]
53172 J. Parker and D. A. Grimaldi. 2014. Specialized myrmecophily at the ecological dawn of modern ants. Current Biology 24:2428-2434 [M. Clapham/M. Clapham]
62315 F. Stebner, H. Singh, J. Rust and D. A. Grimaldi. 2017. Lygistorrhinidae (Diptera: Bibionomorpha: Sciaroidea) in early Eocene Cambay amber. PeerJ 5(e3313):1-25 [M. Clapham/M. Clapham/M. Clapham]
60475 F. Stebner, R. Szadziewski, P. T. Rühr, H. Singh, J. U. Hammel, G. M. Kvifte, and J. Rust. 2016. A fossil biting midge (Diptera: Ceratopogonidae) from early Eocene Indian amber with a complex pheromone evaporator. Scientific Reports 6(34352):1-6 [M. Clapham/M. Clapham]
61370 F. Stebner, R. Szadziewski, H. Singh, S. Gunkel, and J. Rust. 2017. Biting midges (Diptera: Ceratopogonidae) from Cambay amber indicate that the Eocene fauna of the Indian subcontinent was not isolated. PLoS One 12(1):e0169144 [M. Clapham/M. Clapham]
54107 I. M. Vea and D. A. Grimaldi. 2015. Diverse new scale insects (Hemiptera: Coccoidea) in amber from the Cretaceous and Eocene with a phylogenetic framework for fossil Coccoidea. American Museum Novitates 3823:1-80 [M. Clapham/M. Clapham]
84143 R. Wagner, P. Agnihotri, and H. Singh. 2022. A new species of Sycorax (Sycoracinae: Psychodidae) from the Lower Eocene amber of Tadkeshwar, Gujarat, India. Palaeoentomology 5:319-326 [M. Clapham/M. Clapham]
78676 H. M. Wood, H. Singh, and D. A. Grimaldi. 2021. Another Laurasian connection in the Early Eocene of India: Myrmecarchaea spiders (Araneae, Archaeidae). ZooKeys 1071:49-61 [M. Clapham/M. Clapham]
73267 M. Zakrzewska, H. Singh, E. Wagner-Wysiecka and W. Giłka. 2020. Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber. Zoological Journal of the Linnean Society 189:1398-1425 [M. Clapham/M. Clapham/M. Clapham]
64893 M. Zakrzewska, F. Stebner, M. Puchalski, H. Singh, and W. Giłka. 2018. A peculiar leg structure in the first non-biting midge described from Cambay amber, India (Diptera: Chironomidae). Earth and Environmental Science Transactions of the Royal Society of Edinburgh 107:255-261 [M. Clapham/M. Clapham/M. Clapham]