East shore, Ezequiel Ramos Mexía Reservoir: Early Cenomanian, Argentina
collected by R. Spedale 1998 2019

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia - Pterosauria - Azhdarchidae
Azhdarchidae indet. Nessov 1984
Agnolín et al. 2023 1 specimen
MPCA-PV 2003, posterior half of a middle cervical vertebra
Reptilia - Theropoda
Bicentenaria argentina n. gen., n. sp. Novas et al. 2012
131 specimens
MPCA 865, posterior half of skull articulated w/lower jaws; MPCA 866, ~130 bones, incl. frags of 2 articulated premaxillae w/3 pairs of teeth, incomplete R maxilla w/1 tooth, frags of 17 dorsal vertebrae, 14 sacral vertebrae, 20 caudal vertebrae, frags of 2 scapulae, caudodorsal corner of R coracoid, proximal ends of 3 ulnae, distal end of radius, 8 manual unguals, frags of L ilium, proximal frags of 5 pubic shafts, 5 inc. femora, proximal thirds of 2 L tibiae & distal end of R, R astragalus, frags of 5 metatarsals, 15 non-ungual pedal phalanges, 8 pedal unguals, & several isolated rib frags. Collected bones include 3 left femora w/approx. same size (ca. 31 cm long), indicating at least 3 large different specimens. These match with the size observed in several isolated dorsal, sacral and caudal vertebrae w/a closed neurocentral suture. The existence of a R distal femur with an identical morphology, but considerably smaller than the largest ones, indicates the presence of probably younger individuals in the bone assemblage. This statement is supported by the presence of a fragmentary maxilla bearing a tiny tooth with a crown height of 8 mm, obviously corresponding to a juvenile individual.
Reptilia - Testudines - Chelidae
Chelidae indet. Gray 1825
Agnolín et al. 2023
Some turtle plates
see common names

Geography
Country:Argentina State/province:Río Negro
Coordinates: 39.5° South, 68.9° West (view map)
Paleocoordinates:46.5° South, 45.2° West
Basis of coordinate:stated in text
Geographic resolution:small collection
Time
Period: Cretaceous Epoch: Late Cretaceous
Stage: Cenomanian 10 m.y. bin: Cretaceous 5
Key time interval: Early Cenomanian
Age range of interval: 100.5 - 93.9 m.y. ago
Stratigraphy
Geological group:Neuquén Formation:Candeleros
Stratigraphic resolution:bed
Stratigraphy comments: Upper levels of the Candeleros Formation (Cenomanian, early Late Cretaceous; Leanza et al., 2004), Río Limay Subgroup, Neuquén Group
Lithology and environment
Primary lithology:medium,red sandstone
Secondary lithology:medium,red mudstone
Includes fossils?Y
Includes fossils?Y
Lithology description: The rocks of the Candeleros Formation which yielded the remains of Bicentenaria argentina are made up by red medium sized sandstones and mudstones, with abundant dark-brownish paleosoils representing a fluvial environment under braided and meandering regimes, as well as in aeolian conditions (Leanza et al., 2004; Garrido, 2011)
Environment:fluvial indet.
Taphonomy
Modes of preservation:body
Degree of concentration:-single event
Size of fossils:macrofossils
Preservation of anatomical detail:poor
Abundance in sediment:abundant
Disassociated major elements:many
Fragmentation:frequent
Spatial resolution:parautochthonous
Collection methods and comments
Collection excludes:some macrofossils
Collection methods:selective quarrying,surface (in situ),mechanical,field collection
Reason for describing collection:taxonomic analysis
Collectors:R. Spedale Collection dates:1998 and 2019
Collection method comments: MPCA-PV, Vertebrate Paleontology Collection, Museo Provincial “Carlos Ameghino”, Río Negro province, Argentina.

The skeletal remains of *Bicentenaria argentina* were collected by Mr. Raul Spedale in 1998 during an exceptional fall in the water level of the Ezequiel Ramos Mexía Reservoir (Fig. 1). All the bones were collected from a single quarry roughly 40 centimeters long, 30 centimeters wide, and 50 centimeters thick.

The specimen (MPCA-PV 2003) was found during a field trip performed under the project of relocation and georeferencing of fossil sites in the Río Negro province, carried out by the former Dirección de Patrimonio y Museos from the Río Negro goverment and supported by some professionals of the Universidad Nacional de Rio Negro (UNRN) in June, 2019.
Metadata
Database number:132347
Authorizer:R. Benson, E. Vlachos Enterer:R. Benson, F. Aspromonte
Modifier:F. Aspromonte Research group:vertebrate
Created:2012-08-14 18:25:44 Last modified:2023-11-02 11:09:54
Access level:the public Released:2012-08-14 18:25:44
Creative Commons license:CC BY
Reference information

Primary reference:

42624. F. E. Novas, M. D. Ezcurra, F. L. Agnolin, D. Pol, and R. Ortíz. 2012. New Patagonian Cretaceous theropod sheds light about the early radiation of Coelurosauria. Revista del Museo Argentino de Ciencias Naturales, nuevo serie 14(1):57-81 [R. Benson/R. Benson/M. Carrano]

Secondary references:

86257 F. L. Agnolín, S. Rozadilla, R. D. Juárez-Valieri and J. G. Meso. 2023. Oldest azhdarchid (Pterosauria) record from South America. Revista del Museo Argentino de Ciencias Naturales 25(2):309-314 [E. Vlachos/F. Aspromonte]