Bromacker Quarry (upper level of Tambach sandstone): Artinskian, Germany
collected by Schäfer-Gotha 1887, 1893

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Tambachichnium sp. Müller 1954
Voigt et al. 2007
Amphibia
Amphisauropus sp. Haubold 1970
Voigt et al. 2007
Amphibia - Temnospondyli
Dissorophoidea informal new sp.
Martens et al. 2005
"family indeterminate"
Amphibia - Temnospondyli - Trematopidae
Tambachia trogallas n. gen., n. sp. Sumida et al. 1998
Sumida et al. 1998 1 individual
MNG 7722 holotype
Trematopidae informal new sp.
Martens et al. 2005
    = Rotaryus gothae n. gen., n. sp. Berman et al. 2011
Berman et al. 2011
MNG 10182 holotype
Amphibia - Temnospondyli - Dissorophidae
Dissorophidae informal new sp.
Martens et al. 2005
Amphibia - Temnospondyli
Georgenthalia clavinasica n. gen., n. sp. Anderson et al. 2008
Anderson et al. 2008 1 individual
MNG 11135 holotype
unclassified
Ichnium brachydactylum Pabst 1908
Pabst 1908 2 individuals
recombined as Gilmoreichnus brachydactylus
Gotha n. 1841, 1843
Diadectidae
Orobates pabsti n. gen., n. sp. Berman et al. 2004
Berman et al. 2004 4 individuals
MNG 10181 holotype; MNG 8760, 8980, 11133, 11134 paratypes
Ichniotherium cottae (Pohlig 1885)
Pabst 1908 11 specimens
Diadectes absitus n. sp. Berman et al. 1998
4 specimens
MNG 8853 (Holotype: nearly complete articulated skeleton), MNG 8747 (nearly complete skull), MNG 7721 (nearly complete postcranial skeleton), and MNG 8978 (nearly complete but disarticulated postcranial skeleton)
Caseidae
Caseidae informal new sp.
Martens et al. 2005
Martensius bromackerensis n. gen., n. sp. Berman et al. 2020
Berman et al. 2020 1 specimen
MNG 13814 (Holotype), adult and largest of the specimens, consisting of a nearly complete, well–preserved skeleton preserved in several, closely associated, articulated sections
Varanopidae
Varanopseidae informal new sp.
Martens et al. 2005 2 specimens
    = Tambacarnifex unguifalcatus n. gen., n. sp. Berman et al. 2014
Berman et al. 2014
holotype: MNG 10596 (greater part of postcranial skeleton preserved both in bone and as impressions on counterpart blocks); paratype: MNG 15037 (partial left dentary exposed in lateral view)
Sphenacodontidae
Dimetrodon teutonis n. sp. Berman et al. 2001
Berman et al. 2001 1 individual
MNG 10598 holotype
Brachystelechidae
Bromerpeton subcolossus n. gen., n. sp. MacDougall et al. 2024
MacDougall et al. 2024 1 specimen
Holotype: MNG 16545, a partial skull and mandible with left humerus and largely complete right forelimb
Reptilia
Thuringothyris mahlendorffae n. gen., n. sp. Boy and Martens 1991
Müller et al. 2006 8 individuals
MNG 7729 holotype; MNG 10183, 10647, 10652, 11191 referred specimens
Dimetropus sp. Romer and Price 1940
Voigt et al. 2007
Ichnium acrodactylum n. sp. Pabst 1908
Pabst 1908 12 individuals
synonym of Saurichnites leisnerianus
incl. subspecies alternans, curvata, dispar, and microdactylum; Gotha no. 1760, 1762, 1765–1766, 1823, 1825, 1827–1828, 1838, 1983, 1985–1986
Reptilia - Bolosauridae
Eudibamus cursoris n. gen., n. sp. Berman et al. 2000
Berman et al. 2000 1 individual
MNG 8852 holotype
unclassified
Varanopus sp. Moodie 1929
Voigt et al. 2007
Ichnium dolichodactylum Pabst 1908
Pabst 1908 1 individual
synonym of Varanopus microdactylus
Gotha no. 1785
Seymouriidae
Seymouria sanjuanensis Vaughn 1966
Berman et al. 2000 4 individuals
MNG 7727, 8759, 10553, 10554
Myriapoda
Myriapoda sp. (Latreille 1802)
Martens et al. 2005
"family indeterminate, 1-2 undescribed new species"
Insecta
Insecta sp. Linnaeus 1758
Martens et al. 2005
"family indeterminate, 2-3 undescribed new species"
Insecta - Blattodea - Mylacridae
Moravamylacris kukalovae Schneider 1980
Martens et al. 2005
Opsiomylacris sp. Haupt 1952
Martens et al. 2005
Insecta - Phyloblattidae
Phylloblatta sp. (Handlirsch 1906)
Martens et al. 2005
original and current combination Phyloblatta
cf. Anthracoblattina sp. Scudder 1879
Martens et al. 2005
Kunguroblattina sp. Martynov 1930
Martens et al. 2005
Branchiopoda - Lioestheriidae
Lioestheria monticula Martens 1983
Martens et al. 2005
unclassified
Ichnium sphaerodactylum n. sp. Pabst 1908
Pabst 1908 42 specimens
Gotha no. 1351–1352, 1367–1371, 1393–1395, 1401 2, 1747–1749, 1814, 1819, 1840
see common names

Geography
Country:Germany State/province:Thuringia County:Gotha
Coordinates: 50.8° North, 10.6° East (view map)
Paleocoordinates:8.9° North, 22.6° East
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period:Permian Epoch:Cisuralian
Stage:Artinskian 10 m.y. bin:Permian 2
Key time interval:Artinskian
Age range of interval:290.10000 - 283.50000 m.y. ago
Stratigraphy
Geological group:Rotliegend Formation:Tambach Member:Tambach-Sandstein
Stratigraphic resolution:group of beds
Stratigraphy comments: "At the Bromacker quarry site are two superimposed stratigraphic successions that can be characterized by their facies associations and are referred to as the Lower Beds and Upper Beds (Eberth et al. 2000). All the vertebrates from the Bromacker quarry come from the Upper Beds [...]. The vertebrates are almost exclusively restricted to two [... horizons ...] which are separated by 50 cm in a stratigraphic interval of 1.2 m." (Berman et al., 2001)
The Tambach Formation is of Artinskian age according to "Stratigraphische Tabelle von Deutschland 2002", however, a Kungurian age is proposed by Lützner (2006) in LITHOLEX stratigraphic online database [http://www.bgr.de/app/LithoLex/gesamt_ausgabe_neu.php5?id=25]
Lithology and environment
Primary lithology:very fine,brown,red lithified sandstone
Includes fossils?Y
Lithology description: "massive, red-brown, very fine-grained sandstone and siltstone" (Berman et al., 2001). For further details see Eberth et al. (2000).
Environment:fluvial-lacustrine indet. Tectonic setting:intramontane basin
Geology comments: "alluvial paleochannel and sheetflood facies and lacustrine suspension deposits" (Berman et al., 2001).
"During the time that the Lower and Upper beds were deposited, the Tambach Basin was internally-drained and probably characterized by a wet-and-dry tropical or wetter climate that was hot year-round with seasonal-to-subseasonal cycles of rainfall and drying. [...] In the Bromacker area, the floodbasin was never deeply desiccated or heavily vegetated between flooding events." Eberth et al. (2000).
Taphonomy
Modes of preservation:body,permineralized
Size of fossils:macrofossils
Preservation of anatomical detail:excellent
Articulated whole bodies:some
Associated major elements:some
Disassociated major elements:some
Disassociated minor elements:some
Temporal resolution:snapshot
Collection methods and comments
Collection excludes:all macrofossils
Collection methods:bulk,surface (in situ),field collection
Reason for describing collection:taxonomic analysis
Collectors:Schäfer-Gotha Collection dates:1887, 1893
Collection method comments: MNG = Museum der Natur, Gotha
Metadata
Database number:80112
Authorizer:R. Reisz, M. Carrano, J. Mueller, E. Dunne, A. Dunhill Enterer:R. Reisz, B. Allen, T. Liebrecht, M. Carrano, E. Dunne
Modifier:M. Carrano Research group:vertebrate
Created:2008-04-04 13:49:44 Last modified:2021-12-07 11:20:18
Access level:authorizer only Released:2011-04-04 13:49:44
Creative Commons license:CC BY
Reference information

Primary reference:
26922. D.S. Berman, S.S. Sumida, and T. Martens. 1998. Diadectes (Diadectomorpha: Diadectidae) from the Early Permian of Central Germany, with description of a new species. Annals of Carnegie Museum 67(1):53-93 [R. Reisz/D. Mazierski]

Secondary references:

27263ETE J. S. Anderson, A. C. Henrici, S. S. Sumida, T. Martens, and D. S. Berman. 2008. Georgenthalia clavinasica, a new genus and species of dissorophoid temnospondyl from the Early Permian of Germany, and the relationships of the family Amphibamidae. Journal of Vertebrate Paleontology 28(1):61-75 [M. Carrano/M. Carrano]
5765ETE 1524D. S. Berman, A. C. Henrici, S. S. Sumida and T. Martens. 2000. Redescription of Seymouria sanjuanensis (Seymouriomorpha) from the Lower Permian of Germany based on complete, mature specimens with a discussion of paleoecology of Bromacker locality assemblage. Journal of Vertebrate Paleontology 20(2):253-268 [A. Behrensmeyer/A. Behrensmeyer/J. Alroy]
30738 D. S. Berman, A. C. Henrici, R. A. Kissel, S. S. Sumida, and T. Martens. 2004. A New Diadectid (Diadectomorpha), Orobates pabsti, from the Early Permian of Central Germany. Bulletin of Carnegie Museum of Natural History 35(1):1-36 [J. Mueller/T. Liebrecht]
40304 D. S. Berman, A. C. Henrici, T. Martens, S. S. Sumida, and J. S. Anderson. 2011. Rotaryus gothae, a New Trematopid (Temnospondyli: Dissorophoidea) from the Lower Permian of Central Germany. Annals of Carnegie Museum 80(1):49-65 [J. Mueller/T. Liebrecht]
50796 D. S. Berman, A. C. Henrici, S. S. Sumida, T. Martens, and V. Pelletier. 2014. First European record of a Varanodontine (Synapsida: Varanopidae): member of a unique Early Permian upland paleoecosystem, Tambach Basin, central Germany. In C. F. Kammerer, K. D. Angielczyk, J. Fröbsich (eds.), Early Evolutionary History of the Synapsida 69-86 [R. Benson/R. Benson]
72029 D. S. Berman, H. C. Maddin, A. C. Henrici, S. S. Sumida, D. Scott and R. R. Reisz. 2020. New primitive caseid (Synapsida, Caseasauria) from the early Permian of Germany. Annals of Carnegie Museum 86(1):43-75 [E. Dunne/E. Dunne]
28679 D. S. Berman, R. R. Reisz, D. Scott, A. C. Henrici, S. S. Sumida and T. Martens. 2000. Early Permian Bipedal Reptile. Science 290:969-972 [J. Mueller/T. Liebrecht]
28661 D. S. Berman, R. R. Reisz, T. Martens and A. C. Henrici. 2001. A new species of Dimetrodon (Synapsida: Sphenacodontidae) from the Lower Permian of Germany records first occurrence of genus outside of North America. Canadian Journal of Earth Sciences 38(5):803-812 [J. Mueller/T. Liebrecht]
5766ETE 1525D. A. Eberth, D. S. Berman, S.S. Sumida and H. Hopf. 2000. Lower Permian Terrestrial Paleoenvironments and Vertebrate Paleoecology of the Tambach Basin (Thuringia, Central Germany): The Upland Holy Grail. Palaios 15(4):293-313 [A. Behrensmeyer/A. Behrensmeyer/T. Liebrecht]
87733 M. J. MacDougall, A. Jannel, A. C. Henrici, D. S. Berman, S. S. Sumida, T. Martens, N. B. Fröbisch and J. Fröbisch. 2024. A new recumbirostran ‘microsaur’ from the lower Permian Bromacker locality, Thuringia, Germany, and its fossorial adaptations. Scientific Reports 14(4200) [E. Dunne/E. Dunne]
40162 T. Martens, D. S. Berman, A. C. Henrici and S. S. Sumida. 2005. The Bromacker Quarry - the Most Important Locality of Lower Permian Terrestrial Vertebrate Fossils Outside of North America. The Nonmarine Permian. New Mexico Museum of Natural History and Science Bulletin 30:214-215 [J. Mueller/T. Liebrecht]
28678 J. Müller, D. S. Berman, A. C. Henrici, T. Martens, and S. S. Sumida. 2006. The Basal Reptile Thuringothyris mahlendorffae (Amniota: Eureptilia) from the Lower Permian of Germany. Journal of Paleontology 80(4):726-739 [J. Mueller/T. Liebrecht]
78836 W. Pabst. 1908. Die Tierfährten in dem Rotliegenden “Deutschlands” [The animal tracks in the “German” Rotliegenden]. Nova Acta. Abhandlungen der Kaiserlichen Leopoldinisch-Carolinischen Deutschen Akademie der Naturforscher 89(2):315-480 [M. Carrano/M. Carrano]
30596 S. S. Sumida, D. S. Berman, and T. Martens. 1998. A New Trematopid Amphibian from the Lower Permian of Central Germany. Palaeontology 41(4):605-629 [J. Mueller/T. Liebrecht]
67270 S. Voigt, D. S. Berman, and A. C. Henrici. 2007. First well-established track-trackmaker association of paleozoic tetrapods based on Ichniotherium trackways and diadectid skeletons from the Lower Permian of Germany. Journal of Vertebrate Paleontology 27(3):553-570 [A. Dunhill/B. Allen]