Sossis (Eocene of Spain)

Where: Catalonia, Spain (42.3° N, 1.1° E: paleocoordinates 41.0° N, 2.2° W)

• coordinate estimated from map

• outcrop-level geographic resolution

When: Sossís Member (Collegats Formation), Priabonian (38.0 - 33.9 Ma)

• MP17a

Environment/lithology: lacustrine; claystone and sandstone

Size classes: macrofossils, mesofossils

Primary reference: M. Crusafont-Pairo and J. M. Golpe Posse. 1975. Les prosimiens de l'Eocene de la region preaxiale pyrenaique. II. Adapidae. Colloque international CNRS. Problemes actuels de paleontologie-evolution des vertebres (218)851-860 [E. Fara/E. Fara/E. Fara]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 31751: authorized by Emmanuel Fara, entered by Emmanuel Fara on 24.05.2003, edited by Albert Garcia Selles, Terri Cleary and Philip Mannion

Creative Commons license: CC BY (attribution)

Taxonomic list

• incomplete
Mammalia
 Cimolesta - Apatemyidae
Apatemyidae indet.4 Matthew 1909 eutherian
Apatemyidae indet.5 Matthew 1909 eutherian
Heterohyus nanus5 Teihard de Chardin 1916 eutherian
 Theriamorpha - Nyctitheriidae
Euronyctia montana5 Sigé 1997 placental
 Artiodactyla - Anoplotheriidae
Dacrytherium ovinum3 Owen 1857 even-toed ungulate
 Artiodactyla - Xiphodontidae
Leptotheridium lugeoni3 Stehlin 1910 even-toed ungulate
IPS 5817
Xiphodon intermedium3 Stehlin 1910 even-toed ungulate
Haplomeryx euzetensis3 Depéret 1917 even-toed ungulate
IPS 5888, IPS 5842, IPS 14859, IPS 14860
Dichodon cervinum6 even-toed ungulate
 Artiodactyla - Cebochoeridae
Cebochoerus helveticus3 Pictet and Humbert 1869 even-toed ungulate
IPS 5889, 5887, 5867
Acotherulum sp.3 Gervais 1850 even-toed ungulate
IPS 29660-29666
 Perissodactyla - Palaeotheriidae
"Anchilophus dumasi" = Metanchilophus dumasi4 Gervais 1849 odd-toed ungulate
Palaeotherium magnum4 Cuvier 1804 odd-toed ungulate
Palaeotherium medium4 Cuvier 1804 odd-toed ungulate
Palaeotherium crassum4 Cuvier 1805 odd-toed ungulate
Plagiolophus annectens4 Owen 1847 odd-toed ungulate
 Theriamorpha -
Miacis exilis4 placental
 Creodonta - Hyaenodontidae
Hyaenodon minor4 Lange-Badré 1979 creodont
 Rodentia - Gliridae
 Rodentia - Theridomyidae
Estellomys ibericus n. sp.4 Thaler 1966 rodent
Two right M1/2s, Sabadell Museum, holotype, UM.1918, paratype, UM.1887
Elfomys parvulus4 Hartenberger 1971 rodent
Pseudoltinomys cf. phosphoricus4 Hartenberger 1973 rodent
Paradelomys crusafonti n. sp.4 Thaler 1966 rodent
UM 1882 - type
 Rodentia - Pseudosciuridae
Treposciurus mutabilis4 Schmidt-Kittler 1970 rodent
 Primates - Adapidae
Adapis sp.6 Cuvier 1822 primate
Adapis parisiensis Blainville 1841 primate
 Primates - Notharctidae
Nievesia sossisensis n. gen. n. sp.4
Nievesia sossisensis n. gen. n. sp.4 Marigó et al. 2013 primate
 Primates - Omomyidae
Pseudoloris parvulus6 Filhol 1890 tarsier
Microchoerus hookeri n. sp.5 Minwer-Barakat et al. 2017 tarsier
 Theriamorpha - Leptictidae
Leptictis sp.4 Leidy 1868 eutherian
Leptictis sp.5 Leidy 1868 eutherian
 Theriamorpha - Herpetotheriidae
Peratherium lavergnense4 Crochet 1979 marsupial
Reptilia
 Squamata - Anguidae
Glyptosaurinae indet.2 Marsh 1872 squamates
IPS 56167, fragment of ?dentary with two preserved teeth; IPS 56149, fragment of ?maxilla with one preserved tooth; IPS 56150, fragment of maxilla with one preserved tooth.
Glyptosaurini indet.2 Sullivan 1979 squamates
IPS 49845a, hexagonal skull osteoderm; IPS 49845b, skull osteoderm with unusual shape; IPS 49845c, 25 skull osteoderms with unusual shapes; IPS 49845d, >600 hexagonal skull osteoderms; IPS 49845e skull osteoderm; 59477, seven skull osteoderms; IPS 59497, two hexagonal skull osteoderms; IPS 59468, hexagonal skull osteoderm; IPS 59485, 28 skull osteoderms; IPS 59508, two skull osteoderms; IPS 59527, 22 worn skull osteoderms. IPS 49846a, body osteoderm; IPS 49846b, >700 body osteoderms; IPS 49846c, >1300 fragments of osteoderm; IPS 49846d, rounded body osteoderm; IPS 49846e, rounded body osteoderm; IPS 49846f, 50 rounded body osteoderms; IPS 56068, skull bone fragment with two fused osteoderms; IPS 56069, skull bone fragment with one fused osteoderm; IPS 56070, skull bone fragment with two fused osteoderms; IPS 56071, skull bone fragment with one fused osteoderm; IPS 56072, skull bone with one fused osteoderm; IPS 56073, skull bone with one fused osteoderm; IPS 56075, skull bone with one fused osteoderm; IPS 59526, skull osteoderm; IPS 59528, body osteoderm.
Anguinae indet.2 Gray 1825 squamates
IPS 49847a, elongated body osteoderm; IPS 49847b, thousands of elongated body osteoderms; IPS 49851a, rounded body osteoderm; IPS 49851b, more than 100 rounded body osteoderms; IPS 56082, partial dorsal vertebra; IPS 56162, two body osteoderms; IPS 56234, two partial vertebrae; IPS 56085, posterior fragment of parietal; IPS 56086, posterior fragment of a right frontal; IPS 56080, dorsal vertebra; IPS 56081, 37 partial vertebral centra, condyles showing tendency to be triangular in ventral view; IPS 56083, multiple vertebral condyles; IPS 59486, 30 body osteoderms; IPS 56159, body osteoderm; IPS 56231, three body osteoderms; IPS 59490, four fragments of vertebrae; IPS 59504, vertebra condyle; IPS 59509, three body osteoderms; IPS 59487, two vertebral condyles; and less certainly: IPS 59473, vertebral centrum; IPS 59496, four partial vertebrae; IPS 56128, tooth-bearing bone with one broken tooth; 56129, maxilla fragment with one broken tooth; IPS 56130, anterior fragment of left maxilla; IPS 56136, fragment of left dentary with one broken tooth.
 Squamata -
Cordyliformes indet.2 Fitzinger 1826 squamates
IPS 49848, more than 300 body osteoderms; and less certainly IPS 56195, anterior (symphyseal region) fragment of left dentary with two preserved teeth.
Pyrenasaurus evansae1 Bolet and Auge 2014 squamates
IPS 56061, partial left dentary with two teeth; IPS 56168, fragment of right dentary with one tooth
 Squamata - Cordylidae
Pseudolacerta sp.2 De Stefano 1903 spinytail lizard
IPS 56164, partial right dentary with eight teeth (five complete); IPS 56165, fragment of dentary with two teeth; IPS 56152, anterior part of right dentary with five (broken) teeth
 Squamata - Scincidae
? Scincidae indet.2 Gray 1825 skink
IPS 56193, ?dentary fragment with five preserved teeth; ?IPS 56052, fragment of tooth-bearing bone with one tooth.
 Squamata - Iguanidae
Geiseltaliellus sp.2 Kuhn 1944 squamates
IPS 56090, partial left dentary with twelve teeth (eight complete)
Geiseltaliellus pradiguensis2 Auge 2007 squamates
IPS 56093, partial right dentary with four teeth; IPS 56094, partial right maxilla with four teeth (three complete); IPS 59521, fragment of ?dentary with five teeth; IPS 59523, five fragments of dentary with teeth
 Squamata - Lacertidae
? Lacertidae "amblyodont form"2 Bonaparte 1831 squamates
IPS 56169, fragment of left dentary with three preserved teeth; and, less certainly: IPS 56102, posterior left dentary fragment with two preserved teeth; ?IPS 56036, posterior left dentary fragment with two worn teeth; IPS 49913, right dentary fragment with two preserved teeth; IPS 56067, anterior portion of left maxilla with three preserved teeth; and IPS 56170, anterior portion of right maxilla with three preserved teeth.
Lacertidae indet.2 Bonaparte 1831 squamates
IPS 49855, eleven dentary fragments; IPS 56043, partial left frontal; IPS 56044, IPS 56045, ?IPS 56047, fragments of right frontal; ?IPS 56166 (Figure 4.11), partial fused frontal; IPS 59479, fragment of frontal; IPS 56046, fragment of parietal; IPS 56053, fragment of right maxilla with three preserved teeth; IPS 56055, fragment of right dentary with two broken teeth; IPS-56058, fragment of right dentary with one preserved tooth; IPS 56066, anterior region of left dentary with eight preserved teeth.
Gracilicerta sindexi2 Augé 2005 squamates
IPS 56171, partial left dentary with eight preserved teeth (Figure 4.4); IPS 49854, fragment of left dentary with three preserved teeth; IPS 49856, fragment of left dentary with one preserved tooth; IPS 56050, fragment of left maxilla with two preserved teeth.
Dormaalisaurus rossmanni2 Augé and Smith 2002 squamates
IPS 59524, partial left dentary with four preserved teeth; IPS 56048, partial right maxilla with six preserved teeth; IPS 56049, partial right maxilla with four preserved teeth; IPS 56060, four fragments of dentary with few preserved teeth; IPS 56089 partial right maxilla with three preserved teeth + partial left dentary with five broken teeth; IPS 56097, right dentary fragment with one tooth; IPS 56127, posterior fragment of left dentary with two preserved teeth.
Dormaalisaurus girardoti2 Augé and Smith 2002 squamates
IPS 56056, partial right dentary with two teeth; IPS 56051, fragment of left maxilla with four teeth; IPS 56054, fragment of right dentary with three preserved teeth; IPS 56059, anterior end of maxilla with two preserved teeth; IPS 59525, dentary and maxilla fragments
 Squamata -
Iguania indet.2 Cuvier 1817 squamates
Unidentifiable pleurodont iguanian material lacking diagnostic characters: IPS 49853, IPS 56092, IPS 56138- 56144, IPS 56230, IPS 56232, IPS 59481, IPS 59501, IPS 59506, IPS 59522, tooth-bearing bone fragments; ?IPS 56109, partial frontal; ?IPS 56091, partial parietal
 Squamata - Blanidae
cf. Blanidae indet.2 Kearney 2003 worm lizard
IPS 56062, partial left dentary with five broken teeth; IPS 56063, almost complete dorsal vertebra; IPS 56064, almost complete dorsal vertebra; IPS 56155, partial dorsal vertebra.
 Squamata -
Gekkota "Form A"2 Cuvier 1817 squamates
IPS 56176, partial right dentary with one preserved tooth; IPS 56178, partial right dentary with one preserved tooth; IPS 56065, partial right maxilla with one preserved tooth.
Gekkota "Form B"2 Cuvier 1817 squamates
IPS 56132, partial left dentary with one preserved tooth; IPS 56116, fragment of right dentary with one preserved tooth; IPS 56133, fragment of left dentary; IPS 56172, partial right dentary with three broken teeth; IPS 56182, fragment of right dentary; IPS 56183, anterior portion of right dentary with one preserved tooth; IPS 56184, fragment of right dentary with two broken teeth; IPS 56185, anterior portion of right dentary; IPS 56186, fragment of ?right dentary with two broken teeth; IPS 56187, fragment of right dentary with two broken teeth; IPS 56188, fragment of ?right dentary; IPS 56192, anterior portion of right dentary with three broken teeth; IPS 59499, fragment of dentary with one broken tooth; IPS 59511, partial dentaries and fragments of dentaries
Gekkota "Form C"2 Cuvier 1817 squamates
IPS 56117, fragment of right dentary with one preserved tooth; IPS 56173, partial right dentary with five broken teeth; IPS 56174, fragment of right dentary with two teeth; IPS 56175, anterior region of a right dentary with six broken teeth; IPS 56177, partial right dentary with one preserved tooth; IPS 56180, fragment of ?left dentary with five broken teeth; IPS 56190, anterior portion of left dentary with three broken teeth.
Gekkota indet.2 Cuvier 1817 squamates
Additional gekkotan material: IPS 59478, partial fused premaxilla ; IPS 56114, left dentary fragment with two preserved teeth; IPS 56115, left dentary fragment with two teeth; IPS 56120, fragments of dentaries; IPS 56179, fragment of right dentary with four broken teeth; IPS 56181, fragment of right dentary with four broken teeth; IPS 56189, fragment of dentary with two broken teeth; IPS 56191, fragments of dentaries; IPS 59520, fragments of dentaries.