Bitterfeld amber (Hoffeins collection): Priabonian, Germany
collected by C. Hoffeins, H.W. Hoffeins

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Arachnida - Acariformes - Smarididae
Fessonia wunderlichi Bartel et al. 2015
Bartel et al. 2015 1 specimen
Hoffeins 385-3
Arachnida - Araneae - Dictynidae
Balticocryphoeca curvitarsis Wunderlich 2004
Wunderlich 2004 1 specimen
No. 949
Arachnida - Araneae - Uloboridae
? Hyptiomopes sp. Wunderlich 2004
Wunderlich 2004 1 specimen
801a
Arachnida - Araneae - Theridiidae
Theridiidae indet. Sundevall 1833
Wunderlich 2004 1 specimen
801b
Arachnida - Araneae - Cyatholipidae
Spinilipus bispinosus Wunderlich 2004
Wunderlich 2004 1 specimen
No. 680
Insecta - Diptera - Phoridae
Aenigmatias longicornis n. sp. Brown 2016
Brown 2016 2 specimens
Hoffeins 254/3 (1 measurement)
Insecta - Diptera - Anthomyzidae
Protanthomyza tschirnhausi n. sp. Roháček 2013
Roháček 2013 2 specimens
829-8 (collected at Goitzsche mine 2010), 830-7 (purch. 2010) (4 measurements)
Lacrimyza christelae n. sp. Roháček 2013
Roháček 2013 1 specimen
830-5 (collected at Goitzsche mine 1998) (2 measurements)
Insecta - Diptera - Pseudopomyzidae
Eopseudopomyza szasziewskii Hoffeins and Woźnica 2013
Hoffeins and Woźnica 2013 1 specimen
Hoffeins 670-3 (2 measurements)
Insecta - Diptera - Empididae
Heleodromia starki n. sp. Hoffeins et al. 1997
Hoffeins et al. 1997 1 specimen
Hoffeins 707 (Goitsche pit) (2 measurements)
Insecta - Diptera - Anisopodidae
Sylvicola hoffeinsorum Wojtoń et al. 2018
Wojtoń et al. 2018 1 specimen
SDEI 664-2
Sylvicola splendida (Meunier 1904)
Wojtoń et al. 2018 1 specimen
SDEI 664-1
Insecta - Diptera - Keroplatidae
Antlemon sp. Loew 1871
Hoffeins et al. 2010 1 specimen
CCHH 1700-1 (Goitszche mine) (2 measurements)
Insecta - Diptera - Nymphomyiidae
Nymphomyia succina n. sp. Wagner et al. 2000
1 specimen
Hoffeins 381/2 (2 measurements)
Insecta - Lepidoptera - Micropterigidae
Wichardius fossiliphilus Mey 2019
Mey 2019 1 specimen
CCHH 1420/3
Insecta - Mecoptera - Bittacidae
Hylobittacus picteti n. sp. Krzemiński 2007
Krzemiński 2007 1 specimen
DEIM 674/1
Insecta - Coleoptera - Dermestidae
Attagenus balticus Háva et al. 2008
Háva and Alekseev 2015 1 specimen
Hoffeins 115-1
Insecta - Coleoptera - Nitidulidae
Microsoronia hoffeinsorum n. sp. Kirejtshuk and Kurochkin 2010
Kirejtshuk and Kurochkin 2010 1 specimen
DEI 276-2 1
Insecta - Coleoptera - Cyclaxyridae
Neolitochropus bedovoyi (Lyubarsky and Perkovsky 2011)
Gimmel and Szawaryn 2020 1 specimen
CCHH 504-17
Neolitochropus hoffeinsorum n. gen., n. sp. Lyubarsky and Perkovsky 2016
Lyubarsky and Perkovsky 2016 1 specimen
synonym of Neolitochropus bedovoyi
CCHH 1206-1 (2 measurements)
Insecta - Coleoptera - Cryptophagidae
Atomaria (Anchicera) saxonica n. sp. Lyubarsky and Perkovsky 2018
Lyubarsky and Perkovsky 2018 1 specimen
CCHH 1223-4
Insecta - Coleoptera - Erotylidae
Microzavaljus saxonicum n. gen., n. sp. Lyubarsky and Perkovsky 2018
Lyubarsky and Perkovsky 2018 1 specimen
Hoffeins 1223-1
Serramorphus rasnitsyni n. gen., n. sp. Lyubarsky and Perkovsky 2017
Lyubarsky and Perkovsky 2017 1 specimen
CCHH 26-4 (1 measurement)
Triplax contienensis n. sp. Alekseev 2014
Alekseev 2014 1 specimen
Hoffeins 1650-4 (1 measurement)
Insecta - Coleoptera - Anamorphidae
Kleinzaches germanicianus n. gen., n. sp. Alekseev and Tomaszewska 2018
Alekseev and Tomaszewska 2018 1 specimen
CCHH 181-1
Insecta - Coleoptera - Coccinellidae
Baltosidis sp. Szawaryn 2021
Szawaryn 2021 1 specimen
CCHH 743-1
Insecta - Coleoptera - Endomychidae
Trochoideus resinatissimus n. sp. Alekseev and Tomaszewska 2018
Alekseev and Tomaszewska 2018 1 specimen
CCHH 53-2
Insecta - Coleoptera - Zopheridae
? Bitoma sp. Herbst 1793
Alekseev and Bukejs 2016 1 specimen
CCHH 181-3
Insecta - Coleoptera - Aderidae
Palaeocnopus densipunctatus Alekseev and Grzymala 2015
Alekseev and Grzymala 2015 1 specimen
Hoffeins 25-4 (1 measurement)
Palaeocnopus mara n. sp. Alekseev and Grzymala 2015
Alekseev and Grzymala 2015 2 specimens
Hoffeins 890-1, 609-3 (2 measurements)
Palaeocnopus saeticornis n. sp. Alekseev and Grzymala 2015
Alekseev and Grzymala 2015 2 specimens
Hoffeins 890-3, 1049-2 (2 measurements)
Palaeocnopus glabricornis n. sp. Alekseev and Grzymala 2015
Alekseev and Grzymala 2015 1 specimen
Hoffeins 890-2 (1 measurement)
Insecta - Coleoptera - Chrysomelidae
Crepidodera decolorata Nadein and Perkovsky 2010
Bukejs et al. 2016 1 specimen
CCHH 337-3 (1 measurement)
Insecta - Hemiptera - Lithuanicoccidae
Lithuanicoccus damzeni Koteja 2008
Koteja 2008 1 specimen
Hoffeins 1248-1
Insecta - Hemiptera - Kuwaniidae
Hoffeinsia foldii Koteja 2008
Koteja 2008 4 specimens
Hoffeins 1249-1, 0796-1, 0796-2, 0807-3
Insecta - Hemiptera - Aradidae
Calisius balticus Usinger 1941
Heiss 2013 2 specimens
Hoffeins 844/1, 2565 (2 measurements)
Insecta - Thysanoptera - Stenurothripidae
Stenurothrips brachycerus n. sp. Schliephake 1999
Schliephake 1999 1 specimen
Nr. 750/1 (1 measurement)
Insecta - Thysanoptera - Thripidae
Archaeothrips hoffeinsi n. sp. Schliephake 1997
Schliephake 1997 1 specimen
synonym of Archaeothrips latipennis
Hoffeins 704 (2 measurements)
Protoxythrips stenurus Schliephake 1993
Schliephake 2005 1 specimen
Hoffeins 1477/3 (2 measurements)
Protoxythrips ursulae Schliephake 1993
Schliephake 2001 2 specimens
Hoffeins 1368/2; 1477/6 in Schliephake, 2005 (2 measurements)
Protoxythrips conclusus (Priesner 1929)
Schliephake 2005 1 specimen
Hoffeins 1477/5 (2 measurements)
Eochirothrips mirocornis n. gen., n. sp. Schliephake 1999
Schliephake 1999 1 specimen
Nr. 272/2 (1 measurement)
Anaphothrips fuscicaudeus Schliephake 1993
Schliephake 2005 1 specimen
Hoffeins 1477/7 (1 measurement)
Incertothrips hoffeinsorum n. gen., n. sp. Schliephake 2000
Schliephake 2000 1 specimen
Hoffeins 1294 (1 measurement)
Frankliniella intonsoidea n. sp. Schliephake 1999
Schliephake 1999 1 specimen
Nr. 576/1 (1 measurement)
Taeniothrips (Saxonothrips) saxonicus n. sp. Schliephake 1999
Schliephake 1999 1 specimen
synonym of Taeniothrips balticus
Nr. 576/2 (1 measurement)
Taeniothrips (Saxonothrips) amphoroideus Schliephake 1999
Schliephake 1999 1 specimen
synonym of Taeniothrips balticus
Nr. 272/1 (1 measurement)
Taeniothrips (Saxonothrips) goitschei n. sp. Schliephake 1999
Schliephake 1999 1 specimen
Nr. 576/2 (1 measurement)
Taeniothrips usedomi (Schliephake 1999)
Schliephake 2005 1 specimen
original and current combination Taeniothrips (Balticothrips) usedomi
Hoffeins 1477/1 (1 measurement)
Protanaphothrips minoricornis (Schliephake 1999)
Schliephake 2005 1 specimen
Hoffeins 1477/2 (1 measurement)
Insecta - Thysanoptera - Melanthripidae
Archankothrips hoffeinsianus n. sp. Schliephake 2000
Schliephake 2000 1 specimen
Hoffeins 1244/3 (1 measurement)
Eocranothrips leptocerus n. sp. Schliephake 1999
Schliephake 1999 1 specimen
synonym of Melanothrips annulicornis
Nr. 576/3 (1 measurement)
Insecta - Thysanoptera - Phlaeothripidae
Cephenothrips propelaticeps n. sp. Schliephake 2005
Schliephake 2005 1 specimen
Hoffeins 1477/4 (1 measurement)
Prohaplothrips iunctostylosus n. gen., n. sp. Schliephake 2000
Schliephake 2000 1 specimen
Hoffeins 1244/2 (1 measurement)
Necrothrips major n. sp. Schliephake 2000
Schliephake 2000 1 specimen
Hoffeins 1244/1 (1 measurement)
see common names

Geography
Country:Germany State/province:Halle
Coordinates: 51.6° North, 12.4° East (view map)
Paleocoordinates:50.4° North, 8.0° East
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period:Paleogene Epoch:Eocene
Stage:Priabonian 10 m.y. bin:Cenozoic 3
Key time interval:Priabonian
Age range of interval:37.71000 - 33.90000 m.y. ago
Stratigraphy
Stratigraphy comments: Bitterfeld amber occurs in the "Bernsteinschluff" Horizon, i.e. a part of the Upper "Bitterfelder Glimmersand" in the upper part of the Cottbus Formation. Detailed biostratigraphic investigations favor an uppermost Chattian age for the unit. Originally assigned a Miocene age, and then thought to be redeposited Baltic amber. The chemical composition of Bitterfeld amber is distinct from true Baltic amber (e.g., Sodhi et al., 2013). Hydrogen isotopes indicate that Baltic and Bitterfeld ambers have distinct geographic sources, but carbon isotopes and insects suggest Baltic and Bitterfeld ambers are coeval in age (Wolfe et al., 2016).
Lithology and environment
Primary lithology: amber
Environment:terrestrial indet.
Taphonomy
Modes of preservation:body,soft parts,original chitin,amber
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Reason for describing collection:taxonomic analysis
Collectors:C. Hoffeins, H.W. Hoffeins
Collection method comments: Will be deposited in the collection of the Deutsches Entomologisches Institut, Eberswalde, Germany (DEI).
Metadata
Also known as:Saxonian amber
Database number:124273
Authorizer:M. Clapham Enterer:M. Clapham, A. Noshirvan
Modifier:M. Clapham Research group:paleoentomology
Created:2012-02-10 04:39:24 Last modified:2016-04-23 00:29:20
Access level:the public Released:2012-02-10 04:39:24
Creative Commons license:CC BY
Reference information

Primary reference:

40047. R. Wagner, C. Hoffeins, and H. W. Hoffeins. 2000. A fossil nymphomyiid (Diptera) from the Baltic and Bitterfield amber. Systematic Entomology 25:115-120 [M. Clapham/M. Clapham]

Secondary references:

54629 V. I. Alekseev. 2014. A new species of Triplax Herbst, 1793 (Coleoptera: Erotylidae) from Bitterfeld amber. Baltic Journal of Coleopterology 14:171-177 [M. Clapham/M. Clapham]
60670 V. I. Alekseev and A. Bukejs. 2016. New Zopheridae (Coleoptera: Tenebrionoidea) from Baltic amber. Zootaxa 4178:409-427 [M. Clapham/M. Clapham]
56147 V. I. Alekseev and T. L. Grzymala. 2015. New Aderidae (Coleoptera: Tenebrionoidea) from Baltic and Bitterfeld amber. Zootaxa 3956:239-257 [M. Clapham/M. Clapham]
64714 V. I. Alekseev and W. Tomaszewska. 2018. New handsome fungus beetles (Coleoptera: Coccinelloidea: Anamorphidae, Endomychidae) from European amber of the Upper Eocene . Palaeontologia Electronica 21(1.6A):1-23 [M. Clapham/M. Clapham]
57520 C. Bartel, M. Konikiewicz, J. Makol, A. Wohltmann, and J. A. Dunlop. 2015. Smaridid mites in Baltic and Bitterfeld amber, with notes on the fossil record of terrestrial Parasitengona (Trombidiformes: Prostigmata). Annales Zoologici 65:641-659 [M. Clapham/M. Clapham]
61843 B. V. Brown. 2016. Fossil evidence of social insect commensalism in the Phoridae (Insecta: Diptera). Journal of Systematic Palaeontology 15:276-286 [M. Clapham/M. Clapham]
60834 A. Bukejs, M. Biondi, and V. I. Alekseev. 2016. New records and species of Crepidodera Chevrolat (Coleoptera: Chrysomelidae) in Eocene European amber, with a brief review of described fossil beetles from Bitterfeld amber. Zootaxa 4193:390-400 [M. Clapham/M. Clapham]
79079 M. L. Gimmel and K. Szawaryn. 2020. A new genus-level and two new species-level synonyms in the extinct genus Neolitochropus Lyubarsky & Perkovsky (Coleoptera: Cyclaxyridae). Zootaxa 4894:598-599 [M. Clapham/M. Clapham]
55279 J. Háva and V. I. Alekseev. 2015. Contribution to the palaeofauna of Dermestidae (Coleoptera) from Baltic and Bitterfeld amber. Zoology and Ecology 25:154-156 [M. Clapham/M. Clapham]
55455 E. Heiss. 2013. Erstnachweis von Rindenwanzen (Aradidae) in Bitterfelder Bernstein (Insecta, Heteroptera). Linzer Biologische Beiträge 45:741-753 [M. Clapham/M. Clapham]
50252 C. Hoffeins, H. W. Hoffeins, and R. Wagner. 1997. Beschreibung einer Art der Gattung Heleodromia (Diptera, Empididae, Hemerodromiinae) aus dem Bitterfelder Bernstein. Studia Dipterologica 4:441-446 [M. Clapham/M. Clapham]
49473 C. Hoffeins, H. W. Hoffeins, and M. Jaschhof. 2010. First record of Antlemon Loew (Dipera: Keroplatidae) in Baltic amber. Studia Dipterologica 17:143-149 [M. Clapham/M. Clapham]
50014 C. Hoffeins and A. J. Woźnica. 2013. Description of a new species Eopseudopomyza szadziewskii sp. n. and redescription of E. kuehnei Hennig, 1971 from Eocene Baltic amber (Diptera: Pseudopomyzidae). Polish Journal of Entomology 82:317-325 [M. Clapham/M. Clapham]
45473 A. G. Kirejtshuk and A. S. Kurochkin. 2010. New species of sap beetles (Coleoptera: Nitidulidae: Nitidulini) from the Baltic and Bitterfeld ambers. Paleontological Journal 44:53-67 [M. Clapham/M. Clapham]
54131 J. Koteja. 2008. Xylococcidae and related groups (Hemiptera: Coccinea) from Baltic amber. Prace Muzeum Ziemi 49:19-56 [M. Clapham/M. Clapham/M. Clapham]
48707 W. Krzemiński. 2007. A revision of Eocene Bittacidae (Mecoptera) from Baltic amber, with the description of a new species. African Invertebrates 48:153-162 [M. Clapham/M. Clapham/M. Clapham]
60493 G. Y. Lyubarsky and E. E. Perkovsky. 2016. A new genus, Neolitochropus (Coleoptera: Cucujoidea: Phalacridae), from the Upper Eocene Bitterfeld amber. Russian Entomological Journal 25:249-253 [M. Clapham/M. Clapham]
62089 G. Y. Lyubarsky and E. E. Perkovsky. 2017. Serramorphus, a new genus of Erotylidae from Eocene amber (Coleoptera: Clavicornia) from late Eocene Bitterfeldian amber. Russian Entomological Journal 26:37-40 [M. Clapham/M. Clapham]
66936 G. Y. Lyubarsky and E. E. Perkovsky. 2018. Microzavaljus, a second extinct genus of pleasing fungus beetles (Coleoptera: Erotylidae) from Bitterfeld Amber. Russian Entomological Journal 27:271-276 [M. Clapham/M. Clapham]
67619 G. Y. Lyubarsky and E. E. Perkovsky. 2018. First species of Atomaria (Coleoptera: Clavicornia: Cryptophagidae) from Bitterfeld amber and several notes on Miocene and Cretaceous Cryptophagidae. Russian Entomological Journal 27:381-385 [M. Clapham/M. Clapham]
79462 W. Mey. 2019. Die Urmotten (Lepidoptera, Micropterigidae) des Bitterfelder Bernsteins: Beschreibung neuer Gattungen und Arten. Entomologische Nachrichten und Berichte 63:177-185 [M. Clapham/M. Clapham]
46422 J. Roháček. 2013. New amber fossil Anthomyzidae (Diptera): an unexpected Eocene diversity. Journal of Systematic Palaeontology 11:431-473 [M. Clapham/M. Clapham/M. Uhen]
51142 G. Schliephake. 1997. Beitrag zur Kenntnis fossiler Fransenflügler (Thysanoptera) aus dem Bernstein des Tertiär des Bitterfelder Raumes (3. Beitrag: Thripidae, Panchaetothripinae). Entomologische Nachrichten und Berichte 41:66-67 [M. Clapham/M. Clapham]
40610 G. Schliephake. 1999. Fossil Thrips (Thysanoptera, Insecta) of the Baltic (North- and Baltic Sea) and Saxonian (Bitterfeldian) Tertiary Amber from the Collections of Hoffeins. Deutsche Entomologische Zeitschrift 46(1):83-100 [M. Clapham/A. Noshirvan]
50231 G. Schliephake. 2001. Weitere neue Funde fossiler Fransenflügler aus dem Baltischen Bernstein (Insecta, Thysanoptera). Mitteilungen aus dem Geologisch-Paläontologischen Institut der Universität Hamburg 85:197-201 [M. Clapham/M. Clapham]
51551 G. Schliephake. 2005. Weitere fossile Fransenflugler (Thysanoptera, Ins.) der Sammlung Hoffeins. Annals of the Upper Silesian Museum, Entomology 13:61-69 [M. Clapham/M. Clapham]
79377 K. Szawaryn. 2021. The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles. Zoological Journal of the Linnean Society 193:1294-1309 [M. Clapham/M. Clapham]
67519 M. Wojtoń, I. Kania, and K. Kopeć. 2018. Sylvicola Harris, 1780 (Diptera: Anisopodidae) in the Eocene resins. Annales Zoologici 68:849-866 [M. Clapham/M. Clapham/M. Clapham]
47821 J. Wunderlich. 2004. Fossil spiders of the family Uloboridae (Araneae) in Baltic and Dominican amber. In J. Wunderlich (ed.), Beiträge zur Araneologie 3:851-886 [M. Clapham/M. Clapham/M. Clapham]
47828 J. Wunderlich. 2004. The fossil spiders (Araneae) of the family Cyatholipidae in Baltic amber. In J. Wunderlich (ed.), Beiträge zur Araneologie 3:1155-1188 [M. Clapham/M. Clapham]
47837 J. Wunderlich. 2004. Fossil spiders of the family Dictynidae s. l., including Cryphoecinae and Hahniinae in Baltic and Dominican amber and copal from Madagascar, and on selected extant Holarctic taxa, with new descriptions and diagnoses. In J. Wunderlich (ed.), Beiträge zur Araneologie 3:1380-1482 [M. Clapham/M. Clapham]