PBDB Collection

Phosphorites du Quercy (for taxa with non-specific locality data): Late/Upper Eocene - Early/Lower Miocene, France

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Reptilia - Squamata

	Squamata indet. Oppel 1811	Georgalis et al. 2021
	A ?sacral vertebra (NHMW 2019/0095/0002).
Reptilia - Lacertidae

	Mediolacerta sp. Augé 2005	Georgalis et al. 2021
	A left dentary (NHMW 2019/0050/0001).

	Mediolacerta roceki Augé 2005		19 specimens
	twelve right dentaries, five left dentaries, a left dentary with splenial; MNHN QU 17728, a mostly complete right mandible. Most likely Oligocene in age.

	Plesiolacerta lydekkeri (Hoffstetter 1942)		57 specimens
	Iguana europaea
	MNHN QU 17179, seven left dentaries; MNHN QU 17727, four right dentaries; QU 17166, sixteen left maxillae; [no cat no.] twelve right maxillae; QU 17167, eleven dorsal vertebrae; two cervical vertebrae, one caudal vertebra; QU 17168, three parietals, QU 17745, one premaxilla. Age range of other collections: mid Eocene-early Oligocene.

	Dracaenosaurus croizeti Gervais 1848		1 specimen
	MNHN 1903-20, left dentary (previously holotype for Dracaenosaurus sauvegi). Other collections are from the late Oligocene.

	Quercycerta maxima Augé 2005		10 specimens
	MNHN QU 17729 (paratype), right dentary; additional material: three left dentaries, five right dentaries, fragmentary left maxilla. Other collections are mid-Eocene in age.

	Pseudeumeces sp. Hoffstetter 1944	Georgalis et al. 2021
	A left maxilla (NHMW 2019/0051/0004); a left dentary (NHMW 2019/0051/0003); a right dentary (NHMW 2019/0051/0005).

	Pseudeumeces kyrillomethodicus n. sp. Georgalis et al. 2021	Georgalis et al. 2021
	Holotype: NHMW 2019/0051/0001, left dentary; NHMW 2019/0051/0002, right dentary

	Pseudeumeces cadurcensis n. gen., n. sp. Filhol 1877		109 specimens
	MNHN possibly lost holotype (figured in Filhol 1877), dentary; MNHN QU 17169, complete right mandible; left dentary; six maxillae; MNHN QU 17180, hundred dentaries. Probably Oligocene in age.
Reptilia - Teiidae

	Brevisaurus smithi n. gen., n. sp. Augé 2005		5 specimens
	MNHN QU 17742 (holotype), right dentary; QU 17741, right maxilla; additional fragment of right maxilla, left dentary, partial right dentary. Other collections are mid-Eocene in age.
Reptilia - Scincidae

	Ayalasaurus tenuis n. gen., n. sp. Augé 2005		5 specimens
	MNHN QU 17190 (holotype), left dentary with splenial and start of angular; left dentary; MNHN QU 17191,three right dentaries. Certainly from the late Eocene.
Reptilia - Cordylidae

	Pseudolacerta quercyini Augé 2005		2 specimens
	?a fragmentary left dentary, ?a fragmentary right dentary. Probably mid-late Eocene in age.

	Pseudolacerta mucronata n. gen., n. sp. Filhol 1877		5 specimens
	Holotype (MNHN, presumed lost), one dentary; MNHN QU 17159, right dentary with broken posterior; additional three right dentaries. Most likely from the late Eocene.
Reptilia

	Anguimorpha indet. Fürbringer 1900	Georgalis et al. 2021
	Five presacral vertebrae (NHMW 2019/0046/0003- NHMW 2019/0046/0007); a partial pectoral girdle (NHMW 2019/0095/0001).

	Eurheloderma gallicum n. gen., n. sp. Hoffstetter 1957		22 specimens
	MNHN QU 17746 (holotype), right maxilla; additional MNHN material: five maxillae, one left dentary, one left pterygoid, two parietals, five dorsal vertebrae, two posterior cervical vertebrae, two lumbar vertebrae; BMNH R3487, a right dentary; EMP collections: one parietal; FSM coll.: one parietal. Late Eocene (or possibly Oligocene?).
Reptilia - Palaeovaranidae

	Palaeovaranus sp. Zittel 1887	Georgalis et al. 2021
	A left maxilla (NHMW 2019/0048/0002); a left dentary (NHMW 2019/0058/0054); two right dentaries (NHMW2019/0058/0055 and NHMW 2019/0058/0056); 35 presacral verte- brae (NHMW 2019/0058/0001-NHMW 2019/0058/0009, NHMW 2019/0058/0011-NHMW 2019/0058/0027, NHMW 2019/0058/0030, NHMW 2019/0058/0041-NHMW 2019/0058/0046, NHMW 2019/0058/0051, and NHMW 2019/0067/0001); two articulated sacral vertebrae (NHMW 2019/0058/0028); two sacral vertebrae (NHMW 2019/0058/0029 and NHMW 2019/0058/0047); 16 caudal vertebrae (NHMW 2019/0058/0031-NHMW 2019/0058/0040, NHMW 2019/0058/0048-NHMW 2019/0058/0050, NHMW 2019/0058/0052, NHMW 2019/0058/0053, and NHMW 2019/0067/0002).

	Palaeovaranus lismonimenos n. sp. Georgalis et al. 2021	Georgalis et al. 2021
	Holotype: An almost complete parietal (NHMW 2019/0047/0001); A complete parietal (MNHN.F.QU17177) and a partial parietal (UM BFI 1873), both from juvenile individuals. Tentatively also, two frontals (MNHN.F.QU17175 and UM PRA 8).

	Palaeovaranus cayluxi Zittel 1887	Georgalis et al. 2021
	Two almost complete parietals (NHMW 2019/0048/0001 and MNHN.F.QU17176).
Reptilia - Necrosauridae

	Necrosaurus eucarinatus Kuhn 1940		3 specimens
	MNHN QU 17177, one frontal, two parietals. Likely Eocene age.

	Necrosaurus cayluxi n. gen., n. sp. Filhol 1873		18 specimens
	MNHN fig 434 in Filhol 1877 (holotype), dentary; MNHN QU 17172, right dentary; three incomplete right dentaries; one incomplete left dentary; two incomplete maxillae (one left, one right); fragment of jaw; one premaxilla; MNHN QU 17176, parietal; two additional parietals; MNHN QU 17175, frontal; two additional frontals; QU 17173, part of the posterior of a mandible; QU 17174, right quadrate; a number of dorsal and caudal vertebrae (BMNH). Some material from BSPHM collections in Fejervary 1935. Likely late Eocene in age.
Reptilia - Anguidae

	Placosaurus sp. Gervais 1852		3 specimens
	Part of the anterior of a right dentary, part of the anterior of a left dentary, a premaxilla. Likely mid-late Eocene in age. three presacral vertebrae (NHMW 2019/0046/0001, NHMW 2019/0046/0002, and possibly also NHMW 2019/0046/0008); one caudal vertebra (NHMW 2019/0046/0009)

	Placosaurus rugosus Gervais 1852		4 specimens
	MNHN QU 16564, complete right mandible (holotype of Placosaurus europaeus in Filhol 1877); some osteoderms(?); short maxilla fragment; MNHN QU 17178, complete right mandible. Other collections are mid-late Eocene in age.

	Melanosaurini indet. Sullivan 1979	Georgalis et al. 2021
	Four presacral vertebrae (NHMW 2019/0094/0001- NHMW 2019/0094/0003 and NHMW 2019/0094/0005); one sacral vertebra (NHMW 2019/0094/0004).

	Anguinae indet. Gray 1825	Georgalis et al. 2021
	Two presacral vertebrae (NHMW 2019/0093/0001 and NHMW 2019/0093/0002).
Reptilia

	Saniwa sp. Leidy 1870	Georgalis et al. 2021
	Three presacral vertebrae (NHMW 2019/0065/0001-NHMW 2019/0065/0003).

	Placosaurus quercyi n. sp. Filhol 1882		9 specimens
	recombined as Paraplacosauriops quercyi
	MNHN QU 16569 (neotype), right dentary; QU 16565, two left dentaries; QU 17730, three right dentaries; a mandible; two incomplete maxillae. Certainly from the middle or late Eocene. Additional: YPM-PU 11494 from reference 59390, nearly complete right maxilla: it is said to come from "Caylus, Tarn-et-Garonne", but this could mean it is from a number of established Quercy localities.

	Paraplacosauriops quercyi (Filhol 1882)	Georgalis et al. 2021
	An incomplete left maxilla (NHMW 2019/0049/0003); a right dentary (NHMW 2019/0049/0001); a left dentary (NHMW 2019/0049/0002).

	Omoiotyphlops priscus n. gen., n. sp. (de Rochebrune 1884)		1 specimen
	synonym of Omoiotyphlops
	MNHN QU 16573 (holotype), five associated vertebrae.
Reptilia - Gekkonidae

	Cadurcogekko piveteaui n. gen., n. sp. Hoffstetter 1946		12 specimens
	MNHN QU 17163 and QU 17164 (syntypes), left dentary and incomplete maxilla, respectively; seven dentaries, two maxillae, one frontal. Probably mid-late Eocene in age.

	Cadurcogekko cf. piveteaui Hoffstetter 1946	Georgalis et al. 2021
	A frontal (NHMW 2019/0052/0002); a left maxilla (NHMW 2019/0052/0001).
Reptilia - Iguanidae

	Geiseltaliellus lamandini n. sp. (Filhol 1877)		12 specimens
	MNHN No. QU 17739 (holotype), incomplete right mandible with dentary, splenial, coronoid and small part of supra-angular and angular; additional material: seven right dentaries, three left dentaries, and ?a frontal (MNHN QU 17162). Most likely late Eocene in age.
Reptilia - Agamidae

	Quercygama galliae n. gen., n. sp. (Filhol 1877)		1 specimen
	MNHN QU16560 (holotype), mandible. Most likely from the early Oligocene.
Reptilia

	Uromastyx europaeus n. sp. Moody 1980		10 specimens
	MNHN No. QU 17160 (holotype), right dentary; MNHN QU 17161, right and left dentaries; MNHN QU 17183-17189, three incomplete dentaries and four incomplete maxillae. Most likely from the Oligocene.
Reptilia - Colubridae

	Natrix mlynarskii n. sp. Rage 1988	Rage 1988	2 specimens
	MNHN QU 17181 (holotype), one mid-trunk vertebra; MNHN QU 17182, one mid-trunk vertebra; Rage notes that since a Mas de Got specimen exists these specimens are most likely Oligocene in age
Reptilia - Boidae

	Paleryx cayluxi n. sp. De Stefano 1905	Rage 2006

	Plesiotortrix edwardsi n. sp. de Rochebrune 1884	Rage 2006

	Palaeopython cadurcensis n. sp. (Filhol 1877)	Rage 2006
	original and current combination Python cadurcensis

	Palaeopython filholi n. gen., n. sp. de Rochebrune 1880	Rage 2006	1 specimen
	MNHN QU 322 (holotype), vertebra (see also Rage 1974, ref. 16854)

	Palaeopython neglectus n. sp. de Rochebrune 1884	Rage 2006
Reptilia - Testudines - Testudinidae

	Testudo phosphoritarum n. sp. Rage 1988	Rage 2006
Reptilia - Cariamiformes - Cariamidae

	Ameghinornis n. gen. minor (Gaillard 1939)	Mourer-Chauviré 1981
Reptilia - Strigiformes - Sophiornithidae

	Palaeotyto cadurcencis n. gen., n. sp.	Mourer-Chauviré 1987

	Sophiornis quercynus n. gen., n. sp. Mourer-Chauviré 1987	Mourer-Chauviré 1987

	Palaeobyas cracrafti n. gen., n. sp. Mourer-Chauviré 1987	Mourer-Chauviré 1987
Reptilia - Strigiformes - Strigidae

	Selenornis n. gen. henrici (Milne-Edwards 1892)	Mourer-Chauviré 1987
	invalid subgroup of Strigidae

	Necrobyas medius n. sp. Mourer-Chauviré 1987	Mourer-Chauviré 1987
	recombined as Prosybris medius
Amphibia - Temnospondyli - Urodela

	Megalotriton filholi n. sp. Zittel 1890	Rage 2006

	Megalotriton portisi n. sp. De Stefano 1903	Rage 2006

	Phosphotriton sigei n. gen., n. sp. Tissier et al. 2016	Tissier et al. 2016
	MNHN.F.QU 17755 (holotype)
Amphibia - Temnospondyli

	Heteroclitotriton zitteli n. sp. De Stefano 1903	Rage 2006

	Rana plicata n. sp. Filhol 1876	Rage 2006
	synonym of Thaumastosaurus gezei

	Thaumastosaurus bottii n. sp. De Stefano 1903	Rage 2006
Amphibia - Temnospondyli - Bufonidae

	Bufo serratus n. sp. Filhol 1876	Rage 2006
see common names

Geography

Country:	France	State/province:	Languedoc-Roussillon-Midi-Pyrénées	County:	Lot
Coordinates:	44.3° North, 1.6° East (view map)
Paleocoordinates:	43.9° North, 1.1° West
Basis of coordinate:	estimated from map
Geographic resolution:	basin

Time



Key time interval:	Late/Upper Eocene - Early/Lower Miocene
Age range of interval:	37.71000 - 15.98000 m.y. ago

Stratigraphy

Formation:	Quercy Phosphorites
Stratigraphic resolution:	formation
Stratigraphy comments: General collection for specimens from the Phosphorites du Quercy with no specific locality or age data, particularly from older museum collections.

Lithology and environment

Primary lithology:	phosphorite
Environment:	terrestrial indet.

Taphonomy

Modes of preservation:	body
Size of fossils:	macrofossils,mesofossils

Collection methods and comments

Collection methods:	survey of museum collection
Reason for describing collection:	taxonomic analysis

Metadata

Also known as:	Quercy Phosphorites
Database number:	179194
Authorizer:	P. Barrett, P. Mannion, P. Holroyd, M. Uhen	Enterer:	T. Cleary, G. Varnham, P. Holroyd, M. Uhen
Modifier:	G. Varnham	Research group:	vertebrate
Created:	2016-06-03 10:59:07	Last modified:	2022-05-03 08:14:55
Access level:	the public	Released:	2016-06-03 10:59:07
Creative Commons license:	CC BY

Reference information

Primary reference:

59290.

M. L. Augé. 2005. Evolution des lézards du Paléogène en Europe. Mémoires du Muséum national d'histoire naturelle 192:1-369 [P. Barrett/T. Cleary/T. Cleary]

Secondary references:

81564		G. L. Georgalis, A. Čerňanský, and J. Klembara. 2021. Osteological atlas of new lizards from the Phosphorites du Quercy (France), based on historical, forgotten, fossil material. In J. -S. Steyer, M. L. Augé, G. Métais (eds.), Memorial Jean- Claude Rage: A life of paleo-herpetologist. Geodiversitas 43(9):219-293 [P. Mannion/G. Varnham]
77097		C. Mourer-Chauviré. 1981. Première indicaition de la Présence de phorusracidés, famille d'oiseaux géants d'Amérique du sud, dans le Tertiaire Euroéen: Ameghinornis nov. gen. (Aves, Ralliformes) des Phosphorites du Quercy, France. Geobios 14(5):637-647 [M. Uhen/M. Uhen]
77082		C. Mourer-Chauviré. 1987. The owls (Aves: Strigiformes) of Phosphorites du Quercy (France): Systematics, biostratigraphy, and paleobiogeography. Documents des laboratories des géologie, Lyon 99:89-135 [M. Uhen/M. Uhen]
60167		J.-C. Rage. 1988. The oldest known colubrid snakes. The state of the art. Acta Zoologica Cracoviensia 31(13):457-474 [P. Barrett/T. Cleary]
60203		J.-C. Rage. 2006. The lower vertebrates from the Eocene and Oligocene of the Phosphorites du Quercy (France): an overview. Strata 1(13):161-173 [P. Barrett/T. Cleary]
63538		J. Tissier, J.-C. Rage, R. Boistel, V. Fernandez, N. Pollet, G. Garcia, and M. Laurin. 2016. Synchrotron analysis of a ‘mummified’ salamander (Vertebrata: Caudata) from the Eocene of Quercy, France. Zoological Journal of the Linnean Society 177:147-164 [P. Holroyd/P. Holroyd]