Ksar Metlili (KM-A1): Late Tithonian - Early Berriasian, Morocco
collected by Lasseron et al 2010
List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
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unclassified
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Actinopterygii indet.
(Cope 1887)
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| "diverse elements", no specimen no.s given | ||||||||||
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Actinopteri
- Pycnodontiformes
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Pycnodontiformes indet.
Berg 1937
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2 elements | |||||||||
| KM-A1-58, teeth | ||||||||||
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Actinopteri
- Osteoglossiformes
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Osteoglossiformes indet.
Berg 1940
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| no specimen nos in supplementary info | ||||||||||
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Actinopteri
- Siluriformes
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? Siluriformes indet.
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1 element | |||||||||
| "or ? Polypteriformes", KM-A1-40, fin spine | ||||||||||
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Actinopteri
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Ginglymodi indet.
(Cope 1871)
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3495 elements | |||||||||
| KM-A1-41, fin rays; KM-A1-42, teeth; 46, scales; 47, scale; 49, scales; 50, scales | ||||||||||
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Dipnoi
- Ptychoceratodontidae
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cf. Ptychoceratodontidae indet.
Martin 1982
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7 elements | |||||||||
| KM-A1-43, teeth | ||||||||||
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unclassified
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Tetrapoda indet.
Jaekel 1911
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50 elements | |||||||||
| KM-A1-52, diverse elements - "Lissamphibia? Squamata?" | ||||||||||
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Mammalia
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Mammalia indet.
Linnaeus 1758
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2 elements | |||||||||
| KM-A1-16, incisor tooth; KM-A1-17, left dentary; | ||||||||||
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? Mammalia indet.
Linnaeus 1758
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8 elements | |||||||||
| KM-A1-36, mandibular fragments, -37, phalanx | ||||||||||
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Mammalia
- Eutriconodonta
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Eutriconodonta indet.
Kermack et al. 1973
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1 element | |||||||||
| KM-A1-3, tooth | ||||||||||
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Mammalia
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Dryolestoidea indet.
(Butler 1939)
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1 element | |||||||||
| KM-A1-1, left lower molar | ||||||||||
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Mammalia
- Dryolestida
- Donodontidae
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Donodon perscriptoris
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1 element | |||||||||
| KM-A1-2, left upper molar | ||||||||||
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Mammalia
- Symmetrodonta
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"Symmetrodonta" indet.
(Simpson 1925)
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| no specimen nos in supplementary info | ||||||||||
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Reptilia
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Crocodylomorpha indet.
(Hay 1930)
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775 elements | |||||||||
| KM-A1-25, osteoderms; KM-A1-26, 27, 30 to 33, 35, all teeth | ||||||||||
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Atoposauridae indet.
(Gervais 1871)
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385 elements | |||||||||
| KM-A1-28, 29, 34, all teeth | ||||||||||
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Reptilia
- Teleosauridae
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Teleosauridae indet.
Geoffroy 1831
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1 element | |||||||||
| KM-A1-53, tooth | ||||||||||
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Reptilia
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? Dinosauria indet.
(Owen 1842)
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2 elements | |||||||||
| KM-A1-14, phalanges | ||||||||||
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Theropoda indet.
(Marsh 1881)
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23 elements | |||||||||
| KM-A1-9, claw (pedal?); KM-A1-11, claws; 12 to 13, both teeth; 55, claw (pedal?), 56, claw (ungual?) | ||||||||||
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Reptilia
- Dromaeosauridae
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Dromaeosauridae indet.
Colbert and Russell 1969
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1 element | |||||||||
| KM-A1-10, tooth | ||||||||||
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Reptilia
- Pterosauria
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Pterosauria indet.
(Kaup 1834)
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188 elements | |||||||||
| KM-A1-38 to 39, both teeth. Not mentioned in main text but listed in this locality in the supp info | ||||||||||
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Reptilia
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Choristodera indet.
(Cope 1876)
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1 element | |||||||||
| KM-A1-19, dorsal vertebra | ||||||||||
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Sphenodontia indet.
(Jaekel 1910)
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| no specimen nos in supplementary info | ||||||||||
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Testudinata indet.
(Oppel 1811)
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263 elements | |||||||||
| KM-A1-18, KM-A1-20 to 24, all osteoderms | ||||||||||
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Reptilia
- Squamata
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Squamata indet.
Oppel 1811
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3 elements | |||||||||
| KM-A1-5, dorsal vertebra; KM-A1-6, humerus, 8, dorsal vertebra | ||||||||||
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Reptilia
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Scincomorpha indet.
Camp 1923
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1 element | |||||||||
| KM-A1-7, left dentary (fragmentary) | ||||||||||
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Amphibia
- Temnospondyli
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Anura indet.
(Fischer von Waldheim 1813)
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34 elements | |||||||||
| KM-A1-15, dorsal vertebra; KM-A1-51, "diverse elements" | ||||||||||
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Amphibia
- Temnospondyli
- Albanerpetontidae
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Anoualerpeton unicus
Gardner et al. 2003
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1 element | |||||||||
| KM-A1-4, right dentary | ||||||||||
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Chondrichthyes
- Hybodontiformes
- Lonchidiidae
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Lonchidion (Lissodus) marocensis
(Duffin and Sigogneau-Russell 1993)
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2 elements | |||||||||
| KM-A1-54, teeth | ||||||||||
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Chondrichthyes
- Hybodontiformes
- Hybodontidae
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Egertonodus basanus
(Egerton 1845)
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21 elements | |||||||||
| KM-A1-44 and KM-A1-45, both teeth | ||||||||||
| see common names | ||||||||||
Geography
| Country: | Morocco | State/province: | Figuig |
| Coordinates: | 32.7° North, 3.2° West (view map) | ||
| Paleocoordinates: | 16.2° North, 9.7° East (Wright 2013) | ||
| Basis of coordinate: | based on nearby landmark | ||
| Geographic resolution: | small collection |
Time
| Period: | Jurassic - Cretaceous | Epoch: | Late Jurassic - Early Cretaceous |
| Stage: | Tithonian - Berriasian | 10 m.y. bin: | Jurassic 6 - Cretaceous 1 |
| Key time interval: | Late Tithonian - Early Berriasian | ||
| Age range of interval: | 146.3 - 142.7 m.y. ago |
Stratigraphy
| Formation: | Ksar Metlili | ||||
| Local section: | KM | Local bed: | A1 | ||
| Local order: | no particular order | ||||
| Stratigraphy comments: original Fm name "Couches Rouges". Localities KM-A1, A2, B', C, D1 and D2 are fossiliferous loci separated by a few metres in the same calcareous fossiliferous sandstone bed. | |||||
Lithology and environment
| Primary lithology: | lenticular,fine,medium,red calcareous sandstone |
| Includes fossils? | Y |
| Lithology description: The sedimentary facies of the Ksar Metlili Formation are similar to those of the continental lower beds of the Anoual Formation (fine to-medium-grained sandstones, overturned cross-stratifications, ripple marks, root traces). | |
| Environment: | deltaic indet. |
| Geology comments: The Ksar Metlili Formation was likely deposited in a river-dominated deltaic plain, close to the sea. All the vertebrate-bearing beds sampled in the various KM loci show this kind of transitional facies... The KM microvertebrate lens is intercalated between two marine horizons. There is a mixing of continental (wood, mammals, dinosaurs, etc.), brackish (bivalves) and undisputedly marine (echinoderms, coral, brachiopods, etc.) forms in this horizon, which suggests a near-shore, deltaic depositional environment, probably influenced by eustatic variations. | |
Taphonomy
| Modes of preservation: | body |
| Degree of concentration: | concentrated |
| Size of fossils: | macrofossils,mesofossils |
| Preservation of anatomical detail: | variable |
| Articulated whole bodies: | none |
| Associated major elements: | none |
| Disassociated major elements: | many |
| Disassociated minor elements: | many |
| Fragmentation: | frequent |
Collection methods and comments
| Collection methods: | bulk,chemical,sieve | ||
| Reason for describing collection: | general faunal/floral analysis | ||
| Collectors: | Lasseron et al | Collection dates: | 2010 |
| Taxonomic list comments:Lasseron et al (2019) mention the new material comprises 24,541 microvertebrate elements. Here I use the number of elements listed for each taxon, which are split between the specimen numbers listed (often 1000+ per specimen number). How these elements are split between specimens can be found in the supp info of Lasseron et al 2019. | |||
Metadata
| Also known as: | Ksar Met-Lili | ||
| Database number: | 204918 | ||
| Authorizer: | T. Cleary | Enterer: | T. Cleary |
| Modifier: | G. Varnham | Research group: | vertebrate |
| Subset of collection #: | 45726 | ||
| Created: | 2019-09-25 10:31:40 | Last modified: | 2025-02-22 15:12:02 |
| Access level: | the public | Released: | 2019-09-25 10:31:40 |
| Creative Commons license: | CC0 | ||
Reference information
Primary reference:
| 70302. | M. Lasseron, R. Allain, E. Gheerbrant, H. Haddoumi, N.-E. Jalil, G. Métais, J.-C. Rage, R. Vullo, and S. Zouhri. 2019. New data on the microvertebrate fauna from the Upper Jurassic or lowest Cretaceous of Ksar Metlili (Anoual Syncline, eastern Morocco). Geological Magazine 1-26 [T. Cleary/T. Cleary] |