Ksar Metlili (KM-B'): Late Tithonian - Early Berriasian, Morocco
collected by Lasseron et al 2010

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Actinopterygii indet. (Cope 1887)
6582 elements
KM-B'-62, 73-74, 80, scales; KM-B'-63, 67-68, 76-77, 79, all fin rays; 69, vertebrae; 81, mandibular fragments. Not mentioned in main text, from supp info.
Actinopteri - Osteoglossiformes
Osteoglossiformes indet. Berg 1940
6 elements
KM-B'-89, teeth
Actinopteri - Siluriformes
? Siluriformes indet.
67 elements
"or ?Polypteriformes". KM-B'-57, fin spine
Actinopteri
Ginglymodi indet. (Cope 1871)
3905 elements
KM-B'-58 and 60, 66, all ichthyoliths; KM-B'-59, 61, 64-65, 72, 84, all scales
Dipnoi - Ptychoceratodontidae
cf. Ptychoceratodontidae indet. Martin 1982
32 elements
KM-B'-56, teeth
unclassified
cf. Cynodontia informal indet. morphotype 1
3 elements
KM-B'-43-44, 98, single teeth
cf. Cynodontia informal indet. morphotype 2
1 element
Not mentioned in supp, possibly KM-B'-99, which is marked as "Cynodontia".
Mammalia
Mammalia indet. Linnaeus 1758
9 elements
KM-B'-31-32, both single incisor teeth; 33-34, left dentary; 35, dentary; 38, right dentary; 50, mandibular fragment. Not mentioned in main text, from supp info.
Mammalia - Eutriconodonta
Eutriconodonta indet. Kermack et al. 1973
4 elements
KM-B'-3, 6, single lower molars; 8, 30, single teeth
Mammalia
Hypomylos phelizoni Sigogneau-Russell 1992
3 elements
KM-B'-4, left lower molar; 9-10, single right lower molars
Hypomylos micros Sigogneau-Russell 1995
[no mention of specimen no.s in supp info]
Tribotherium africanum Sigogneau-Russell 1991
1 element
KM-B'-5, right upper molar
? Dryolestoidea indet. (Butler 1939)
1 element
"or ?Eutriconodonta". KM-B'-7, tooth. Not mentioned in main text, from supp info.
Mammalia - Dryolestida - Donodontidae
Donodon perscriptoris
3 elements
KM-B'-2, left lower molar; 11, right upper molar; 95, right upper molar
Mammalia - Dryolestidae
Dryolestidae indet. Marsh 1879
4 specimens
Probably the same as "Dryolestoidea" in the supplement. KM-B'-1, left lower molar; 37, teeth; 96-97, single left lower molars
Mammalia - Symmetrodonta
"Symmetrodonta" indet. (Simpson 1925)
2 elements
KM-B'-36, teeth. Not mentioned in main text, from supp info.
unclassified
Sauropsida indet. Huxley 1864
137 elements
KM-B'-87, teeth
Reptilia
Crocodylomorpha indet. (Hay 1930)
444 elements
KM-B'-40, osteoderms; KM-B'-45, mandibular fragment; 46, 52, 91, all teeth
Atoposauridae indet. (Gervais 1871)
2165 elements
KM-B'-47, 48, teeth
cf. Theriosuchus indet. Owen 1878
1 element
KM-B'-42, premaxillary
Reptilia - Teleosauridae
Teleosauridae indet. Geoffroy 1831
74 elements
KM-B'-51, teeth; KM-B'-53, tooth
Reptilia
Theropoda indet. (Marsh 1881)
22 elements
KM-B'-21, claw; KM-B'-23-25, teeth; 94, claw (ungual?)
Maniraptora indet. Gauthier 1986
1 element
KM-B'-22, tooth. Not mentioned in main text, from supp info.
Reptilia - Dromaeosauridae
Dromaeosauridae indet. Colbert and Russell 1969
4 elements
KM-B'-26, teeth. Not mentioned in main text, from supp info.
Reptilia - Pterosauria
Pterosauria indet. (Kaup 1834)
138 elements
KM-B'-54-55, teeth
Reptilia
Choristodera indet. (Cope 1876)
9 elements
KM-B'-49, dorsal vertebrae
Sphenodontia indet. (Jaekel 1910)
1 element
KM-B'-14, dorsal vertebra
Testudinata indet. (Oppel 1811)
57 elements
KM-B'-39, 41, osteoderms
Reptilia - Squamata
Squamata indet. Oppel 1811
5 elements
KM-B'-28, dorsal vertebrae; 29, jaw fragment; 90, 92, single teeth
Reptilia
Scincomorpha indet. (Camp 1923)
1 element
KM-B'-19, right dentary
Tarratosaurus anoualensis Broschinksi and Sigogneau-Russell 1996
1 element
KM-B'-20, right dentary. Not mentioned in main text, from supp info.
Amphibia - Temnospondyli
Anura indet. (Fischer von Waldheim 1813)
21 elements
KM-B'-15, right ilium; 16, scapula; 17, dorsal vertebrae; 27, dorsal vertebrae of a hatchling or neotenic; 86, long bone fragments
Amphibia - Temnospondyli - Alytidae
cf. Alytidae indet. Fitzinger 1843
15 elements
KM-B'-18, dorsal vertebrae
Amphibia - Temnospondyli - Albanerpetontidae
Anoualerpeton unicus Gardner et al. 2003
2 elements
KM-B'-12-13, left dentary
Chondrichthyes - Hybodontiformes - Hybodontidae
Egertonodus basanus (Egerton 1845)
see common names

Geography
Country:Morocco State/province:Figuig
Coordinates: 32.7° North, 3.2° West (view map)
Paleocoordinates:16.2° North, 9.7° East (Wright 2013)
Basis of coordinate:based on nearby landmark
Geographic resolution:small collection
Time
Period: Jurassic - Cretaceous Epoch: Late Jurassic - Early Cretaceous
Stage: Tithonian - Berriasian 10 m.y. bin: Jurassic 6 - Cretaceous 1
Key time interval: Late Tithonian - Early Berriasian
Age range of interval: 146.3 - 142.7 m.y. ago
Stratigraphy
Formation:Ksar Metlili
Local section:KM Local bed:B'
Local order:no particular order
Stratigraphy comments: original Fm name "Couches Rouges". Localities KM-A1, A2, B', C, D1 and D2 are fossiliferous loci separated by a few metres in the same calcareous fossiliferous sandstone bed.
Lithology and environment
Primary lithology:lenticular,fine,medium,red calcareous sandstone
Includes fossils?Y
Lithology description: The sedimentary facies of the Ksar Metlili Formation are similar to those of the continental lower beds of the Anoual Formation (fine to-medium-grained sandstones, overturned cross-stratifications, ripple marks, root traces).
Environment:deltaic indet.
Geology comments: The Ksar Metlili Formation was likely deposited in a river-dominated deltaic plain, close to the sea. All the vertebrate-bearing beds sampled in the various KM loci show this kind of transitional facies... The KM microvertebrate lens is intercalated between two marine horizons. There is a mixing of continental (wood, mammals, dinosaurs, etc.), brackish (bivalves) and undisputedly marine (echinoderms, coral, brachiopods, etc.) forms in this horizon, which suggests a near-shore, deltaic depositional environment, probably influenced by eustatic variations.
Taphonomy
Modes of preservation:body
Degree of concentration:concentrated
Size of fossils:macrofossils,mesofossils
Preservation of anatomical detail:variable
Articulated whole bodies:none
Associated major elements:none
Disassociated major elements:many
Disassociated minor elements:many
Fragmentation:frequent
Collection methods and comments
Collection methods:bulk,chemical,sieve
Reason for describing collection:general faunal/floral analysis
Collectors:Lasseron et al Collection dates:2010
Taxonomic list comments:Lasseron et al (2019) mention the new material comprises 24,541 microvertebrate elements. Here I use the number of elements listed for each taxon, which are split between the specimen numbers listed (often 1000+ per specimen number). How these elements are split between specimens can be found in the supp info of Lasseron et al 2019.
Metadata
Also known as:Ksar Met-Lili
Database number:204936
Authorizer:T. Cleary Enterer:T. Cleary
Modifier:G. Varnham Research group:vertebrate
Subset of collection #:45726
Created:2019-09-27 08:55:58 Last modified:2025-02-22 15:12:02
Access level:the public Released:2019-09-27 08:55:58
Creative Commons license:CC0
Reference information

Primary reference:
70302. M. Lasseron, R. Allain, E. Gheerbrant, H. Haddoumi, N.-E. Jalil, G. Métais, J.-C. Rage, R. Vullo, and S. Zouhri. 2019. New data on the microvertebrate fauna from the Upper Jurassic or lowest Cretaceous of Ksar Metlili (Anoual Syncline, eastern Morocco). Geological Magazine 1-26 [T. Cleary/T. Cleary]