#NEXUS
[MacClade 4.05 registered to Peter Wagner, Field Museum of Natural History]
BEGIN MacCladeStart;
Extended;
END;
BEGIN DATA;
DIMENSIONS NTAX=57 NCHAR=119;
[!Lü, J., D. M. Unwin, X. Jin, Y. Liu, and Q. Ji. 2010. Evidence for modular evolution in a long-tailed pterosaur with a pterodactyloid skull. Proceedings of the Royal Society B: Biological Sciences 277:383-389.]
FORMAT SYMBOLS= " 0 1 2 3 4 5 6 7 8 9 A B C D E" MISSING=? GAP=- ;
CHARSTATELABELS
118 FA / Triassic_4 Jurassic_1 Jurassic_2 Jurassic_3 Jurassic_4 Jurassic_5 Jurassic_6 Cretaceous_1 Cretaceous_2 Cretaceous_3 Cretaceous_4 Cretaceous_5 Cretaceous_6 Cretaceous_7 Cretaceous_8,
119 LA / Triassic_4 Jurassic_1 Jurassic_2 Jurassic_3 Jurassic_4 Jurassic_5 Jurassic_6 Cretaceous_1 Cretaceous_2 Cretaceous_3 Cretaceous_4 Cretaceous_5 Cretaceous_6 Cretaceous_7 Cretaceous_8
;
MATRIX
[ 10 20 30 40 50 60 70 80 90 100 110 ]
[ . . . . . . . . . . . ]
Outgroup 000(01)000000000000(01)0000000000000000000?000000000000000000000000000000000000?0000000000000000000000000000000?0(01)000000000??
Preondactylus 0000000000?0001000000?10000000?00??0?0000000000101000000000000??0000?000??00?0010??1?00000000101000001001?0000??1100?00
Dimorphodon 000000000000001000000010000000000000?100000000010000000000000001000?0001??1000011001000(01)0000011100000100100000000000012
Peteinosaurus ???????0?00??????????????????????????1000000000100000000000000??0?0?0001?01000011?010000000001110000010?10000??00100000
Anurognathus 00?000000100000000000?0000000000??10?100000000?0101101000000000?010?0010??10?001100200111000001101002010??0000001100066
Batrachognathus 00?000000100000000000?00000000?00?10?1000000000010000100000000??0???0010101?0001??0200??1??0??1???????1?????00??1100055
Dendrorhynchoides 00?000?001??0000??0?0??????00??0??1????00?0000?010????000000000?01000010??1000012?0200101000001102?0201???0100?01100089
Jeholopterus 00?0000001????0????00?????000??0???0???00?0000?010????00000000??01?20010??1000012?020011200?00110200101??00000?01100045
Austriadactylus 0001010020?1111110000011000000?0?0?0?1?00???0?0001000000000000??00000000?????001????00???0????1?0?00?1??????0???1????00
Campylognathoides 000101000001111110000001000000100100?11000010000000000000000000100000001401000013003000010(01)0011202000111210000000110033
Eudimorphodon 000101000001111110000001000000100100?11000010000010000000000000?00000000401100012003000(01)(01)010011(12)0(12)000101210(01)00001110000
Raeticodactylus 00010000201?11111000000100?000000100?1000?02010001000000000000010?000??????????1?0?30??01??0????0100?1?????000001????00
Scaphognathus 00010100001111111000000100000000010001000100001000111100000000??000000012010?001200400022011011101000110210000?01110166
Sordes 000101000011111110000001000000000100?1000100001000111100000000??000000012?100001300400021011011101000110210000001(01)10155
Pterorhynchus 0001010020111111101000010000000001?0?1000?00001000110100000000??0?00?001???????12???00022?1?01?1010001????0?0??????0?45
Cacibupteryx 00010000001111111010011100000000?0000??????????00?1?0?000???0????????????????????????????????????????????????????????55
Dorygnathus 000101000011111111100111000000000100?1000110001000100100000000??0000000120100001200400022010011101000110210000001110033
Nesodactylus ???????????????????00???????????????????????????????????????????0000???120100001?00400?2?0?101?20?01001020?0?????????55
Rhamphorhynchus 000102000011111111100101000000000100010001100010001001000000000100000001(12)0100001300400022011011203010010210000001110056
Darwinopterus 00010210201122110001111000000000010111000100001000000100000000??10120001301000012?01000(12)001101010000011?210100?01110145
Pterodactylus 000102000011221100010001000011010201010001000010000001000000000011120110201000012005000(01)002102110(01)00(01)011210000001111?55
Cycnorhamphus 001102000011221??0?1000?0000110102?201001100001000110100000000000100011030110001200500013021020101001011210000?0101??56
Eosipterus ????????????????????????????????????????????????????????????????11120110??11000110050000001102010100001???0000001111?89
Pterodactylus_longicollum 011112000011221??0?10?0?00001?010201010111000010000001000001001?11120110??1100010005000030?1120001001011211000?01011?56
Gnathosaurus 0111121020112211?001000100001101020101011100001000000100000100???????????????????????????????????????????????????????67
Cearadactylus 01111210001?221??0?1100?10?????0?1?2?1011100001000000000000100111110?????????????????????????????????????????????????AA
Gegepterus 0011121000112211?0010001000011010201?10?110?0010000001000001101111120??????10??1?????????0??????0100?01?????0??011???89
Ctenochasma 00111210(02)011221??001000?000011010201?1001100001000000100000110??11120110?01?00011?05000(01)0?1102010100101?210000001111?67
Pterodaustro 001112100011221??001000?000011010201?1001100001000000100000110111112?01030110001000500?000?1120101001011210000001111?9A
Istiodactylus 001101000011221??001101?000000000102010011000010000001000000011101011????0200111301611?211?012?1020010?????100002????89
Nurhachius 0011021000?122110001101100?0??0??1020100110000100000010?000001?101011?????200111?0?6?1?2???112?10200201????100??201??99
Haopterus 001101?000??221??0010??????0?????10??1001100001000000100100001??010?0???3020?111???611?1?11002110?0010????????????11?89
Ornithocheirus 010102101011221??0?1011?10010000010211010100011000000100100000???????????????????????????????????????????????????????8A
Anhanguera 010102101011221100010111100000000102110101000(01)10000001001000001101011110312011113016111(12)11211211020010101?11000020???AB
Coloborhynchus 010102101011221??0?1011?100000000102110101000110000001001000001101011110312011113016111111?11211020010101?0100002011?AB
Ludodactylus 010102100011221100010111100100000102?10?0100001000000100100000???????????????????????????????????????????????????????9A
Boreopterus 00011210(01)011221??001011?100000000102?10001000010000001001000001?0101?110???????11????1111?111211020010????0100??2011?89
Nyctosaurus 000112100011221??0?1010?100200000102010001001010000000100000001001011110102101112107?112212?23?(01)03002011120100002111?CE
Muzquizopteryx ???0???00011221??0?1?10?1002000001?2?1??????101000000010000000??010?111011?1??11??07?11?????2?????????1??20100002??1?CC
Pteranodon 000112100011221??0?1011?10020000010201000100101000000010000000110101111011200111110611110111231103001010120100001011?CD
Germanodactylus_cristatus 0001020020012211000100110000000011?1?1000100000000000001011000?101000?10??1?00001?05001130?102010(01)0010????1011011111?66
Germanodactylus_rhamphastinus 000102002001221100011011000000001101?1000100000000000001010000?101000110??1000011?05001200??020000001012?11011??1011?66
Dsungaripterus 000102002001221??0?1001?20030000110101000100000000000001011000110110111010110000?00500???0???20?0?00??123?1011011????9A
Tatal_pterosaur 000102002001221??0?1001?200300001101?10001000000000000010110001101100?????11?000200500?130?1120?0100?0????10110110???77
Noripterus ?????????0???????????????????????????1?0?1????000???000??1100?11010????????????0?00500?030?1120?0100?01????0110?1111?9A
Lonchodectes 000?????(01)0????1?????????????????????110011000(01)0000000000000000101112???????1?0?0?1050??????????????????????0011??????7B
Tapejara 100200004011221??0?1101?011300000102010001000100000000100000001011120???10110001?00500?1?0??120?1?00?0102??0000?10?1?AA
Tupandactylus 100200104011221??0?1101?0114000001?2?1000100010000000010000000???????????????????????????????????????????????????????99
Sinopterus 1002000000?1221??0?1101?011300000101?10001000100000000100000001?111201101011??011??5?0113021?20011002010211001?11011?99
Huaxiapterus 1002000040?1221??0?1100?01140000010101000100010000000010000000??1112?110??110??111050011302112001100(12)0???11001??10???99
Shenzhoupterus 000202110011221??0?1101?0115000001?2?1000100100000000010000000??11120?10??11?0011??5001230?12200110020????100????0?1?99
Chaoyangopterus 0002?2?100??221????1?????????????1???100010010?000000010000000??1112??1???1????11??????13?2122?0100020????100???10?1?99
Tupuxuara 000201103011221??0?1101?011500000102110001000000000000100000001101101110101100011105001030212200110020102?1001111011?AA
Thalassodromeus 000201103011221??0?1101?01150000010211000100000000000010000000???????????????????????????????????????????????????????AA
Quetzalcoatlus 00021210501?221??0?1001?011??????102010001001000000000100000001021121????0110001110500?1302?2300111020?????0011120?1?EE
Zhejiangopterus 000202100011221??0?1100?0110000101?2?1000100100000000010000000??21121110?011?0011?05?0123?21?3001110?010201001?12011?DD
Azhdarcho 000??????0????1???????????????????????000100???0000000100000001021121??????????1?1050????0????????10???????00111?????CD
;
END;
BEGIN ASSUMPTIONS;
OPTIONS DEFTYPE=unord PolyTcount=MINSTEPS ;
TYPESET * UNTITLED = unord: 1-117 119, strat: 118;
WTSET * UNTITLED = 1: 1-117, 0: 118-119;
END;
BEGIN TREES;
TRANSLATE
1 Outgroup,
2 Preondactylus,
3 Dimorphodon,
4 Peteinosaurus,
5 Anurognathus,
6 Batrachognathus,
7 Dendrorhynchoides,
8 Jeholopterus,
9 Austriadactylus,
10 Campylognathoides,
11 Eudimorphodon,
12 Raeticodactylus,
13 Scaphognathus,
14 Sordes,
15 Pterorhynchus,
16 Cacibupteryx,
17 Dorygnathus,
18 Nesodactylus,
19 Rhamphorhynchus,
20 Darwinopterus,
21 Pterodactylus,
22 Cycnorhamphus,
23 Eosipterus,
24 Pterodactylus_longicollum,
25 Gnathosaurus,
26 Cearadactylus,
27 Gegepterus,
28 Ctenochasma,
29 Pterodaustro,
30 Istiodactylus,
31 Nurhachius,
32 Haopterus,
33 Ornithocheirus,
34 Anhanguera,
35 Coloborhynchus,
36 Ludodactylus,
37 Boreopterus,
38 Nyctosaurus,
39 Muzquizopteryx,
40 Pteranodon,
41 Germanodactylus_cristatus,
42 Germanodactylus_rhamphastinus,
43 Dsungaripterus,
44 Tatal_pterosaur,
45 Noripterus,
46 Lonchodectes,
47 Tapejara,
48 Tupandactylus,
49 Sinopterus,
50 Huaxiapterus,
51 Shenzhoupterus,
52 Chaoyangopterus,
53 Tupuxuara,
54 Thalassodromeus,
55 Quetzalcoatlus,
56 Zhejiangopterus,
57 Azhdarcho
;
TREE * UNTITLED = [&R] (2,((4,3),((7,(8,(6,5))),((11,10),((12,9),(((14,13),(15,((19,18),(17,16)))),(20,(21,(23,((27,(28,(29,(24,(25,26))))),(22,((32,((31,30),((40,(39,38)),(37,(36,(33,(35,34))))))),((42,((44,43),(45,41))),((49,(47,(48,50))),((52,51),((55,(56,57)),(46,(54,53))))))))))))))))));
END;
BEGIN NOTES;
TEXT TAXON=2 CHARACTER=118 TEXT= Alaunian;
TEXT TAXON=3 CHARACTER=118 TEXT= Hettangian;
TEXT TAXON=3 CHARACTER=119 TEXT= 'Pliensbachian_-_Toarcian';
TEXT TAXON=4 CHARACTER=118 TEXT= Alaunian;
TEXT TAXON=5 CHARACTER=118 TEXT= Early_Tithonian;
TEXT TAXON=6 CHARACTER=118 TEXT= 'Oxfordian_-_Kimmeridgian';
TEXT TAXON=7 CHARACTER=118 TEXT= 'Barremian_-_Early_Aptian';
TEXT TAXON=8 CHARACTER=118 TEXT= 'Bathonian_-_Kimmeridgian';
TEXT TAXON=9 CHARACTER=118 TEXT= 'Alaunian_-_Sevatian';
TEXT TAXON=9 CHARACTER=119 TEXT= Sevatian;
TEXT TAXON=10 CHARACTER=118 TEXT= Early_Toarcian;
TEXT TAXON=11 CHARACTER=118 TEXT= Alaunian;
TEXT TAXON=11 CHARACTER=119 TEXT= Sevatian;
TEXT TAXON=13 CHARACTER=118 TEXT= Early_Tithonian;
TEXT TAXON=14 CHARACTER=118 TEXT= 'Oxfordian_-_Kimmeridgian';
TEXT TAXON=15 CHARACTER=118 TEXT= 'Bathonian_-_Kimmeridgian';
TEXT TAXON=16 CHARACTER=118 TEXT= 'Middle_Oxfordian_-_Late_Oxfordian';
TEXT TAXON=17 CHARACTER=118 TEXT= Early_Toarcian;
TEXT TAXON=18 CHARACTER=118 TEXT= Late_Oxfordian;
TEXT TAXON=19 CHARACTER=118 TEXT= Late_Callovian;
TEXT TAXON=19 CHARACTER=119 TEXT= Early_Tithonian;
TEXT TAXON=20 CHARACTER=118 TEXT= 'Late_Bathonian_-_Early_Kimmeridgian';
TEXT TAXON=21 CHARACTER=118 TEXT= Kimmeridgian;
TEXT TAXON=22 CHARACTER=118 TEXT= Late_Kimmeridgian;
TEXT TAXON=22 CHARACTER=119 TEXT= Early_Tithonian;
TEXT TAXON=23 CHARACTER=118 TEXT= 'Barremian_-_Early_Aptian';
TEXT TAXON=24 CHARACTER=118 TEXT= Late_Kimmeridgian;
TEXT TAXON=24 CHARACTER=119 TEXT= Early_Tithonian;
TEXT TAXON=25 CHARACTER=118 TEXT= Early_Tithonian;
TEXT TAXON=25 CHARACTER=119 TEXT= Berriasian;
TEXT TAXON=26 CHARACTER=118 TEXT= Early_Albian;
TEXT TAXON=27 CHARACTER=118 TEXT= 'Barremian_-_Early_Aptian';
TEXT TAXON=28 CHARACTER=118 TEXT= Early_Tithonian;
TEXT TAXON=28 CHARACTER=119 TEXT= Berriasian;
TEXT TAXON=29 CHARACTER=118 TEXT= Aptian;
TEXT TAXON=29 CHARACTER=119 TEXT= Albian;
TEXT TAXON=30 CHARACTER=118 TEXT= Barremian;
TEXT TAXON=30 CHARACTER=119 TEXT= Aptian;
TEXT TAXON=31 CHARACTER=118 TEXT= Aptian;
TEXT TAXON=32 CHARACTER=118 TEXT= 'Barremian_-_Early_Aptian';
TEXT TAXON=33 CHARACTER=118 TEXT= Early_Hauterivian;
TEXT TAXON=33 CHARACTER=119 TEXT= Early_Albian;
TEXT TAXON=34 CHARACTER=118 TEXT= Early_Albian;
TEXT TAXON=34 CHARACTER=119 TEXT= Cenomanian;
TEXT TAXON=36 CHARACTER=118 TEXT= 'Aptian_-_Albian';
TEXT TAXON=38 CHARACTER=118 TEXT= 'Late_Coniacian_-_Early_Santonian';
TEXT TAXON=39 CHARACTER=118 TEXT= 'Late_Turonian_-_Early_Coniacian';
TEXT TAXON=40 CHARACTER=118 TEXT= Early_Coniacian;
TEXT TAXON=40 CHARACTER=119 TEXT= Early_Campanian;
TEXT TAXON=41 CHARACTER=118 TEXT= Early_Tithonian;
TEXT TAXON=42 CHARACTER=118 TEXT= Tithonian;
TEXT TAXON=43 CHARACTER=118 TEXT= 'Late_Aptian_-_Albian';
TEXT TAXON=44 CHARACTER=118 TEXT= 'Berriasian_-_Valanginian';
TEXT TAXON=46 CHARACTER=118 TEXT= 'Late_Berriasian_-_Valanginian';
TEXT TAXON=46 CHARACTER=119 TEXT= 'Cenomanian_-_Turonian';
TEXT TAXON=47 CHARACTER=118 TEXT= Early_Albian;
TEXT TAXON=48 CHARACTER=118 TEXT= Aptian;
TEXT TAXON=49 CHARACTER=118 TEXT= Aptian;
TEXT TAXON=50 CHARACTER=118 TEXT= Aptian;
TEXT TAXON=51 CHARACTER=118 TEXT= Aptian;
TEXT TAXON=53 CHARACTER=118 TEXT= Early_Albian;
TEXT TAXON=54 CHARACTER=118 TEXT= Early_Albian;
TEXT TAXON=55 CHARACTER=118 TEXT= Late_Maastrichtian;
TEXT TAXON=56 CHARACTER=118 TEXT= Early_Campanian;
TEXT TAXON=57 CHARACTER=118 TEXT= 'Middle_Turonian_-_Late_Turonian';
END;
BEGIN MacClade;
Version 4.0 84;
LastModified -908786448;
FileSettings editor '1' '1' '118' '1';
Singles 110;
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