#NEXUS
[MacClade 4.08 registered to Peter Wagner, Field Museum of Natural History]
BEGIN MacCladeStart;
Extended;
END;
BEGIN DATA;
DIMENSIONS NTAX=79 NCHAR=71;
FORMAT SYMBOLS= " 0 1 2 3 4 5 6 7 8 9 A B C D E F G H J" MISSING=? GAP=- ;
CHARSTATELABELS
1 GL_Prom / _ fine sharp coarse,
2 Imb_GLs / _ absent present,
3 Sinus / _ absent present,
4 Sinus_Width / _ wide narrow very_narrow,
5 Sinus_Depth / _ very_deep deep moderate shallow very_shallow,
6 Sinus_Shape / _ curved straight,
7 Lunulae / _ sharp_crescent straightened_crescent,
8 ImbLunulae / _ absent present,
9 ∂ImbLun / _ _ juvenile_whorls_only all_whorls,
10 ImbLun_Type / _ deep_and_obtuse narrow_and_sharp,
11 SK / _ absent present,
12 Periph_Lira / _ absent present 10 _ absent present,
13 ApexThread / _ absent present,
14 Selen_Width / _ narrow moderate wide,
15 StrDullPL / _ sharp_threads dull_strong_threads,
16 SK_Prom / _ slight strong,
17 AL_Strength / _ Weak Same_as_P.L. Strong,
18 SeratedSK / _ absent present,
19 IntChan / _ Absent Moderate Strong,
20 Selen_Att / _ parallel abapical_hook,
21 Slit / _ Absent Present,
22 RRShape / _ Concave Sl._Concave Flat Convex Globular,
23 LRShape / _ Flat Sl._Convex Very_Convex Globular,
24 RRC / _ absent present,
25 RRCType / _ weak_thread sharp_thread thick_sharp_thread,
26 ∆RRC / _ absent present,
27 RRC_Loc / _ '~' 75Ëš_from_SK '~' 45Ëš_from_SK,
28 RRC_Att / _ perpendicular_to_ramp abapical_attitude,
29 Serr_RRC / _ thread serrated,
30 SC / _ absent present,
31 SC_Strength / _ weak strong,
32 LRC / _ absent present,
33 LRC_Type / _ protrusion actual_carina,
34 LRCType / _ thick_dull_carina very_weak_thread sharp_thread strong_sharp_thread,
35 Serr_LRC / _ plain serrated,
36 2LRC / _ one_carina two_carinae,
37 UC / _ absent dull_protrusion sharp_lirae,
38 UC_Loc / _ '~' 90Ëš_below_exhalent '~' 120Ëš_below_exhalent,
39 UCMag / _ weak strong,
40 ChanUC / _ Solid Channeled,
41 LR_Orn / _ absent present,
42 LR_Orn_Type / _ Thin_Threads Thick_Lirae,
43 LROrn_Density / _ Sparse Moderate Dense,
44 RR_Orn / _ absent present,
45 RR_Orn_Type / _ Thin_Threads Thick_Lirae,
46 RROrn_Density / _ Sparse Moderate Dense,
47 ∆RROrn / _ Constant Fading_with_Age,
48 Cren_Aperutre / _ smooth crenulate,
49 E / _ 0.04_to_0.05 0.06_to_0.08 0.085_to_0.1,
50 K / _ high moderate low,
51 T / _ low moderate high very_high,
52 ∆T / _ constant increasing_on_older_whorls,
53 Mag_∆T / _ lower_suture_point uncoiling_of_shell,
54 ß / _ ß_ '>' _100˚ ß_@_90˚ ß_@_75˚ ß_@_60˚ ß_@_45˚,
55 Theta / _ '~' 45Ëš '~' 70Ëš '~' 90Ëš,
56 Phi / _ '~' 45Ëš '~' 60Ëš '~' 90Ëš,
57 Base_Sh / _ flat round acute,
58 Extended_UR / _ Absent Present,
59 Tangent / _ Absent '~10Ëš-15Ëš' '>' 25Ëš,
60 Inclined_Base / _ Absent Present,
61 'L-R Inclination' / _ Absent Present,
62 ColLip_Sh / _ 'upside-down_"7"_shape' '"L"-shape',
63 Round_Lip / _ '"L"-shape' 'rounded_"V"-shape_pointing_->_u' 'half-circle',
64 U_Lip / _ 'upside-down_"7"' '"U"-shape',
65 Lip_Thick / _ No_Thicker Slightly_Thicker Much_Thicker Extremely_Thick,
66 Lip_Reflex / _ Absent Present,
67 PI_Thick / _ Absent Thinner Same_as_Shell,
68 Size / '<10_mm3' '<100_mm3' '<1000_mm3' '<10000_mm3',
69 Origin / _ Late_Tremadoc Early_Arenig Middle_Arenig Late_Arenig Llanvirn Llandeilo Early_Caradoc Middle_Caradoc Late_Caradoc Ashgill Early_Llandovery Late_Llandovery Early_Wenlock Middle_Wenlock Late_Wenlock Early_Ludlow Late_Ludlow Pridoli,
70 Extinction / _ Late_Tremadoc Early_Arenig Middle_Arenig Late_Arenig Llanvirn Llandeilo Early_Caradoc Middle_Caradoc Late_Caradoc Ashgill Early_Llandovery Late_Llandovery Early_Wenlock Middle_Wenlock Late_Wenlock Early_Ludlow Late_Ludlow Pridoli,
71 Genus / Outgroup Donaldiella Lophospira '"MetaLoph"' Zimmerspira Paupospira Arjamannia Ruedemannia Schizolopha Eunema Trochonema Trochonemella Gyronema Longstaffia Loxoplocus
;
MATRIX
[ 10 20 30 40 50 60 70]
[ . . . . . . . ]
Sinuopea_sweeti 11213111--1?1?-???1?1531-----1?21-?11---1--1---13221-122211113-11132110
Hormotoma_confusa 11212111--221211??111521-----1?21-?11---1--1---11241-222211111-121?2110
Ectomaria_adelina 11212111--221311??111111-----1?21-?11---1--1---11241-211111111-12221310
Pagodispira_cicelia 11212111--22211121211111-----1?21-111---1--1---11241-211111111-12221253
Lophospira_perangulata 11212111--22211121211111-----1?21-111---1--1---11231-211111111-122212A3
Lophospira_kangyaoensis 11212111--22211121211311-----1?21-111---1--1---11221-211111111-12222363
Lophospira_grabaui 21223111--22211121211211-----1?223111---1--1---12121-211111112112221362
Lophospira_milleri 21223111--22211121211212211111?223111---1--1---12121-2121111121122214A2
Proturritella_bicarinata 21223111--2121-131211212211111?223111---222223112121-212111112112221462
Pagodispira_derwiduii 11212111--22211221211111-----1?21-111---1--1---11241-211111111-1--21561
Pagodispira_dorothea 11212111--22211221211312111111?21-111---1--1---11241-211111111-1--21561
Trochonemella_knoxvillensis 21223111--22211121311212211111?2231121111--1---13312141211212211213256B
Lophospira_centralis 21224221--22211121211312211111?223111---1--1---12121-212111212112221779
Trochonemella_trochonemoides 21223111--2121-131311312211111?2221131111--1---1331?-42211212221113277A
Trochonemella_montrealensis 21233111--221111?131131221111212241122111--1---13312153221212221213277B
Trochonema_bellulum 21223111--2121-13131131221211222221131211--1---13311-53311312221113277A
Lophospira_helicteres 21224221--2221112121131231111212231121111--1---121(23)22212111212112232779
Trochonemella_n.sp._A 2123311221221111?131141231221212241122111--1---13312253321212221213277B
Lophospira_serrulata 212121122-2221112221111231111212231121111--1---112(23)22211111121-22221773
Lophospira_ventricosa 11212111--22211121211411-----1?21-111---1--1---12231-211111111-12221783
Trochonemella_notablis 2123311231221111?131141231221212241122111--1---13312253321212221213277B
Eunema_strigillatum 21224221--2121112131131231111212231121111--1---121(23)22312111112112232779
Trochonema_umbilicatum 21224111--2121-13131131221211222231132211--1---13311-5332131222111327AA
Trochonema_wilsonae 21224211--2121-13131131221211222231132211--1---13311-53321312221113277A
Gyronema_pulchellum 11234221--2121-121211312211111?22312?---1--221112121-21211121211222177C
Gyronema_semicarinatum 11234221--2121-121211312212111?22312?---221221112121-21211121211222188C
Gyronema_liljevalli 21224211--2121-13131141231211222231212221--1---13311-53321312221113188A
Lophospira_burginensis 11212111--2221112121121?111111?21-111---1--1---11231-211111111-231218A5
Lophospira_spironema 21222111--22211121211212211111?223111---213213112121-212111112112221882
Lophospira_oweni 11212111--22211211211312121111?21-111---1--1---12221-211112111-241228A5
Trochonema_madisonense 21223111--2121-13131131231111212231132211--1---13311-5331121223111338AA
Lophospira_sumnerensis 11212111--22211211211312121112221-111---1--1---12131-421112111-241229A5
Lophospira_decursa 11212111--22212121211111-----1?21-111---1--1---11231-111111111-12221994
Lophospira_conoidea 11212111--22212111211311-----1?21-111---1--1---11231-111111111-12222994
Lophospira_producta 11212111--22212121211211-----1?21-111---1--1---11241-111111111-12222994
Lophospira_Curdsville_sp. 11212111--22212121211212111111?21-111---1--1---112422111111111-12222994
Ruedemannia_humilis 21223111--22221121211422212111?1--111---1--1---13121-222211112112221997
Schizolopha_textilis 21223111--22211121212422312111?1--111---212212113121-22221111?-?--22997
Ruedemannia_lirata 11223111--22221121211422212111?1--111---2--1---13121-222211112112221977
Lophospira_tenuistriata 11212111--22211121211212211111?21-111---212212111231-211111111-23-21995
Lophospira_tropidophora 11212111--22211211211312121111?21-111---1--1---12221-211112111-241229A5
Schizolopha_moorei 21212121--22211211212321-----1?21-??1---1--1---12221-211?12111-2--?29A5
Donaldiella_filosa 11212111--22211221211311111111?21-111---1--1---11241-211111111-12221991
Lophospira_pulchella 21233111--22211231221122211111?221111---1--1---12131-323121112112221996
Lophospira_bellicarinata 21233111--22211231221122211111?221111---1--1---12131-3131211121122229A6
Donaldiella_bowdeni 11212111--22211221211312111111?21-111---1--1---11241-211111111-122229A1
Lophospira_aff._L._serrulata 212331123222211122211212312221?2242131111--1---122221333212122212121AAB
Globonema_kokeni 211???11--2121-231311311-----222131131221--1---12121-322211122311132AAA
Ruedemannia_Boda_Ls._sp. 11223111--22221121211421-----1?1--111---212212113221-222211112-?--?2AA7
'Trochonemella "G. pulchellum"' 212331123222211122211212312221?2242231111--1---122221333212122212122AAB
Trochonemella_churkini 2122311231221111?1311412312221?2241122111--1---133122433212122212132AAB
Trochonemella_reusingi 2122311231221111?1311412312221?2242122111--1---13312-433212122212132AAB
Trochonema_Alaska_sp. 21225211--2121-13131131221211222241131211--1---12221-333212122311132AAA
'Trochonema (Eunema) salteri' 21224211--2121-131311311-----222131131211--1---12221-322211122311132AAA
Trochonema_panderi 21222111--2121-13131131231111212231132211--1---13311-533112122311132AAA
Lophospira_quadrisulcata 21234221--2221112121121231111212231231111--1---121221212121212112232AA9
Lophospira_sedgwicki 21233111--22211231221121-----21221111---1--1---12131-323121112112221AA6
Lophospira_ornate_sp. 21233111--22211231221121-----21221111---212112112131-323121112112221AA6
Arjamannia_thraivensis 21233111--22211231221212211111?221111---2221---12131-323121112112222AA6
Arjamannia_cancellatula 21233111--22211231221312212111?223111---223223112131-323211112112222BC6
Longstaffia_cyclonema 21223121--22221221211432212111?223111---1--1---13121-222211112112222BCD
Arjamannia_woodlandi 21233111--22211231221312222111?221111---223223212121-313211112112222BC6
Arjamannia_inexpectans 21233111--22211231221312222111?221111---223221212121-313211112112222CC6
Lophospira_holmi 21233111--22211231221121-----??1--???---2--1---12131-313321112112212CE6
Lophospira_gothlandica 21233121--22211231221121-----1?1--??31111--1---121321423211112113213CJ6
Kiviasukkaan_nelsonae 21233121--22211231221121-----1?1--??31111--1---121322423211112113213DD6
Frodospira_imbricata 322331123222211232221311-----22223111---222222121141-321112112112210DH6
Longstaffia_laqueata 32223111--22221121211432212111?223111---1--1---23121-222211112112222DGD
Trochonema_turritum 311???11--2121-23131142221211222231132222221---22131-233311122311132DEA
Ruedemannia_laevissima 11223111--22221111211431-----1?1--??1---212113113121-222211112111212EE7
Ruedemannia_robusta 11223111--22221111211432112111?1--??1---213212113121-222211112111213EH7
Longstaffia_tubulosa 322231223222221132211332212111?223121---222121123121-222211112112222GJD
Loxoplocus_soluta 212331?1--22211231221121-----1?1--??31211--1---1214?-42321111211--?3GHE
'Trochonema (Eunema) kaysi' 311???11--2121-23131142221211222231132222121---22131-233311112311132GGA
Trochonema_muricatum 311???11--2121-23131142221211222231132221--1---22121-233311122311132GHA
Ruedemannia_subrobusta 11223111--22221111211432112111?1--??1---213211113121-222211112111213GJ7
Frodospira_munda 212331123222211232221311-----22223111---1--1---21141-321122112113210GH6
Frodospira_cochleata 312331223222211231221111-----22223111---212212121141-321122112113210GH6
Ptychozone_aberrans 11223111--22231111111541-----1?1--??1---222222113121-222212112111-12HJ7
;
END;
BEGIN SETS;
TAXSET Outgroup = 1-3;
TAXSET CohortA = 1-11;
TAXSET CohortB = 12-22;
TAXSET CohortC = 23-44;
TAXSET bicincta = 5 7-8 11-18 20-22 31-33 37-43;
TAXSET perangulata = 2-6 9-10 18-19 23-30 34-36 44;
END;
BEGIN ASSUMPTIONS;
OPTIONS DEFTYPE=unord PolyTcount=MAXSTEPS ;
TYPESET mytypes = unord: 1-3 5-8 10-13 15-36 38 40-48 52-64 66-71, ord: 4 9 14 37 39 49-51 65;
TYPESET * UNTITLED = unord: 1-3 5-8 10-13 15-36 38 40-48 52-64 66-68 71, ord: 4 9 14 37 39 49-51 65 69-70;
WTSET * UNTITLED = 1: 1-68, 0: 69-71;
END;
BEGIN TREES;
TRANSLATE
1 Sinuopea_sweeti,
2 Hormotoma_confusa,
3 Ectomaria_adelina,
4 Pagodispira_cicelia,
5 Lophospira_perangulata,
6 Lophospira_kangyaoensis,
7 Lophospira_grabaui,
8 Lophospira_milleri,
9 Proturritella_bicarinata,
10 Pagodispira_derwiduii,
11 Pagodispira_dorothea,
12 Trochonemella_knoxvillensis,
13 Lophospira_centralis,
14 Trochonemella_trochonemoides,
15 Trochonemella_montrealensis,
16 Trochonema_bellulum,
17 Lophospira_helicteres,
18 Trochonemella_n.sp._A,
19 Lophospira_serrulata,
20 Lophospira_ventricosa,
21 Trochonemella_notablis,
22 Eunema_strigillatum,
23 Trochonema_umbilicatum,
24 Trochonema_wilsonae,
25 Gyronema_pulchellum,
26 Gyronema_semicarinatum,
27 Gyronema_liljevalli,
28 Lophospira_burginensis,
29 Lophospira_spironema,
30 Lophospira_oweni,
31 Trochonema_madisonense,
32 Lophospira_sumnerensis,
33 Lophospira_decursa,
34 Lophospira_conoidea,
35 Lophospira_producta,
36 Lophospira_Curdsville_sp.,
37 Ruedemannia_humilis,
38 Schizolopha_textilis,
39 Ruedemannia_lirata,
40 Lophospira_tenuistriata,
41 Lophospira_tropidophora,
42 Schizolopha_moorei,
43 Donaldiella_filosa,
44 Lophospira_pulchella,
45 Lophospira_bellicarinata,
46 Donaldiella_bowdeni,
47 Lophospira_aff._L._serrulata,
48 Globonema_kokeni,
49 Ruedemannia_Boda_Ls._sp.,
50 'Trochonemella "G. pulchellum"',
51 Trochonemella_churkini,
52 Trochonemella_reusingi,
53 Trochonema_Alaska_sp.,
54 'Trochonema (Eunema) salteri',
55 Trochonema_panderi,
56 Lophospira_quadrisulcata,
57 Lophospira_sedgwicki,
58 Lophospira_ornate_sp.,
59 Arjamannia_thraivensis,
60 Arjamannia_cancellatula,
61 Longstaffia_cyclonema,
62 Arjamannia_woodlandi,
63 Arjamannia_inexpectans,
64 Lophospira_holmi,
65 Lophospira_gothlandica,
66 Kiviasukkaan_nelsonae,
67 Frodospira_imbricata,
68 Longstaffia_laqueata,
69 Trochonema_turritum,
70 Ruedemannia_laevissima,
71 Ruedemannia_robusta,
72 Longstaffia_tubulosa,
73 Loxoplocus_soluta,
74 'Trochonema (Eunema) kaysi',
75 Trochonema_muricatum,
76 Ruedemannia_subrobusta,
77 Frodospira_munda,
78 Frodospira_cochleata,
79 Ptychozone_aberrans
;
TREE Template = [&R] (1,(2,(3,(4,(5,((7,(8,((12,((14,(16,23)),(15,(18,(((51,52),(47,50)),21))))),((59,45),44),29,(13,(56,(22,17))),((39,38),37)))),19),20,(33,(35,34,36)),(6,(((41,42),(30,32)),(40,28)))),(((11,46),43),10)))));
TREE Truth = [&R] (1,(2,(3,(4,(5,(7,(8,((12,((14,(16,23)),(15,(18,(((51,52),(47,50)),21))))),((59,45),44),29,(13,(56,22,17)),((39,38),37)))),19,20,(33,(35,34,36)),6,(((41,42),(30,32)),(40,28))),(((11,46),43),10)))));
TREE Stratocladogram = [&R] (1,(2,(3,(4,(((5,(7,(8,((12,((14,(16,23)),(15,(18,(((51,52),(47,50)),21))))),((13,(56,(17,22))),(((59,45),44),((38,(39,37)),29))))))),((20,19),(6,(((41,42),(30,32)),(40,28))))),((33,(35,(36,34))),((11,(46,43)),10)))))));
TREE AllLophospirids_1 = [&R] (1,(2,(3,(4,((5,(7,(8,9,(12,((14,(16,(55,31),(23,((((48,((69,74),75)),54),53),24,27)))),(15,(18,(((51,52),(47,50)),21))))),(13,(26,25),(56,17,22)),((((59,((63,62),60)),45),((58,57),((64,((78,67),77)),(73,65)))),44),((38,((39,(((70,(71,76)),((72,68),61)),49)),37)),29))),20,19,6,(33,((35,36),34)),(((41,42),30,32),40,28)),((11,(46,43)),10))))));
TREE AllLophospirids_2 = [&R] (1,(2,(3,(4,((5,(7,(8,9,(12,((14,(16,(55,31),(23,((((48,((69,74),75)),54),53),24,27)))),(15,(18,(((51,52),(47,50)),21))))),(13,(26,25),(56,17,22)),(((59,((63,62),60)),45),(58,57,(64,((78,67),77),(73,65))),44),((38,(39,((((70,79),(71,76)),((72,68),61)),49))),37),29)),20,19,6,(33,((35,36),34)),(((41,42),30,32),40,28)),((11,(46,43)),10))))));
TREE All_through_Silurian = [&R] (1,(2,(3,(4,((10,(11,(43,46))),(5,(7,(8,9,(12,((15,(18,(21,((51,52),(47,50))))),(14,(16,(55,31),(23,(53,(54,(48,(75,(69,74))))),24,27))))),(13,(26,25),(56,17,22)),((45,(59,(60,(63,62)))),(58,57,(64,((78,67),77),(73,65))),44),(37,(38,(39,(49,((61,(72,68)),((70,79),(71,76))))))),29)),20,19,6,(33,(34,(35,36))),(((41,42),30,32),40,28)))))));
TREE * All_Lophospirids_3 = [&R] (1,(2,(3,(4,((10,(11,(43,46))),(5,(7,(8,9,(12,((15,(18,(21,((51,52),(47,50))))),(14,(16,(55,31),(23,((53,(54,(48,(75,(69,74))))),24,27)))))),(13,(26,25),(56,17,22)),((45,(59,(60,(63,62)))),(58,57,(64,(77,(78,67)),((73,66),65))),44),(37,(38,(39,(49,((61,(72,68)),((70,79),(71,76))))))),29)),20,19,6,(33,(34,(35,36))),(((41,42),30,32),40,28)))))));
END;
BEGIN NOTES;
TEXT TAXON=47 TEXT= Rohr_1988;
TEXT TAXON=53 TEXT= Rohr_1988;
TEXT TAXON=48 TEXT= was_Globonema_bicarinatus;
TEXT TAXON=3 TEXT= Mislabelled_Ectomaria_ada_in_paper.;
END;
BEGIN MacClade;
Version 4.0 87;
LastModified -851371076;
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ListWindow Trees closed Gadget '12' '50' '10' '276' '490' ;
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Continuous '0' '3' 1;
Calculations 1000101;
SummaryMode '0' '0' 0;
Charts Arial '12' (normal) 0010;
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ChartBiggestSpot 1;
ChartPercent 10;
ChartBarWidth '10' 1;
ChartVerticalAxis 10101;
ChartMinMax '0';
TraceAllChangesDisplay '1' 1;
BarsOnBranchesDisplay '0' '0' '60000' '10000' '10000' '10000' '10000' '60000' '65000' '65000' '65000' '6' '1' 0000101;
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AllStatesBranchLabels 1;
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ColorOptions 00;
TreeTools '0' '5' '4' '0' '10' '4' '0' 00100111111101110;
EditorTools '0' '0' '0' '1000' '0' '0' '6' '3' '0' 100000101110001;
PairAlign '2' '2' '3' '2' '1' '1' '2' '1' '3' 1010;
BothTools '1';
END;