Rio Cocles [Uscari Fm] (Miocene of Costa Rica)

Where: Costa Rica (9.6° N, 82.8° W: paleocoordinates 9.4° N, 81.5° W)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Uscari Formation, Late/Upper Miocene (11.6 - 5.3 Ma)

• LITHOSTRATIGRAPHY: The Uscari Formation which sits diachronously with abrupt lithological transition below the Rio Banano Formation, mostly conformably and sometimes disconformably. AGE: Late Miocene. STRATIGRAPHIC POSITION: Collection is representative of formation.

• formation-level stratigraphic resolution

Environment/lithology: marine; siliciclastic sediments

• No paleoenvironmental data provided.
• LITHIFICATION: Unknown. LITHOLOGY: No lithologic data provided, but assumed to be siliciclastic.

Size class: macrofossils

Reposited in the PRI

Collection methods: surface (in situ),


Primary reference: A. A. Olsson. 1922. The Miocene of Northern Costa Rica. Bulletins of American Paleontology 9(39):1-309 [A. Hendy/A. Hendy]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 134681: authorized by Austin Hendy, entered by Austin Hendy on 12.10.2012

Creative Commons license: CC BY (attribution)

Taxonomic list

• NOMENCLATURE: Authoritative publication, with species-resolution identifications, and antiquated nomenclature. COVERAGE: Exhaustive for molluscs.
 Pectinida - Pectinoidae
 Neogastropoda - Buccinidae
"Ptychosalpinx dentalis" = Amarophos dentalis
"Ptychosalpinx dentalis" = Amarophos dentalis Olsson 1922 true whelk
 Neogastropoda - Volutidae
Scaphella costaricana n. sp. Olsson 1922 volute
 Tonnoidea - Cassidae
Sconsia cocleana Olsson 1922 snail
 Dentaliida - Dentaliidae
"Dentalium uscarianum" = Fissidentalium uscarianum
"Dentalium uscarianum" = Fissidentalium uscarianum Olsson 1922 tusk shell