Sample B, Titusville Site (1967 excavation) (Pleistocene of the United States)

Where: Pennsylvania (41.6° N, 79.6° W: paleocoordinates 41.6° N, 79.6° W)

• coordinate stated in text

• hand sample-level geographic resolution

When: Titusville Till Formation, Late/Upper Pleistocene (0.1 - 0.0 Ma)

Environment/lithology: mire or swamp; unlithified peat

• Like Zone I, analogues to the Zone III fauna occur today in forest–tundra environments of North America where mean July temperatures range from 10 to 13C.

Size class: mesofossils

Collected in 1967

Collection methods: quarrying, sieve

• All specimens, including those examined by Totten and Coope in 1967, were mounted onto micropaleontological slides and reposited in the Quaternary Entomology Laboratory at North Dakota State University.

Primary reference: S. Cong, A. C. Ashworth, D. P. Schwert and S. M. Totten. 1996. Fossil Beetle Evidence for a Short Warm Interval near 40,000 yr B.P. at Titusville, Pennsylvania. Quaternary Research 45:216-225 [M. Clapham/J. Karr/J. Karr]more details

Purpose of describing collection: paleoecologic analysis

PaleoDB collection 137210: authorized by Matthew Clapham, entered by Jered Karr on 12.12.2012

Creative Commons license: CC BY (attribution)

Taxonomic list

 Coleoptera - Bostrichidae
Bostrichidae indet. Latreille 1802 false powderpost beetle
 Coleoptera - Helophoridae
Helophorus sempervarians Angus 1970 water scavenger beetle
 Coleoptera - Staphylinidae
Olophrum rotundicolle Sahlberg 1817 rove beetle
Boreaphilus henningianus Sahlberg 1832 rove beetle
 Coleoptera - Carabidae
"Carabus maeander" = Carabus (Homoeocarabus) maeander, "Bembidion mutatum" = Bembidion (Bembidion) mutatum, "Agonum simile" = Agonum (Europhilus) simile
"Carabus maeander" = Carabus (Homoeocarabus) maeander Fischer von Waldheim 1822 ground beetle
"Bembidion mutatum" = Bembidion (Bembidion) mutatum Gemminger and Harold 1868 ground beetle
"Agonum simile" = Agonum (Europhilus) simile Kirby 1837 ground beetle