Los Santos de la Humosa (Miocene of Spain)

Also known as Henares 1

Where: Spain (40.5° N, 3.3° W: paleocoordinates 40.7° N, 4.4° W)

• coordinate based on nearby landmark

• small collection-level geographic resolution

When: MN 6 mammal zone, Serravallian (13.8 - 11.6 Ma)

• Late Aragonian

• bed-level stratigraphic resolution

Environment/lithology: terrestrial; lithology not reported

Size class: macrofossils

Collection methods: MNCN

Primary reference: A. Pérez-García and E. Vlachos. 2014. New generic proposal for the European Neogene large testudinids (Cryptodira) and the first phylogenetic hypothesis for the medium and large representatives of the European Cenozoic record. Zoological Journal of the Linnean Society 172:653-719 [E. Vlachos/E. Vlachos]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 187336: authorized by Evangelos Vlachos, entered by Evangelos Vlachos on 21.07.2017

Creative Commons license: CC BY (attribution)

Taxonomic list

 Testudines - Pantestudinidae
Titanochelon bolivari Hernandez-Pacheco 1917 turtle
Los Santos de la Humosa (Madrid, Spain). MNCN collection (17 specimens, Fig. 9) 49366, left humerus; 49367, right radius; 49369, humerus head; 49370, scapula; 49370, right ulna; 49372, humerus head; 49373, humerus? diaphysis; 49374, femur diaphysis; 49375, humerus diaphysis; 49379, humerus head; 49381, pelvis fragment; 49385, right epiplastron; 73733, left humerus; 73734, right epiplastron; 73740, osteoderm; 73750, osteoderm; 73767, ungual.