Kirtlington 3p (Mammal Bed) (Jurassic of the United Kingdom)

Also known as Kirtlington Cement Quarry

Where: England, United Kingdom (51.9° N, 1.3° W: paleocoordinates 41.5° N, 9.1° E)

• coordinate based on nearby landmark

• small collection-level geographic resolution

When: Forest Marble Formation (Great Oolite Group), Late/Upper Bathonian (167.7 - 164.7 Ma)

• Oppelia aspidoides zone

• bed-level stratigraphic resolution

Environment/lithology: mire or swamp; unlithified marl

• "deposited under non-marine conditions in a swampy coastal region" (Gardner et al. 2003)
• "nonmarine clay" and "unconsolidated marl"

Size classes: macrofossils, mesofossils

Collected by Team from University College, London, led by the late Prof. K.A. Kermack; reposited in the BMNH

Collection methods: bulk, sieve,

• BDUC collection, Biology Department, University College, London

Primary reference: E. P. Freeman. 1976. Mammal teeth from the Forest Marble (Middle Jurassic) of Oxfordshire, England. Science 194(4269):1053-1055 [M. Carrano/M. Carrano/R. Benson]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 39075: authorized by Matthew Carrano, entered by Matthew Carrano on 19.05.2004, edited by Kaitlin Maguire, Terri Cleary, John Alroy, Roger Benson, Richard Butler and Philip Mannion

Creative Commons license: CC BY (attribution)

Taxonomic list

unclassified
  -
Chondrichthyes
 Hybodontiformes - Hybodontidae
Hybodus sp.12 Agassiz 1837 elasmobranch
 Hybodontiformes - Acrodontidae
Asteracanthus sp.12 Agassiz 1837 elasmobranch
 Hybodontiformes - Lonchidiidae
Lissodus sp.9 Brough 1935 elasmobranch
Gnathostomata
  -
Actinopterygii indet.12 ray-finned fish
197+ scales, 6+ vertebrae
Actinopteri
 Holostei -
 Amiiformes -
? Amioidea indet.9 Bonaparte 1838 bowfin
 Lepisosteiformes -
cf. Lepidotes sp.9 Agassiz 1843 gar
Osteichthyes
 Temnospondyli - Albanerpetontidae
Anoualerpeton priscus n. sp.13 Gardner et al. 2003 tetrapod
 Salientia - Discoglossidae
Eodiscoglossus oxoniensis n. sp.11 Evans et al. 1990 painted frog
 Caudata -
Caudata "indet. A"11 salamander
"Salamander A (common form)"
Caudata "indet. B"11 salamander
"Salamander B (small form)"
Marmorerpeton sp.10 Evans et al. 1988 salamander
Marmorerpeton kermacki n. gen. n. sp.10 Evans et al. 1988 salamander
Marmorerpeton freemani n. sp.10 Evans et al. 1988 salamander
Reptilia
 Testudines - Pleurosternidae
 Testudines -
Cryptodira "histomorph II"16 turtle
Histomorph II is represented only by a single specimen (IPB R589)
 Saurischia -
? Cetiosaurus sp.12 Owen 1841 sauropod
 Avetheropoda -
Maniraptora indet.1 Gauthier 1986 coelurosaur
 Saurischia - Megalosauridae
Megalosaurus sp.1 Buckland 1824 tetanuran
 Ornithischia -
cf. Alocodon sp.8 Thulborn 1975 ornithischian
 Pterosauria -
Pterosauria indet.20 Kaup 1834 pterosaur
Rhamphorhynchoidea indet.9 Plieninger 1901 pterosaur
Teeth
 Loricata - Goniopholididae
cf. Goniopholis sp.8 Owen 1842 crocodilian
 Loricata -
cf. Theriosuchus sp.19 Owen 1878 crocodilian
 Eosuchia -
Marmoretta oxoniensis n. gen. n. sp.5
Marmoretta oxoniensis n. gen. n. sp.5 Evans 1991 crown diapsid
BMNH R.12020–12146
 Squamata -
Anguimorpha indet.8 Fürbringer 1900 lizard
Eophis underwoodi n. gen. n. sp.3 Caldwell et al. 2015 snake
NHMUK R12355, symphyseal portion of right dentary; NHMUK R12354, midsection right dentary; NHMUK R12370, posterior portion of right dentary; NHMUK R12352, fragment of left maxilla
? Gekkota indet.9 Cuvier 1817 lizard
Scincomorpha indet.8 Camp 1923 lizard
Bellairsia gracilis n. gen. n. sp.7 Evans 1998 lizard
BMNH R12678 (holotype), partial left dentary; BMNH R12679, fragment of right maxilla; BMNH R12680, left premaxilla; further dentary specimens not further elaborated on
Balnealacerta silvestris n. gen. n. sp.7 Evans 1998 lizard
BMNH R12669 (holotype), partial left dentary; BMNH R12670, right maxilla; BMNH R12671, R12672, two left premaxillae; BMNH R12673, R12634, R12635 three frontals; further specimens not further elaborated on
Parviraptor cf. estesi6 Evans 1994 lizard
Series of dissociated elements including BMNH R12353, a left parietal; R12356, a partial left frontal; R12357, left frontal of a juvenile; R12358, another left frontal of a juvenile; R12359, partial left frontal; R12360, axis vertabra; R12361, partial dorsal vertebra; R12362, partial dorsal vert.; R12363, dorsal vert. of juvenile; R12364, fragment of vertebral condyle; R12365, dorsal vert.; R12366, anterior caudal vert.; R12367, anterior caudal vert.; R12368, anterior part of an autotomized caudal vert.; R12369, posterior part of an autotomized vert.; further collection of about 100 dissociated jaws, skull bones and vertebrae. Other specimens from the same reference have subseq. been erected under new taxon Eophis underwoodi
Oxiella tenuis n. gen. n. sp.7 Evans 1998 lizard
BMNH R12689 (holotype), left dentary
 Squamata - Paramacellodidae
Saurillodon sp.9 Estes 1983 lizard
Saurillodon marmorensis n. sp.7 Evans 1998 lizard
BMNH R12650 (holotype), right dentary; BMNH R12651, symphyseal region of right dentary; BMNH R12653, right maxilla; BMNH R12652, left premaxilla; plus "range of highly distinctive vertebrae including BMNH R12654-12668"
 Rhynchocephalia -
Sphenodontia indet.8 rhynchocephalian
 Choristodera -
Cteniogenys sp.4 Gilmore 1928 choristodere
"Reptile B"; NHMUK specimens - skull and postcranial remains
Osteichthyes
 Therapsida - Tritylodontidae
Tritylodontidae indet.12 Cope 1884 therapsid
four cusps or parts of cusps, and a complete incisor similar to Oligokyphus (FM/K 6 [=BMNH M36506])
Stereognathus ooliticus15 Charlesworth 1854 therapsid
NHMUK, OUMNH
 Cynodontia -
Haramiyida sp.2 mammaliaform
BDUC J 562 is a relatively small molariform tooth: Interpretation of J 562 must await the discovery of further material; at present it can only be classified incertae sedis
 Therapsida -
Kirtlingtonia catenata n. gen. n. sp.2
Kirtlingtonia catenata n. gen. n. sp.2 Butler and Hooker 2005 mammaliaform
BMNH M46497, the crown of a right upper molar, probably unerupted; BMNH M46579, right upper molar with root; BMNH M46818, left upper premolar (bed 3p(iii))
Mammalia
 Morganucodonta - Morganucodontidae
Wareolestes rex n. gen. n. sp.12
Wareolestes rex n. gen. n. sp.12 Freeman 1979 mammal
 Theriiformes - Dryolestidae
? Dryolestidae indet. Marsh 1879 mammal
 Pantotheria -
Eupantotheria indet. Kermack and Musset 1958 mammal
 Theriiformes -
Palaeoxonodon ooliticus n. gen. n. sp.
Palaeoxonodon ooliticus n. gen. n. sp. Freeman 1976 mammal
 Prototribosphenida -
Amphitheria indet.12 Kermack et al. 1968 mammal
FM/K 32 (=BMNH M36532)
 Theriiformes -
? Amphitherium sp.12 de Blainville 1838 mammal
FM/K 16 (=BMNH M36516), a left lower molar minus its talonid
 Shuotheridia - Shuotheriidae
Shuotherium sp.17 Chow and Rich 1982 mammal
J.439, a lower left tooth
? Shuotherium sp.17 Chow and Rich 1982 mammal
J.789, right molar; J.808, left upper molar; J.235, left upper molar
 Docodonta - Tegotheriidae
"Shuotherium kermacki n. sp." = Paritatodon kermacki17, Krusatodon kirtlingtonensis n. gen. n. sp.18
"Shuotherium kermacki n. sp." = Paritatodon kermacki17 Sigogneau-Russell 1998 dodocont
J.612, left lower molar; J226, right lower molar
Krusatodon kirtlingtonensis n. gen. n. sp.18 Sigogneau-Russell 2003 dodocont
 Docodonta - Docodontidae
Borealestes serendipitus18 Waldman and Savage 1972 dodocont
Borealestes mussetti n. sp.18 Sigogneau-Russell 2003 dodocont
Cyrtlatherium canei n. gen. n. sp.12 Freeman 1979 dodocont
Peraiocynodon major n. sp.18 Sigogneau-Russell 2003 dodocont
 Docodonta - Simpsonodontidae
Simpsonodon oxfordensis n. gen. n. sp.18
Simpsonodon oxfordensis n. gen. n. sp.18 Kermack et al. 1987 dodocont
Holotpe: J. 100, fragment of right ramus of lower jaw with three postcanine teeth
 Allotheria -
Allotheria indet.2 mammal
BMNH M46234, left I2
Eleutherodon oxfordensis n. gen. n. sp.14 Kermack et al. 1998 mammal
 Multituberculata - Paulchoffatiidae
? Paulchoffatiidae indet.12 Hahn 1969 multituberculate
FM/K 43 (=BMNH M36543): a partial upper incisor referred to as a 'multituberculate or tritylodontid incisor' by Freeman (1979); transferred to just Paulchoffatiidae indet. by Hahn (1993)
 Multituberculata - Hahnotheriidae
Hahnotherium antiquum n. gen. n. sp.2
Hahnotherium antiquum n. gen. n. sp.2 Butler and Hooker 2005 multituberculate
BMNH M46797, a left second upper molar. Described by Kermack et al. 1998: 596, figs. 19B, 21 as Eleutherodon oxfordensis, group
 Multituberculata - Kermackodontidae
Kermackodon multicuspis n. gen. n. sp.2
Kermackodon multicuspis n. gen. n. sp.2 Butler and Hooker 2005 multituberculate
BMNH M46822, a left upper molar (M2)
unclassified
  -
Charophyta indet.9 Migula 1890
Pteridopsida
  -
"Triletes" sp.9 Reinsch 1881
megaspores