PBDB Collection

Shanwang, Linqu County, collection s (Shandong Provincial Museum) (Miocene of China)

Where: Shandong, China (36.6° N, 118.7° E: paleocoordinates 36.5° N, 116.3° E)

• coordinate based on nearby landmark

• local area-level geographic resolution

When: Shanwang Formation, Burdigalian (20.4 - 16.0 Ma)

• Paleomagnetic and radiometric evidence correlates the Shanwang Formation to subchron 5En of Chron 5. The main part of the Shanwang Formation covers 18.524 Ma to 18.056 Ma, with the Shanwang Formation as a whole covering 18.5 ma to 17.5 Ma. Recent paleontological results show that the Shanwang Formation can be correlated with the European mammalian zone MN instead of the previous MN5.

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine - large; lithified, diatomaceous, tuffaceous mudstone

• Two minor volcanic faulted basins, the Linqu and Shanwang Basins, are present in the study area. The former lies near the county town of Linqu, and the latter in the Shanwang National Geopark, east of this town. The faulted basins are the result of intense Mesozoic-Cenozoic tectonic extension and rifting; the development of these basins was closely related in time with the nearby NE-SW and NW-SE trending faults in the Yishu Fault Zone.

Size classes: macrofossils, mesofossils

Preservation: adpression

Collection methods: Repository: Shandong Provincial Museum. This collection contains specimens labeled with five-digit numbers prefaced by letter s.

Primary reference: J. F. Zhang. 1989. Fossil Insects From Shanwang, Shandong, China 1-459 [D. Smith/E. Leckey/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 128552: authorized by Matthew Clapham, entered by Matthew Clapham on 11.06.2012

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Insecta
Ephemeroptera - Heptageniidae	Heptagenia sp. Heptagenia sp. Walsh 1863 stream mayfly s82707, s82712, s82724
Odonata - Coenagrionidae	Agriocnemis lacteola Agriocnemis lacteola Selys 1877 wisp s82682
Odonata - Macromiidae	Epophthalmia zotheca n. sp. Epophthalmia zotheca n. sp. Zhang 1989 true dragonfly s82725/6
Odonata - Aeshnidae	"Mediaeschna lucida n. sp." = Epiaeschna lucida "Mediaeschna lucida n. sp." = Epiaeschna lucida Zhang 1989 darner dragonfly s82681
Hemiptera - Aphididae	"Cinara elegans n. sp." = Cinara (Cinara) elegans, "Cinara limnogena n. sp." = Cinara (Cinara) limnogena "Cinara elegans n. sp." = Cinara (Cinara) elegans Zhang 1989 aphid s82674 "Cinara limnogena n. sp." = Cinara (Cinara) limnogena Zhang 1989 aphid s82673
Hemiptera - Tingidae	Agramma tertiarium n. sp. Agramma tertiarium n. sp. Zhang 1989 lace bug s82730
Hemiptera - Nepidae	Laccotrephes cf. robustus Laccotrephes cf. robustus Stal 1870 true bug s82717
Hemiptera - Corixidae	Corixa triquetra n. sp. Corixa triquetra n. sp. Zhang 1989 water boatman s82679
Hemiptera - Rhopalidae	Stictopleurus breviantennus n. sp. Stictopleurus breviantennus n. sp. Zhang 1989 scentless plant bug s82703
Hemiptera - Urostylididae	? Urochela sp., Urochela pardalina n. sp., Urochela falsirubra n. sp. ? Urochela sp. Dallas 1851 true bug s82674 Urochela pardalina n. sp. Zhang 1989 true bug s82678 Urochela falsirubra n. sp. Zhang 1989 true bug s82677
Hemiptera - Pentatomidae	Stollia rubibrunneus n. sp., Carbula aff. crassiventris, Carbula pustulosa n. sp. Stollia rubibrunneus n. sp. Zhang 1989 stink bug s82716 Carbula aff. crassiventris Dallas 1849 stink bug s82676 Carbula pustulosa n. sp. Zhang 1989 stink bug s82675
Hemiptera - Fulgoridae	Limois shanwangensis Limois shanwangensis Hong 1979 fulgorid planthopper s82668, s82671
Dicondylia -	Oiophassus nycterus n. gen. n. sp. Oiophassus nycterus n. gen. n. sp. Zhang 1989 winged insect s82702
Blattodea - Kalotermitidae	"Kalotermes shandongianus n. sp." = Glyptotermes shandongianus "Kalotermes shandongianus n. sp." = Glyptotermes shandongianus Zhang 1989 termite s82767
Blattodea - Blattidae	Periplaneta lacera n. sp. Periplaneta lacera n. sp. Zhang 1989 cockroach s82696
Orthoptera - Acrididae	Tumearolium humifuse n. gen. n. sp. Tumearolium humifuse n. gen. n. sp. Zhang 1989 slant-faced grasshopper s82697
Dermaptera - Forficulidae	Allodahlia shanwangensis Allodahlia shanwangensis Zhou 1986 common earwig s82727, s82729
Hymenoptera - Tiphiidae	Tiphia mediovena n. sp., Tiphia varia n. sp., Tiphia brevala n. sp. Tiphia mediovena n. sp. Zhang 1989 wasp s82642 Tiphia varia n. sp. Zhang 1989 wasp s82633 Tiphia brevala n. sp. Zhang 1989 wasp s82641
Hymenoptera - Vespidae	"Vespa magnifica" = Vespa mandarinia magnifica "Vespa magnifica" = Vespa mandarinia magnifica Smith 1852 Asian giant hornet s82640
Hymenoptera - Formicidae	Formica ovala, Formica linquensis, "Camponotus microthoracus" = Camponotites microthoracus, "Camponotus ambon n. sp." = Camponotites ambon, "Camponotus gracilis n. sp." = Camponotites gracilis, "Camponotus ampullosus n. sp." = Camponotites ampullosus, "Camponotus shanwangensis" = Camponotites shanwangensis, Camponotus festinus, Dolichoderus evolans n. sp., Dolichoderus lacinius n. sp., Alloiomma changweiensis n. sp., Technomyrmex septentrionalis n. sp., Elaphrodites scutulatus n. gen. n. sp., Myopopone sinensis n. sp. Formica ovala Zhang 1989 ant s82624 Formica linquensis Zhang 1989 ant s82623 "Camponotus microthoracus" = Camponotites microthoracus Zhang 1989 ant s82620 "Camponotus ambon n. sp." = Camponotites ambon Zhang 1989 ant s82621, s82622 "Camponotus gracilis n. sp." = Camponotites gracilis Zhang 1989 ant s82606 "Camponotus ampullosus n. sp." = Camponotites ampullosus Zhang 1989 ant s82603 "Camponotus shanwangensis" = Camponotites shanwangensis Hong 1984 ant s82601, s82604, s82605 Camponotus festinus Smith 1857 ant s82602 Dolichoderus evolans n. sp. Zhang 1989 ant s82625 Dolichoderus lacinius n. sp. Zhang 1989 ant s82627 Alloiomma changweiensis n. sp. Zhang 1989 ant s82629 Technomyrmex septentrionalis n. sp. Zhang 1989 ant s82626 Elaphrodites scutulatus n. gen. n. sp. Zhang 1989 ant s82636 Myopopone sinensis n. sp. Zhang 1989 ant s82628
Hymenoptera - Melittidae	"Dasypoda basaltica n. sp." = Macropis basaltica "Dasypoda basaltica n. sp." = Macropis basaltica Zhang 1989 bee s82637
Hymenoptera - Apidae	Xylocopa diatoma n. sp.², Apis longitibia n. sp.², Bombus luianus n. sp.² Xylocopa diatoma n. sp.² Zhang 1990 large carpenter bee s82772 Apis longitibia n. sp.² Zhang 1990 long-legged honey bee s82773 Bombus luianus n. sp.² Zhang 1990 bumble bee s82771
Hymenoptera - Megachilidae	Megachile shanwangae n. sp. Megachile shanwangae n. sp. Zhang 1989 mason bee s82638
Hymenoptera - Ichneumonidae	Exetastes areolus n. sp., Corynobatopsis anfracta n. gen. n. sp., Agrothereutes townesi n. sp., Mesoleptus stigmus n. sp., ? Ovipterygium flexuosum n. sp., Yezoceryx brachycoleos n. sp. Exetastes areolus n. sp. Zhang 1989 ichneumon wasp s82645 Corynobatopsis anfracta n. gen. n. sp. Zhang 1989 ichneumon wasp s82649 Agrothereutes townesi n. sp. Zhang 1989 ichneumon wasp s82648 Mesoleptus stigmus n. sp. Zhang 1989 ichneumon wasp s82646 ? Ovipterygium flexuosum n. sp. Zhang 1989 ichneumon wasp s82647 Yezoceryx brachycoleos n. sp. Zhang 1989 ichneumon wasp s82644
Hymenoptera -	"Protelenomus miocenicus n. gen. n. sp." = Sinoprotelenomus miocenicus "Protelenomus miocenicus n. gen. n. sp." = Sinoprotelenomus miocenicus Zhang 1989 wasp s82643
Hymenoptera - Tenthredinidae	Selandria perelegans n. sp. Selandria perelegans n. sp. Zhang 1989 wood wasp s82715
Coleoptera - Carabidae	Calosoma cf. maderae, Bembidion strenum n. sp. Calosoma cf. maderae Fabricius 1775 ground beetle s82751 Bembidion strenum n. sp. Zhang 1989 ground beetle s82706
Coleoptera - Dytiscidae	Laccophilus pusulatus n. sp., Laccophilus corystes n. sp. Laccophilus pusulatus n. sp. Zhang 1989 predaceous diving beetle s82721 Laccophilus corystes n. sp. Zhang 1989 predaceous diving beetle s82711
Coleoptera - Scarabaeidae	? Phalops sp., Onthophagus sp., Anomala sp., Anomala scia n. sp., Anomala furva ? Phalops sp. Erichson 1848 scarab beetle s82760 Onthophagus sp. Latreille 1802 scarab beetle s82761 Anomala sp. Samouelle 1819 scarab beetle s82762, s82763 Anomala scia n. sp. Zhang 1989 scarab beetle s82710 Anomala furva Hong 1984 scarab beetle s82755
Coleoptera -	Scelocopris enertheus n. gen. n. sp. Scelocopris enertheus n. gen. n. sp. Zhang 1989 scarab beetle s82754
Coleoptera - Staphylinidae	Lathrobium gensium n. sp., Medon stenus n. sp. Lathrobium gensium n. sp. Zhang 1989 rove beetle s82698 Medon stenus n. sp. Zhang 1989 rove beetle s82699
Coleoptera - Hydrophilidae	Enochrus lapsus n. sp., Helochares mydaleus n. sp., "Hydrous pergrandis n. sp." = Hydrophilus pergrandis Enochrus lapsus n. sp. Zhang 1989 water scavenger beetle s82714 Helochares mydaleus n. sp. Zhang 1989 water scavenger beetle s82720 "Hydrous pergrandis n. sp." = Hydrophilus pergrandis Zhang 1989 water scavenger beetle s82719
Coleoptera - Cleridae	Cleridae indet. Cleridae indet. Latreille 1802 checkered beetle s82748
Coleoptera - Tenebrionidae	Dascillus exiguus n. sp. Dascillus exiguus n. sp. Zhang 1989 comb-clawed beetle s82731
Coleoptera - Endomychidae	"Engonius alviolatus" = Gerstaeckerus alviolatus "Engonius alviolatus" = Gerstaeckerus alviolatus Hong 1984 handsome fungus beetle s82743, s82744, 82745, s82746
Coleoptera - Coccinellidae	Scymnus (Pullus) aff. kawamurai Scymnus (Pullus) aff. kawamurai Ohta 1929 ladybird beetle s82722
Coleoptera - Cerambycidae	Protaxis chinensis n. sp., Delostromatium curvum n. gen. n. sp., Alifasciola sirdena n. gen. n. sp., Mesosa viaria n. sp., Stromatium cf. barbatum, Trosites foveolatus n. gen. n. sp. Protaxis chinensis n. sp. Zhang 1989 longhorned beetle s82694 Delostromatium curvum n. gen. n. sp. Zhang 1989 longhorned beetle s82705 Alifasciola sirdena n. gen. n. sp. Zhang 1989 longhorned beetle s82693 Mesosa viaria n. sp. Zhang 1989 longhorned beetle s82709 Stromatium cf. barbatum Fabricius 1775 longhorned beetle s82708 Trosites foveolatus n. gen. n. sp. Zhang 1989 longhorned beetle s82695
Coleoptera - Chrysomelidae	Chrysomelidae indet., Chrysothoracus tropicus n. gen. n. sp. Chrysomelidae indet. Latreille 1802 leaf beetle s82749 Chrysothoracus tropicus n. gen. n. sp. Zhang 1989 leaf beetle s82701
Coleoptera - Attelabidae	Oedeuops ephemera n. gen. n. sp. Oedeuops ephemera n. gen. n. sp. Zhang 1989 leaf rolling weevil s82689
Coleoptera - Curculionidae	Hylobius turfaceus n. sp., Hylobius plenus Hylobius turfaceus n. sp. Zhang 1989 snout beetle s82691 Hylobius plenus Zhang 1989 snout beetle s82690
Coleoptera - Brentidae	"Balanobius crassipedalis n. sp." = Longifistulia crassipedalis, "Balanobius edentis n. sp." = Longifistulia edentis "Balanobius crassipedalis n. sp." = Longifistulia crassipedalis Zhang 1989 weevil s82688 "Balanobius edentis n. sp." = Longifistulia edentis Zhang 1989 weevil s82683, s82684, s82685, s82687
Coleoptera - Dascillidae	Dascillus sp., Dascillus flatus n. sp., Dascillus shandongianus Dascillus sp. Latreille 1796 soft bodied plant beetle s82750 Dascillus flatus n. sp. Zhang 1989 soft bodied plant beetle s82734, s82735, s82736 Dascillus shandongianus Zhang 1989 soft bodied plant beetle s82732, s82733, s82740, s82741, s82742
Coleoptera - Elateridae	Elater asmodeus n. sp., Elater mitrus n. sp. Elater asmodeus n. sp. Zhang 1989 click beetle s82747 Elater mitrus n. sp. Zhang 1989 click beetle s82713
Coleoptera - Cantharidae	"Curticantharis sp." = Themus "Curticantharis sp." = Themus Motschulsky 1858 soldier beetle s82753
Diptera - Chironomidae	Bythomyia oryctes n. gen. n. sp.¹ Bythomyia oryctes n. gen. n. sp.¹ Zhang 1989 non biting midge s32469
Diptera - Bibionidae	Plecia spinula n. sp., Plecia aculeolata n. sp., Plecia villosa n. sp., "Bibio abdominalis" = Bibio brunettii, Bibio tranopterus, Bibio varineurus, Bibio ventricosus, Bibio expansus n. sp., Bibio colossoceps n. sp. Plecia spinula n. sp. Zhang 1989 love bug s82652, s82675 Plecia aculeolata n. sp. Zhang 1989 love bug s82651 Plecia villosa n. sp. Zhang 1989 love bug s82650, s82766 "Bibio abdominalis" = Bibio brunettii Hardy 1973 march fly s82659 Bibio tranopterus Zhang 1989 march fly s82653 Bibio varineurus Zhang 1989 march fly s82718 Bibio ventricosus Zhang 1989 march fly s82655 Bibio expansus n. sp. Zhang 1989 march fly s82654 Bibio colossoceps n. sp. Zhang 1989 march fly s82658
Diptera - Pallopteridae	Palloptera hypolithica n. sp. Palloptera hypolithica n. sp. Zhang 1989 fly s82665
Diptera - Asilidae	Asilus angustialius n. sp., Asilus brunneitibius n. sp. Asilus angustialius n. sp. Zhang 1989 robber fly s82663 Asilus brunneitibius n. sp. Zhang 1989 robber fly s82664
Arachnida
Araneae - Thomisidae	Miothomisus subnudus n. sp.³, "Ectona brunnea n. sp." = Ecotona brunnea³ Miothomisus subnudus n. sp.³ Zhang et al. 1994 crab spider s83401 "Ectona brunnea n. sp." = Ecotona brunnea³ Zhang et al. 1994 crab spider s85611
Araneae - Araneidae	Araneus luianus n. sp.³ Araneus luianus n. sp.³ Zhang et al. 1994 orb-weaving spider s82704

¹Zhang 1989; ²Zhang 1990; ³Zhang et al. 1994