East shore, Ezequiel Ramos Mexía Reservoir (Cretaceous of Argentina)

Where: Río Negro, Argentina (39.5° S, 68.9° W: paleocoordinates 46.5° S, 45.2° W)

• coordinate stated in text

• small collection-level geographic resolution

When: Candeleros Formation (Neuquén Group), Early/Lower Cenomanian (99.6 - 93.5 Ma)

• Upper levels of the Candeleros Formation (Cenomanian, early Late Cretaceous; Leanza et al., 2004), Río Limay Subgroup, Neuquén Group

• bed-level stratigraphic resolution

Environment/lithology: fluvial; medium-grained, red sandstone and medium-grained, red mudstone

• The rocks of the Candeleros Formation which yielded the remains of Bicentenaria argentina are made up by red medium sized sandstones and mudstones, with abundant dark-brownish paleosoils representing a fluvial environment under braided and meandering regimes, as well as in aeolian conditions (Leanza et al., 2004; Garrido, 2011)

Size class: macrofossils

Collected by R. Spedale in 1998 and 2019

Collection methods: quarrying, surface (in situ), mechanical,

• MPCA-PV, Vertebrate Paleontology Collection, Museo Provincial “Carlos Ameghino”, Río Negro province, Argentina.

•The skeletal remains of *Bicentenaria argentina* were collected by Mr. Raul Spedale in 1998 during an exceptional fall in the water level of the Ezequiel Ramos Mexía Reservoir (Fig. 1). All the bones were collected from a single quarry roughly 40 centimeters long, 30 centimeters wide, and 50 centimeters thick.

•The specimen (MPCA-PV 2003) was found during a field trip performed under the project of relocation and georeferencing of fossil sites in the Río Negro province, carried out by the former Dirección de Patrimonio y Museos from the Río Negro goverment and supported by some professionals of the Universidad Nacional de Rio Negro (UNRN) in June, 2019.

Primary reference: F. E. Novas, M. D. Ezcurra, F. L. Agnolin, D. Pol, and R. Ortíz. 2012. New Patagonian Cretaceous theropod sheds light about the early radiation of Coelurosauria. Revista del Museo Argentino de Ciencias Naturales, nuevo serie 14(1):57-81 [R. Benson/R. Benson/M. Carrano]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 132347: authorized by Roger Benson, entered by Roger Benson on 14.08.2012, edited by Franco Aspromonte

Creative Commons license: CC BY (attribution)

Taxonomic list

Reptilia
 Pterosauria - Azhdarchidae
Azhdarchidae indet. Nessov 1984 pterosaur
MPCA-PV 2003, posterior half of a middle cervical vertebra
 Theropoda -
Bicentenaria argentina n. gen. n. sp.
Bicentenaria argentina n. gen. n. sp. Novas et al. 2012 coelurosaur
MPCA 865, posterior half of skull articulated w/lower jaws; MPCA 866, ~130 bones, incl. frags of 2 articulated premaxillae w/3 pairs of teeth, incomplete R maxilla w/1 tooth, frags of 17 dorsal vertebrae, 14 sacral vertebrae, 20 caudal vertebrae, frags of 2 scapulae, caudodorsal corner of R coracoid, proximal ends of 3 ulnae, distal end of radius, 8 manual unguals, frags of L ilium, proximal frags of 5 pubic shafts, 5 inc. femora, proximal thirds of 2 L tibiae & distal end of R, R astragalus, frags of 5 metatarsals, 15 non-ungual pedal phalanges, 8 pedal unguals, & several isolated rib frags. Collected bones include 3 left femora w/approx. same size (ca. 31 cm long), indicating at least 3 large different specimens. These match with the size observed in several isolated dorsal, sacral and caudal vertebrae w/a closed neurocentral suture. The existence of a R distal femur with an identical morphology, but considerably smaller than the largest ones, indicates the presence of probably younger individuals in the bone assemblage. This statement is supported by the presence of a fragmentary maxilla bearing a tiny tooth with a crown height of 8 mm, obviously corresponding to a juvenile individual.
 Testudines - Chelidae
Chelidae indet. Gray 1825 sideneck turtle
Some turtle plates