Xiaheyan, Zhongwei County (CNU coll) (Carboniferous to of China)

Where: Ningxia, China (37.4° N, 105.0° E: paleocoordinates 15.7° N, 71.0° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Yanghugou Formation, Westphalian B to Westphalian B (316.9 - 309.8 Ma)

• In bed 6, the conodont Declinognathodus marginodosus is a cosmopolitan species that defines the homonymous biozone of latest Bashkirian to earliest Moscovian age (Fig. 15; Nemyrovska, 1999; Davydov et al., 2010; 2012). This age range for bed 6 can be limited to the latest Bashkirian (Duckmantian), as plesiomorphic features (adcarinal grooves, narrow platform and parapets) of co-occurring Neognathodus specimens suggest (Nemyrovska, 1999). A latest Bashkirian age is additionally supported by the discovery of a specimen of Gastrioceras cf. listeri (Fig. 14) in the same bed. Gastrioceras species with this conch morphology (coronate shape with angular umbilical margin and coarse ribs) are representatives of the globally applicable late Bashkirian Branneroceras-Gastrioceras genus zone (Davydov et al., 2012; Korn and Klug, 2015; Fig. 15). Concerning bed 36, the conodont species Idiognathodus praeobliquus has been reported from the Donets and Midcontinent basins, where it indicates a middle Moscovian (Bolsovian) age (Fig. 15; Nemyrovska et al., 1999; Marshall, 2010; Nemyrovska, 2017). A post-Bashkirian age for bed 36 is also indicated by juvenile Idiognathodus specimens, because their prominent rostral ridges, which reach far beyond the anterior margin of the platform, appear to be too advanced for Bashkirian Idiognathodus. A late Moscovian or even Kasimovian age for the uppermost insect-bearing strata cannot be excluded given the fact that more than 45 % of unit 3 is positioned above the uppermost sampled limestone horizon. (Trümper et al., 2019)

• group of beds-level stratigraphic resolution

Environment/lithology: interdistributary bay; lithified siliciclastic sediments

• Western margin of Ordos basin, developed on North China craton

Size class: macrofossils

Preservation: mold/impression

Collection methods: Repository: Capital Normal University, Beijing

Primary reference: J. Prokop and D. Ren. 2007. New significant fossil insects from the Upper Carboniferous of Ningxia in northern China (Palaeodictyoptera, Archaeorthoptera). European Journal of Entomology 104:267-275 [M. Clapham/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 139576: authorized by Matthew Clapham, entered by Matthew Clapham on 14.02.2013, edited by Jered Karr

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Meganisoptera - Meganeuridae
Tupus orientalis12, Shenzhousia qilianshanensis12, "Sinomeganeura huangheensis n. gen. n. sp." = Oligotypus huangheensis18
Tupus orientalis12 Zhang et al. 2012 griffinfly
CNU-NX1-406, 408, 409-428,430,439,448,449, 457
Shenzhousia qilianshanensis12 Zhang and Hong 2006 griffinfly
CNU-NX1-400, 401, 402, 403, 404, 405
"Sinomeganeura huangheensis n. gen. n. sp." = Oligotypus huangheensis18 Prokop et al. 2008 griffinfly
CNU-NX2006003; additional NX1-407,431-436,453,455 in Li et al., 2013
 Dicondylia - Erasipteridae
Erasipterella jini12 Zhang et al. 2012 winged insect
CNU-NX1-438, 440, 441, 442, 443, 444, 447, 458, 459
 Dicondylia -
Sylphalula laliquei n. gen. n. sp.12, Aseripterella sinensis n. gen. n. sp.12
Sylphalula laliquei n. gen. n. sp.12 Li et al. 2013 winged insect
CNU-NX1-452
Aseripterella sinensis n. gen. n. sp.12 Li et al. 2013 winged insect
CNU-NX1-450, 451, 454
 Plecoptera - Gulouidae
Gulou carpenteri n. gen. n. sp.1
Gulou carpenteri n. gen. n. sp.1 Béthoux et al. 2011 stonefly
CNU-NX1-137 - 159
 Cnemidolestodea - Spanioderidae
Sinopteron huangheense n. gen. n. sp., Miamia maimai n. sp.3, Chenxiella liuae n. gen. n. sp.15, Longzhua loculata n. gen. n. sp.8
Sinopteron huangheense n. gen. n. sp. Prokop and Ren 2007 winged insect
CNU-A-NX2006001, NX2006002-1(2)
Miamia maimai n. sp.3 Béthoux et al. 2012 winged insect
CNU-NX1-272 - 280, 319, 284, 296, 299, 313, 316, 481-486
Chenxiella liuae n. gen. n. sp.15 Liu et al. 2009 winged insect
CNU-A-NX2008001
Longzhua loculata n. gen. n. sp.8 Gu et al. 2011 winged insect
CNU-NX1
 Cnemidolestodea - Ctenoptilidae
Ctenoptilus frequens n. sp.4 Chen et al. 2021 winged insect
CNU-NX1-326, 198, 199, 731-759, 764
 Cnemidolestodea -
Xixia huban n. gen. n. sp.9
Xixia huban n. gen. n. sp.9 Gu et al. 2014 winged insect
CNU-NX1-381, 380, 382, 383, 384, 385, 387, 388, 389, 390, 391, 392
 Cnemidolestodea - Parmapteridae
Heterologus duyiwuer n. sp.2 Béthoux et al. 2012 winged insect
CNU-NX1-266
 Dicondylia -
Phtanomiamia gui n. gen. n. sp.5, Sinogerarus pectinatus n. gen. n. sp.10, Protomiamia yangi n. gen. n. sp.7
Phtanomiamia gui n. gen. n. sp.5 Chen et al. 2020 winged insect
CNU-NX1-324
Sinogerarus pectinatus n. gen. n. sp.10 Gu et al. 2017 winged insect
CNU-NX1-343
Protomiamia yangi n. gen. n. sp.7 Du et al. 2016 winged insect
CNU-NX1-703, 701, 25, 95, 331, 702, 704-727
 Grylloblattodea - Sinonamuropteridae
Sinonamuropteris ningxiaensis6 Peng et al. 2005 ice crawler
CNU-NX1
 Dicondylia -
Dictyoptera "indet. 1"19 Latreille 1829 winged insect
CNU-NX1-347
Dictyoptera "indet. 2"19 Latreille 1829 winged insect
CNU-NX1-346
Dictyoptera "indet. 3"19 Latreille 1829 winged insect
CNU-NX1-308
Qilianiblatta namurensis11 Zhang et al. 2013 winged insect
CNU-NX1-303; 301, 302, 304, 336, 337 in Wei et al., 2013
Kinklidoblatta youhei n. sp.19 Wei et al. 2013 winged insect
CNU-NX1-307, 305, 306
 Palaeodictyoptera - Spilapteridae
Tytthospilaptera wangae n. gen. n. sp.14, Sinodunbaria jarmilae n. gen. n. sp.13
Tytthospilaptera wangae n. gen. n. sp.14 Liu et al. 2015 winged insect
CNU-NX1-165 ("Phoenix 1" trench)
Sinodunbaria jarmilae n. gen. n. sp.13 Li et al. 2013 winged insect
CNU-NX1-338
 Palaeodictyoptera - Namuroningxiidae
Namuroningxia elegans n. gen. n. sp.
Namuroningxia elegans n. gen. n. sp. Prokop and Ren 2007 winged insect
CNU-P-NX2006001
 Megasecoptera - Brodiopteridae
Brodioptera sinensis n. sp.17 Pecharová et al. 2015 winged insect
CNU-NX1-600, 601-629, 632-639, 651-657, 659-666
 Megasecoptera - Aykhalidae
Sinopalaeopteryx splendens n. gen. n. sp.16, Sinopalaeopteryx olivieri n. sp.16, Namuroptera minuta n. gen. n. sp.16
Sinopalaeopteryx splendens n. gen. n. sp.16 Pecharová et al. 2015 winged insect
CNU-NX1-641, 667, 672
Sinopalaeopteryx olivieri n. sp.16 Pecharová et al. 2015 winged insect
CNU-NX1-642, 644, 647
Namuroptera minuta n. gen. n. sp.16 Pecharová et al. 2015 winged insect
CNU-NX1-646
 Diaphanopterodea - Sinodiaphidae
Sinodiapha ramosa n. gen. n. sp.20
Sinodiapha ramosa n. gen. n. sp.20 Yang et al. 2020 winged insect
CNU-NX1-348, 349