Also known as ETB H-02
Where: Villaverde, Spain (40.3° N, 3.7° W: paleocoordinates 40.4° N, 3.8° W)
• coordinate estimated from map
• outcrop-level geographic resolution
When: Pleistocene (2.6 - 0.0 Ma)
• middle Pleistocene.
•"Laplana et al. (2015) suggest a late middle Pleistocene age (MIS 6) for H-02 (ETB) because of the occurrence of the proboscidean Palaeoloxodon antiquus together with the rodent Microtus brecciensis. P. antiquus is a species present in the middle Pleistocene to early Late Pleistocene terraces of the Manzanares and Jarama Rivers to the southeast of Madrid (Sesé and Soto, 2002a, 2002b). The latest record of M. brecciensis in the Iberian Peninsula is probably from Sala de los Huesos in the Cueva de Maltravieso (Cáceres, western Spain), dated to MIS 6 or, with less probability, to the base of MIS 5 (Hanquet, 2011). Such a chronological interpretation is also supported by the presence of Microtus arvalis in H-02 (ETB), a species that has recently appeared in small-mammal associations from other localities of the Jarama and Manzanares River terraces. The record of M. arvalis from H-02 (ETB) may represent the oldest for this species in the area (Laplana et al., 2015). In addition, Laplana et al. (2015) stress that the occurrence in H-02 (ETB) of both M. arvalis and Bison priscus suggests cold climatic conditions, later than MIS 8, documented in other localities in the area such as Valdocarros (Panera et al., 2011; Sesé et al., 2011). Thus, this would constitute an additional argument that H-02 (ETB) formed during MIS 6." - Blain et al, 2017
Environment/lithology: fluvial; silty claystone
•bones lying in sandy bars below. The whole unit disappears westward (Fig. 2)." - Blain et al, 2017
Size class: mesofossils
Primary reference: H.-A. Blain, S. Rubio-Jara, J. Panera, D. Uribelarrea, C. Laplana, E. Herráez, and A. Pérez-González. 2017. A new middle Pleistocene (Marine Oxygen Isotope Stage 6) cold herpetofaunal assemblage from the central Iberian Peninsula (Manzanares Valley, Madrid). Quaternary Research 87:499-515 [G. Benevento/G. Benevento]more details
Purpose of describing collection: general faunal/floral analysis
PaleoDB collection 206172: authorized by Gemma Benevento, entered by Gemma Benevento on 01.11.2019
Creative Commons license: CC BY (attribution)
Taxonomic list
| Amphibia | |
|
| |
|
Pelobates cultripes Tschudi 1838 spadefoot toad two maxillae, eight frontoparietals, one sphenethmoid, two squamosals, one vomer, one exoccipital, one parasphenoid, 19 indeterminate cranial fragments with dermal bone ornamentation, one atlas, nine vertebrae, one sacrum, three scapulae, one coracoid, five humeri, one radioulna, three ischio-pubes, 13 tibiofibulae, one tarsal, and one phalanx. Although most of the fossils are markedly incomplete, all of them are very characteristic of this species
| |
|
Bufo cf. spinosus toad One exoccipital, one atlas, 14 vertebrae, one sacrum, one urostyle, seven ilia, two ischio-pubes, three scapulae, one coracoid, five humeri, four radioulnae, five femurs, seven tibiofibulae, one tarsal, and 10 phalanxes
| |
|
Pelophylax perezi Frost et al. 2006 frog 107 bones: one premaxilla, eleven maxillae, one articular sensu lato, one exoccipital, one atlas, 10 vertebrae, one sacrum, four urostyles, three scapulae, three coracoids, one clavicule, one parasternum, 10 humeri, eight radioulnae, eight ilia, three ischio-pubes, one femur, 10 tibiofibulae, and 28 phalanxes.
| |
|
| |
| Reptilia | |
|
Lacertidae indet. Bonaparte 1831 squamates a medium-size lacertid (three vertebrae, one femur, and one tibia) and a small-size lacertid (two maxillae, six dentaries, one fragment of indeterminate tooth-bearing bone, three vertebrae, one femur, two tibiae, and two hemipelves). Although rather incomplete, all these elements, in particular the maxillae and dentaries, are characteristic of the family Lacertidae
| |
|
Coronella girondica Daudin 1803 southern smooth snake one cervical and three small-sized (centrum length <3 mm) fossil trunk vertebrae
| |
|
Natrix natrix Linnaeus 1758 water snake two fragments of maxillae, one cervical vertebra, 39 trunk vertebrae, 13 caudal vertebrae, and three fragments of undetermined vertebrae.
| |
|
Testudines indet. Batsch 1788 turtle Two peripheral plates and various fragments from two indeterminate plates. very probably Emys orbicularis or Mauremys leprosa, the only turtles currently living in the Iberian Peninsula.
| |