Spilia 1 (Pliocene of Greece)

Where: Greece (41.2° N, 23.5° E: paleocoordinates 41.0° N, 23.3° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: MN14 mammal zone, Spilia Formation, Ruscinian (5.3 - 3.2 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: delta plain; sandstone and shelly/skeletal, silty, sandy mudstone

• Sediments have textural relationships and cross-bedding and channel bedding that lead them to be interpreted as being deposited on a prograding delta which developed into an alluvial plain.
• white, well sorted sandstones sometimes with lenses of well rounded pebbles and contain very little calcite. In the center of the area the sandstones interfinger with carbonaceous mudstones, siltstones and fine sandstones with occasiona marls; these sometimes contain fresh water molluscs. Sediments have cross-bedding and channel bedding.

Size class: mesofossils

Primary reference: A. Armour-Brown, H. de Bruijn, C. Maniati, G. Siatos, and P. Niesen. 1977. The Geology of the Neogene sediments north of Serrai and the use of rodent faunas for biostratigraphic control. VI Colloquium on the Geology of the Aegean Region; Athens 1977, Proceedings 2:615-622 [J. Alroy/E. Leckey/E. Leckey]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 31229: authorized by John Alroy, entered by Erin Leckey on 24.04.2003, edited by Evangelos Vlachos and Lars van den Hoek Ostende

Creative Commons license: CC BY (attribution)

Taxonomic list

Amphibia
 Salientia - Pelobatidae
Pelobates aff. praefuscus2 spadefoot toad
two humeri (UU SP1 1144)
Reptilia
 Squamata - Natricidae
Natrix aff. rudabanyaensis2 water snake
one caudal vertebra (UU SP1 1012);
unclassified
  -
Mollusca indet. Linnaeus 1758
"freshwater molluscs"
Amphibia
 Salientia -
Anura indet.2 frog
one premaxilla (UU SP1 1085), 14 maxillae (UU SP1 1045, UU SP1 1051, UU SP1 1052, UU SP1 1055, UU SP1 1056, UU SP1 1060, UU SP1 1062–UU SP1 1064, UU SP1 1111, and UU SP1 1138), six angulars (UU SP1 1106, UU SP1 1117, UU SP1 1139, and UU SP1 1149), one sculptured bone fragment (UU SP1 1020), two atlantes (UU SP1 1039 and UU SP1 1142), 10 trunk vertebrae (UU SP1 1054, UU SP1 1061, and UU SP1 1151), seven fragments of vertebra (UU SP1 1102, UU SP1 1104, UU SP1 1114, UU SP1 1116, UU SP1 1121, and UU SP1 1128), one sacral vertebra (UU SP1 1040), two urostyles (UU SP1 1137 and UU SP1 1148), one scapula (UU SP1 1124), three coracoids (UU SP1 1025, UU SP1 1118, and UU SP1 1153), 11 humeri (UU SP1 1019, UU SP1 1059, UU SP1 1105, UU SP1 1131, UU SP1 1132, UU SP1 1140, UU SP1 1146, and UU SP1 1147), 16 radioulnae (UU SP1 1050, UU SP1 1099, UU SP1 1103, UU SP1 1113, UU SP1 1122, UU SP1 1130, and UU SP1 1135), 13 ilia (UU SP1 1029, UU SP1 1030, UU SP1 1032, UU SP1 1036, UU SP1 1037, UU SP1 1042, and UU SP1 1123), eight femora (UU SP1 1058, UU SP1 1120, UU SP1 1129, UU SP1 1136, and UU SP1 1152), eight tibiofibulae (UU SP1 1101, UU SP1 1107, UU SP1 1119, UU SP1 1125, and UU SP1 1150), nine phalanges (UU SP1 1047–UU SP1 1049, UU SP1 1053, UU SP1 1112, UU SP1 1126, and UU SP1 1127), and 252 indeterminate elements (UU SP1 1023, UU SP1 1084, UU SP1 1086, UU SP1 1091, and UU SP1 1095); and 252 indeterminate elements (UU SP1 1023, UU SP1 1084, UU SP1 1086, UU SP1 1091, and UU SP1 1095
 Salientia - Pelobatidae
Pelobates cf. syriacus4 Boettger 1889 Syrian spadefoot toad
 Salientia - Bufonidae
Bufo cf. viridis4 Laurenti 1768 European green toad
Bufotes viridis2 Laurenti 1768 toad
one femur (UU SP1 1057)
 Salientia - Ranidae
Ranidae indet.2 Rafinesque 1814 frog
one sacral vertebra (UU SP1 1143) and three humeri (UU SP1 1115 and UU SP1 1145);
Rana sp.2 Linnaeus 1758 frog
six humeri (UU SP1 1035, UU SP1 1038, and UU SP1 1108) and one ilium (UU SP1 1109)
Pelophylax sp.2 Fitzinger 1843 frog
13 ilia [UU SP1 1031, UU SP1 1033, and UU SP1 1034 (11 elements)
 Salientia - Hylidae
Hyla sp.4 Laurenti 1768 tree frog
Hyla arborea2 Linnaeus 1758 tree frog
one humerus (UU SP1 1100) and one ilium (UU SP1 1043)
 Salientia - Discoglossidae
Latonia sp.4 von Meyer 1843 painted frog
Latonia cf. ragei2 Hossini 1993 painted frog
one frontoparietal (UU SP1 1022), four maxillae (UU SP1 1018, UU SP1 1024, UU SP1 1044, and UU SP1 1046), two atlantes (UU SP1 1028), three trunk vertebrae [UU SP1 1021 (two elements) and UU SP1 1027], one urostyle (UU SP1 1134), and five ilia (UU SP1 1017, UU SP1 1026, UU SP1 1110, UU SP1 1133, and UU SP1 1141)
 Caudata - Urodela
Urodela indet.2 Duméril 1805 salamander
three fragmentary vertebrae (UU SP1 1066–UU SP1 1068) and one humerus (UU SP1 1065);
Reptilia
 Squamata - Lacertidae
Lacertidae indet.2 Bonaparte 1831 squamates
one dentary (UU SP1 1001);
cf. Lacerta sp.2 Linnaeus 1758 squamates
one dentary (UU SP1 1002)
 Testudines -
Testudinoidea indet.2 Gray 1825 turtle
two peripherals (UU SP1 1087 and UU SP1 1098), many shell fragments (UU SP1 1070, UU SP1 1071, UU SP1 1072, UU SP1 1089, UU SP1 1090a, UU SP1 1093a, UU SP1 1096, and UU SP1 1097), a ?coracoid fragment (UU SP1 1074), a scapula/acromion fragment (UU SP1 1088), a scapula (UU SP1 1092), nine ungual phalanges [UU SP1 1075, UU SP1 1076, UU SP1 1077 (six elements), and UU SP1 1094], two cervical vertebrae (UU SP1 1073 and UU SP1 1090b), three caudal vertebrae [UU SP1 1078 and UU SP1 1079 (two elements)], two vertebral fragments (UU SP1 1093b), and an unidentifiable fragment (UU SP1 1080);
Mammalia
 Lagomorpha - Ochotonidae
 Lagomorpha - Leporidae
 Rodentia - Muridae
 Soricomorpha - Talpidae
Desmana verestchagini3 Topachevski 1961 Russian desman
Colubriformes
 Colubriformes -
Colubriformes indet.2 Günther 1864
seven trunk vertebrae (UU SP1 1004–UU SP1 1008, UU SP1 1013, and UU SP1 1016), three fragments of trunk vertebrae (UU SP1 1014), and one caudal vertebra (UU SP1 1015);
Colubroides
 Colubriformes -
? Paraxenophis spanios2 Georgalis et al. 2019
three trunk vertebrae (UU SP1 1003, UU SP1 1009, and UU SP1 1010);
Periergophis micros2 Georgalis et al. 2019
one posterior mid- or posterior trunk vertebra (UU SP1 1011);