Subfamily Varuninae Milne-Edwards 1853 (crab)
Malacostraca - Decapoda - Varunidae
Parent taxon: Varunidae according to C. E. Schweitzer et al. 2023
See also Karasawa 1993, Rathbun 1909 and Schweitzer et al. 2010
Sister taxa: Asthenognathinae, Cyclograpsinae, Gaeticinae, Sakakurapsus, Thalassograpsinae
Subtaxa: Acmaeopleura Brachynotus Eriocheir Hemigrapsus Utica Varuna
Type: Varuna
Ecology: infaunal detritivore
Distribution:
• Quaternary of United States (6: California collections)
• Pliocene to Holocene of United States (1: California)
• Miocene of Hungary (6)
• Oligocene of Italy (1), Japan (2)
• Eocene of Italy (4)
Total: 20 collections including 23 occurrences