Malacostraca - Decapoda - Varunidae
Parent taxon: Grapsoidea according to C. E. Schweitzer et al. 2023
See also Schweitzer et al. 2010
Sister taxa: Daragrapsus, Daranyia, Gecarcinidae, Glyptograpsidae, Grapsidae, Percnidae, Plagusiidae, Pseudodaranyia, Sesarmidae, Xenograpsidae
Subtaxa: Asthenognathinae Cyclograpsinae Gaeticinae Sakakurapsus Thalassograpsinae Varuninae
Type: Varuna
Ecology: infaunal detritivore
Distribution:
• Quaternary of United States (7: California, Florida collections)
• Pliocene to Holocene of United States (1: California)
• Pliocene of Italy (1), New Zealand (1)
• Miocene of Argentina (2), Hungary (7), Japan (6), United States (2: Oregon)
• Oligocene of Italy (2), Japan (2), United States (3: Washington)
• Eocene of Argentina (1), Italy (4)
Total: 39 collections including 42 occurrences