Family Callionymidae (ray-finned fish)
Actinopteri - Syngnathiformes - Callionymidae
Parent taxon: Callionymoidei according to R. Betancur et al. 2013
See also Jordan 1963 and Nelson et al. 2016
Sister taxon: Gilmourella
Subtaxa: Callionymus Protonymus
Ecology: nektonic omnivore
Distribution:
• Miocene of India (1 collection), Italy (2), Kazakhstan (1), New Zealand (1), Romania (1), the Russian Federation (1), Serbia and Montenegro (1)
• Waitakian of New Zealand (1)
• Eocene of Belgium (1), the United Kingdom (5)
Total: 15 collections including 18 occurrences