Insecta - Coleoptera - Carabidae

Parent taxon: Harpalinae according to P. Bouchard et al. 2011

See also LeConte 1861

Sister taxa: Abacetini, Aethocarabus, Chlaeniini, Dryptini, Galeritini, Harpalini, Lebiini, Licinini, Odacanthini, Panagaeini, Pseudomorphini, Pterostichini, Sinis, Sphodrini, Zabrini, Zuphiini, Oodini

Subtaxa: Omphrina

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Ecology:

Distribution: found only at East Africa Copal (Coburg collection) (Quaternary of Tanzania)

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